taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03E487C4B22DFFA818B2F919FC21FBF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112938/files/figure.png	https://doi.org/10.5281/zenodo.19112938	Fig. 4. Shells of Corrosella ballestae Delicado & Ramos sp. nov. A–C, F–H. Paratypes (MNCN 15.05/63703P). D–E. Holotype (MNCN 15.05/63703H), frontal and lateral views. F–G. Details of the protoconch. H. Protoconch microsculpture.	Fig. 4. Shells of Corrosella ballestae Delicado & Ramos sp. nov. A–C, F–H. Paratypes (MNCN 15.05/63703P). D–E. Holotype (MNCN 15.05/63703H), frontal and lateral views. F–G. Details of the protoconch. H. Protoconch microsculpture.	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
03E487C4B22DFFA818B2F919FC21FBF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112942/files/figure.png	https://doi.org/10.5281/zenodo.19112942	Fig. 5. Anatomy of Corrosella ballestae Delicado & Ramos sp. nov., paratype (MNCN 15.05/63704), from a pond near Játar. A. Operculum, inner side. B. Operculum, outer side. C. Overview of radular teeth rows. D–E. Detailed view of the central and outer marginal teeth, respectively. F. Ctenidium and osphradium. G. Stomach. H. Partial nervous system. I. Pallial oviduct. J. Bursa copulatrix and seminal receptacle. K. Head of male and penis. Abbreviations: Ag = albumin gland; Bc = bursa copulatrix; Bd = bursal duct; Cg = capsule gland; Ct = ctenidium; Os = osphradium; Ro = renal oviduct; Sr = seminal receptacle.	Fig. 5. Anatomy of Corrosella ballestae Delicado & Ramos sp. nov., paratype (MNCN 15.05/63704), from a pond near Játar. A. Operculum, inner side. B. Operculum, outer side. C. Overview of radular teeth rows. D–E. Detailed view of the central and outer marginal teeth, respectively. F. Ctenidium and osphradium. G. Stomach. H. Partial nervous system. I. Pallial oviduct. J. Bursa copulatrix and seminal receptacle. K. Head of male and penis. Abbreviations: Ag = albumin gland; Bc = bursa copulatrix; Bd = bursal duct; Cg = capsule gland; Ct = ctenidium; Os = osphradium; Ro = renal oviduct; Sr = seminal receptacle.	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
03E487C4B22DFFA818B2F919FC21FBF7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112930/files/figure.png	https://doi.org/10.5281/zenodo.19112930	Fig. 1. Bayesian phylogeny of species of Corrosella Boeters, 1970 based on a concatenated dataset of COI, 16S and 28S sequences. Bayesian posterior probabilities>0.90 are indicated at nodes above the species level. Vertical bars on the right denote groups identified by molecular delimitation methods: Assemble Species by Automatic Partitioning (ASAP), the single-threshold generalized mixed Yulecoalescent (GMYC) model and the Bayesian Poisson Tree Processes (PTP) method. Major clades are distinguished by colour coding, with an inset showing their geographic distribution and representative photo vouchers for species within Clade 1. Species names are followed by locality codes, detailed in Supp. file 1: Table S1. Scale bar below the topology: substitutions per site.	Fig. 1. Bayesian phylogeny of species of Corrosella Boeters, 1970 based on a concatenated dataset of COI, 16S and 28S sequences. Bayesian posterior probabilities>0.90 are indicated at nodes above the species level. Vertical bars on the right denote groups identified by molecular delimitation methods: Assemble Species by Automatic Partitioning (ASAP), the single-threshold generalized mixed Yulecoalescent (GMYC) model and the Bayesian Poisson Tree Processes (PTP) method. Major clades are distinguished by colour coding, with an inset showing their geographic distribution and representative photo vouchers for species within Clade 1. Species names are followed by locality codes, detailed in Supp. file 1: Table S1. Scale bar below the topology: substitutions per site.	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
03E487C4B229FFB71B16FBB1FDF1F91C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112944/files/figure.png	https://doi.org/10.5281/zenodo.19112944	Fig. 6. Shells of Corrosella hinzi (Boeters, 1986). A, E–I. Balsa de Vargas, Borja, Zaragoza (MNCN 15.05/63698). B. Spring of river park in Calamocha, Teruel (MNCN 15.05/63701). C. Prado Spring, Caminreal, Teruel (MNCN 15.05/63702). D. Cazuelas Spring, Borja, Zaragoza (MNCN 15.05/63699). A–E. Frontal views. F. Lateral view. G–H. Details of the protoconch. I. Protoconch microsculpture.	Fig. 6. Shells of Corrosella hinzi (Boeters, 1986). A, E–I. Balsa de Vargas, Borja, Zaragoza (MNCN 15.05/63698). B. Spring of river park in Calamocha, Teruel (MNCN 15.05/63701). C. Prado Spring, Caminreal, Teruel (MNCN 15.05/63702). D. Cazuelas Spring, Borja, Zaragoza (MNCN 15.05/63699). A–E. Frontal views. F. Lateral view. G–H. Details of the protoconch. I. Protoconch microsculpture.	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
03E487C4B229FFB71B16FBB1FDF1F91C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112950/files/figure.png	https://doi.org/10.5281/zenodo.19112950	Fig. 7. Anatomy of Corrosella hinzi (Boeters, 1986). A–J, L. Balsa de Vargas, Borja, Zaragoza (MNCN 15.05/63698). K, N. Spring of river park in Calamocha, Teruel (MNCN 15.05/63701). M. Cazuelas Spring, Borja, Zaragoza (MNCN 15.05/63699). A. Operculum, inner side. B. Operculum, outer side. C. Overview of radular teeth rows. D–E. Detailed views of the central and outer marginal teeth, respectively. F. Ctenidium and osphradium. G. Stomach. H. Partial nervous system. I. Pallial oviduct. J–K. Bursa copulatrix and seminal receptacle. L. Head of male and penis. M–N. Penises.	Fig. 7. Anatomy of Corrosella hinzi (Boeters, 1986). A–J, L. Balsa de Vargas, Borja, Zaragoza (MNCN 15.05/63698). K, N. Spring of river park in Calamocha, Teruel (MNCN 15.05/63701). M. Cazuelas Spring, Borja, Zaragoza (MNCN 15.05/63699). A. Operculum, inner side. B. Operculum, outer side. C. Overview of radular teeth rows. D–E. Detailed views of the central and outer marginal teeth, respectively. F. Ctenidium and osphradium. G. Stomach. H. Partial nervous system. I. Pallial oviduct. J–K. Bursa copulatrix and seminal receptacle. L. Head of male and penis. M–N. Penises.	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
03E487C4B236FFBE1B0AF906FC86FD7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112952/files/figure.png	https://doi.org/10.5281/zenodo.19112952	Fig. 8. Shells of Corrosella navasiana (Fagot, 1907). A–B. Fonnueva Spring, Bulbuente, Zaragoza (MNCN 15.05/53718). C–D. Ojos de Cimballa Wetland, Zaragoza (MNCN 15.05/63617). E. A spring in Arbujuelo, Soria (MNCN 15.05/63625). F–G. Tinte Spring, Medinaceli, Soria (MNCN 15.05/63643). H–I. Manadero River, Guadalajara (MNCN 15.05/63654). J. Tía Perra Spring, El Hosquillo, Cuenca (MNCN 15.05/63695). K–L. Pozo Azul Spring, Covanera, Burgos (MNCN 15.05/63675). M. A stream in Tubilleja, Burgos (MNCN 15.05/63677). N. La Toba Spring, Tubilla del Agua, Burgos (MNCN 15.05/63673). O. A stream in Valtubilla, Burgos (MNCN 15.05/63678). P–Q. María Spring, Ontígola, Toledo (MNCN 15.05/63670). R. A ditch in Borox, Toledo (MNCN 15.05/63668). S–T. A spring in Peralejos de las Truchas, Guadalajara (MNCN 15.05/63653). U–V. A spring next to the Sanctuary Virgen de la Fuencisla, Segovia (MNCN 15.05/97257). W–X. Aceña Spring, Quintanilla de Onésimo, Valladolid (MNCN 15.05/97200).	Fig. 8. Shells of Corrosella navasiana (Fagot, 1907). A–B. Fonnueva Spring, Bulbuente, Zaragoza (MNCN 15.05/53718). C–D. Ojos de Cimballa Wetland, Zaragoza (MNCN 15.05/63617). E. A spring in Arbujuelo, Soria (MNCN 15.05/63625). F–G. Tinte Spring, Medinaceli, Soria (MNCN 15.05/63643). H–I. Manadero River, Guadalajara (MNCN 15.05/63654). J. Tía Perra Spring, El Hosquillo, Cuenca (MNCN 15.05/63695). K–L. Pozo Azul Spring, Covanera, Burgos (MNCN 15.05/63675). M. A stream in Tubilleja, Burgos (MNCN 15.05/63677). N. La Toba Spring, Tubilla del Agua, Burgos (MNCN 15.05/63673). O. A stream in Valtubilla, Burgos (MNCN 15.05/63678). P–Q. María Spring, Ontígola, Toledo (MNCN 15.05/63670). R. A ditch in Borox, Toledo (MNCN 15.05/63668). S–T. A spring in Peralejos de las Truchas, Guadalajara (MNCN 15.05/63653). U–V. A spring next to the Sanctuary Virgen de la Fuencisla, Segovia (MNCN 15.05/97257). W–X. Aceña Spring, Quintanilla de Onésimo, Valladolid (MNCN 15.05/97200).	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
03E487C4B236FFBE1B0AF906FC86FD7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112954/files/figure.png	https://doi.org/10.5281/zenodo.19112954	Fig. 9. Protoconch and radula of Corrosella navasiana (Fagot, 1907). A–B, D–F. Fonnueva Spring, Bulbuente, Zaragoza (MNCN 15.05/53718). C, G–I. Pozo Azul Spring, Covanera, Burgos (MNCN 15.05/63675). J–K. María Spring, Ontígola, Toledo (MNCN 15.05/63670). L. Tía Perra Spring, El Hosquillo, Cuenca (MNCN 15.05/63695).A. Details of the protoconch. B–C. Protoconch microsculpture. D–L. Overview of radular teeth rows and detailed view of the central, lateral and marginal teeth.	Fig. 9. Protoconch and radula of Corrosella navasiana (Fagot, 1907). A–B, D–F. Fonnueva Spring, Bulbuente, Zaragoza (MNCN 15.05/53718). C, G–I. Pozo Azul Spring, Covanera, Burgos (MNCN 15.05/63675). J–K. María Spring, Ontígola, Toledo (MNCN 15.05/63670). L. Tía Perra Spring, El Hosquillo, Cuenca (MNCN 15.05/63695).A. Details of the protoconch. B–C. Protoconch microsculpture. D–L. Overview of radular teeth rows and detailed view of the central, lateral and marginal teeth.	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
03E487C4B236FFBE1B0AF906FC86FD7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112961/files/figure.png	https://doi.org/10.5281/zenodo.19112961	Fig. 10. Heads and penises of Corrosella navasiana (Fagot, 1907). A. Fonnueva Spring, Bulbuente, Zaragoza (MNCN 15.05/53718). B. Ojos de Cimballa Wetland, Zaragoza (MNCN 15.05/53717). C. A spring in Arbujuelo, Soria (MNCN 15.05/63625). D. Tinte Spring, Medinaceli, Soria (MNCN 15.05/63643). E. Tía Perra Spring, El Hosquillo, Cuenca (MNCN 15.05/63695). F–H. Pozo Azul Spring, Covanera, Burgos (MNCN 15.05/63675). I. La Toba Spring, Tubilla del Agua, Burgos (MNCN 15.05/63673). J. A stream in Valtubilla, Burgos (MNCN 15.05/63678). K–L. A ditch in Borox, Toledo (MNCN 15.05/63668). M–N. A spring in Peralejos de las Truchas, Guadalajara (MNCN 15.05/63653). O–P. Aceña Spring, Quintanilla de Onésimo, Valladolid (MNCN 15.05/97200). Q–S. A spring next to the Sanctuary Virgen de la Fuencisla, Segovia (MNCN 15.05/97257). All penises are shown in dorsal view except L, which is presented in ventral view.	Fig. 10. Heads and penises of Corrosella navasiana (Fagot, 1907). A. Fonnueva Spring, Bulbuente, Zaragoza (MNCN 15.05/53718). B. Ojos de Cimballa Wetland, Zaragoza (MNCN 15.05/53717). C. A spring in Arbujuelo, Soria (MNCN 15.05/63625). D. Tinte Spring, Medinaceli, Soria (MNCN 15.05/63643). E. Tía Perra Spring, El Hosquillo, Cuenca (MNCN 15.05/63695). F–H. Pozo Azul Spring, Covanera, Burgos (MNCN 15.05/63675). I. La Toba Spring, Tubilla del Agua, Burgos (MNCN 15.05/63673). J. A stream in Valtubilla, Burgos (MNCN 15.05/63678). K–L. A ditch in Borox, Toledo (MNCN 15.05/63668). M–N. A spring in Peralejos de las Truchas, Guadalajara (MNCN 15.05/63653). O–P. Aceña Spring, Quintanilla de Onésimo, Valladolid (MNCN 15.05/97200). Q–S. A spring next to the Sanctuary Virgen de la Fuencisla, Segovia (MNCN 15.05/97257). All penises are shown in dorsal view except L, which is presented in ventral view.	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
03E487C4B236FFBE1B0AF906FC86FD7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112963/files/figure.png	https://doi.org/10.5281/zenodo.19112963	Fig. 11. Distal female genitalia of Corrosella navasiana (Fagot, 1907). A. Fonnueva Spring, Bulbuente, Zaragoza (MNCN 15.05/53718). B. Dulce River, Cabrera, Guadalajara (MNCN 15.05/63656). C. Canalejas Spring, Somolinos, Guadalajara (MNCN 15.05/63660). D. Tía Perra Spring, El Hosquillo, Cuenca (MNCN 15.05/63695). E–F. Pozo Azul Spring, Covanera, Burgos (MNCN 15.05/63675). G. La Toba Spring, Tubilla del Agua, Burgos (MNCN 15.05/63673). H. A stream in Valtubilla, Burgos (MNCN 15.05/63678). I. A ditch in Borox, Toledo (MNCN 15.05/63668). J–K. A spring in Peralejos de las Truchas, Guadalajara (MNCN 15.05/63653). L–M. A spring next to the Sanctuary Virgen de la Fuencisla, Segovia (MNCN 15.05/97257). N–O. Aceña Spring, Quintanilla de Onésimo, Valladolid (MNCN 15.05/97200).	Fig. 11. Distal female genitalia of Corrosella navasiana (Fagot, 1907). A. Fonnueva Spring, Bulbuente, Zaragoza (MNCN 15.05/53718). B. Dulce River, Cabrera, Guadalajara (MNCN 15.05/63656). C. Canalejas Spring, Somolinos, Guadalajara (MNCN 15.05/63660). D. Tía Perra Spring, El Hosquillo, Cuenca (MNCN 15.05/63695). E–F. Pozo Azul Spring, Covanera, Burgos (MNCN 15.05/63675). G. La Toba Spring, Tubilla del Agua, Burgos (MNCN 15.05/63673). H. A stream in Valtubilla, Burgos (MNCN 15.05/63678). I. A ditch in Borox, Toledo (MNCN 15.05/63668). J–K. A spring in Peralejos de las Truchas, Guadalajara (MNCN 15.05/63653). L–M. A spring next to the Sanctuary Virgen de la Fuencisla, Segovia (MNCN 15.05/97257). N–O. Aceña Spring, Quintanilla de Onésimo, Valladolid (MNCN 15.05/97200).	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
03E487C4B236FFBE1B0AF906FC86FD7A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19112932/files/figure.png	https://doi.org/10.5281/zenodo.19112932	Fig. 2. Morphological differentiation among species of Corrosella Boeters, 1970. The left column shows canonical variate analysis (CVA) results for all species across the major clades, while the right column focuses on species within Clade 1. Analyses were conducted using four datasets. A. Principal components of shell shape. B. Shell measurements. C. Female genitalia measurements. D. Male genitalia measurements. The sample size (n) for each dataset is provided in the top right corner of each plot.	Fig. 2. Morphological differentiation among species of Corrosella Boeters, 1970. The left column shows canonical variate analysis (CVA) results for all species across the major clades, while the right column focuses on species within Clade 1. Analyses were conducted using four datasets. A. Principal components of shell shape. B. Shell measurements. C. Female genitalia measurements. D. Male genitalia measurements. The sample size (n) for each dataset is provided in the top right corner of each plot.	2026-02-24	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten		Zenodo	biologists	Delicado, Diana;Miller, Jonathan P.;Boulaassafer, Khadija;Ramos, Marian A.;Hauffe, Torsten			
