identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E787B2241DFF9B8AE1CFC91D3413CE.text	03E787B2241DFF9B8AE1CFC91D3413CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexasepalum bacigalupoae Cabana Fader & E. L. Cabral 2026	<div><p>Hexasepalum bacigalupoae Cabaña Fader &amp; E.L.Cabral sp. nov.</p><p>urn:lsid:ipni.org:names:77382787-1</p><p>Figs 1–3</p><p>Diagnosis</p><p>Hexasepalum bacigalupoae sp. nov. is similar to H. apiculatum (Willd. ex Roem. &amp; Schult.) Delprete &amp; J.H.Kirkbr. but differs by the leaf blade margins without an epidermal thickening (vs leaf blade margins with a white epidermal thickening in H. apiculatum), conspicuous secondary veins on the abaxial side (vs inconspicuous), and mericarps with flat ventral faces (vs mericarps with two depressions on ventral faces, one on each side of the midline). Hexasepalum bacigalupoae is also similar to H. radulum (Willd.) Delprete &amp; J.H.Kirkbr. but differs by the 3–5-fimbriate stipular sheath (vs 6–12-fimbriate), glabrous fimbriae (vs pilose fimbriae), dorsal face of the mericarps with 3–4 soft ribs (vs dorsal face of the mericarps smooth), and exine of pollen grains covered by spines and small granules (vs only microspines).</p><p>Etymology</p><p>The specific epithet honours the renowned Rubiaceae specialist Professor Nélida Bacigalupo, researcher at the Darwinion Institute in Buenos Aires, Argentina, who pioneered the study of the Spermacoceae tribe in South America.</p><p>Material examined</p><p>Type</p><p>BRAZIL – Bahia • Palmeiras, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.561943&amp;materialsCitation.latitude=-12.639444" title="Search Plazi for locations around (long -41.561943/lat -12.639444)">Volta da Serra na estrada de Palmeiras para a Guiné</a>; 12°38′22″ S, 41°33′43″ W; 1135 m a.s.l.; 4 May 2009; Queiroz et al. 14587; holotype: HUEFS [HUEFS 156605]; isotypes: CTES, SI .</p><p>Paratypes</p><p>BRAZIL – Bahia • Capão da Volta, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.406418&amp;materialsCitation.latitude=-13.444305" title="Search Plazi for locations around (long -41.406418/lat -13.444305)">estrada para Jussiape</a>; 13°26′39.5″ S, 41°24′23.1″ W; 1030 m a.s.l.; 18 May 1999; Souza et al. 22723; CTES, ESA [ESA 061228] • Mucugê, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.438335&amp;materialsCitation.latitude=-13.024445" title="Search Plazi for locations around (long -41.438335/lat -13.024445)">estrada para Barra da Estiva ao sul de Mucugê</a>; 13°01′28″ S, 41°26′18″ W; 1030 m a.s.l.; 17 Aug. 2008; Queiroz et al. 13709; CTES, HUEFS [HUEFS 141925] • same data as for preceding; Queiroz et al. 13711; CTES, HUEFS [HUEFS 141927] .</p><p>Description</p><p>Subshrubs, up to 50 cm tall. Stems decumbent, internodes 1.5–2.5 cm long, tetragonal to cylindrical, glabrous or with few isolated hairs, basally decorticate. Stipular sheath glabrous, 3–5-fimbriate, fimbriae 1–2.5 mm long, succulent, glabrous. Leaves opposite, sessile; blade 15–30 × 3–5 mm, elliptic to lanceolate, apex acute to mucronate, base attenuate, margins slightly revolute, succulent, coriaceous when dry, glabrous, 3–4 secondary veins on each side of the midrib, conspicuous on the abaxial side. Inflorescences axillary, 5–12 glomeruli per flowering branch, glomerules 1–15-flowered, foliaceous bracts, succulent. Flowers sessile; hypanthium obovate, glabrous; calyx 4-lobed, tube absent, lobes 0.8–1.5 mm long, glabrous, sometimes trichomes at the margin; corolla 4.8–5 mm long, 4-lobed, infundibuliform, lilac with pink lobes, externally glabrous, internally with a ring of moniliform trichomes near the base of the tube; stamens exserted, slightly shorter than the corolla lobes; pollen grains 19–20-zonocolporate, suboblate to oblate-spheroidal, endoaperture forming an endocingulum, spines and small granules irregularly distributed over the grain; style 5.5 mm long, glabrous, stigma bilobate, each lobe globose; nectariferous disk entire. Schizocarps 3.5–4 mm long, globose, glabrous, mericarps with dorsal face convex, with 3–4 soft longitudinal ribs, ventral face flat, without depressions. Seeds 1.9–2.1 mm long, subtriangular in outline, brownish–black, dorsal face convex, with longitudinal grooves, ventral face flat, Y-shaped, partially covered by the strophiole, with numerous raphides; testa reticulate-areolate or pitted, elongated or isodiametric cells.</p><p>Distribution, habitat, and phenology</p><p>Hexasepalum bacigalupoae sp. nov. is endemic to Bahia and grows on white sand and rocky soils in the ʻcampo rupestreʼ vegetation of the ʻChapada Diamantinaʼ (Fig. 3A). It is known from the municipalities of Capão da Volta, Mucugê, and Palmeiras (Fig. 4). Specimens with flowers and fruits were collected in May and August.</p><p>Preliminary conservation status</p><p>Hexasepalum bacigalupoae sp. nov. is a rare species, known only from four herbarium specimens representing three locations (Fig. 4). Furthermore, to our knowledge, H. bacigalupoae does not occur within the limits of the ʻParque Nacional da Chapada Diamantinaʼ, a conservation unit of integral protection located in the region.</p><p>The impacts on the ʻcampo rupestreʼ, the vegetation where H. bacigalupoae sp. nov. occurs, are usually associated with opencast mining, annual anthropogenic burnings to support the cattle industry, wood extraction, invasive species, harvesting ornamental plants (orchids, bromeliads, everlastings), road construction, and uncontrolled urbanization, especially linked to tourism expansion, and eucalyptus plantations (Silveira et al. 2016). Specifically in the context of the ʻChapada Diamantinaʼ, severe impacts are caused by the ever-expanding, irrigated, heavily mechanised, and agrochemical-based cropland system developments in the region (Funch et al. 2009).Also, road construction and urban expansion in the valleys are quite noticeable in the analysis of satellite images on Google Earth.</p><p>Hexasepalum bacigalupoae sp. nov. has EOO and AOO equal to 240.640 km 2 and 12 km 2, respectively (kml file available in Supp. file 1). Therefore, the three occurrences represent three locations, and the species is preliminary assessed as Endangered EN B1ab(iii)+B2ab(iii), based on the EOO (less than 5000 km 2) and AOO (less than 500 km 2), number of locations (less than five), and an inferred continuing decline of the quality of the habitat.</p></div>	https://treatment.plazi.org/id/03E787B2241DFF9B8AE1CFC91D3413CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	do Carmo, João A. M.;Salas, Roberto M.;Cabral, Elsa L.;Cabaña Fader, Andrea A.	do Carmo, João A. M., Salas, Roberto M., Cabral, Elsa L., Cabaña Fader, Andrea A. (2026): Two new species of Hexasepalum (Rubiaceae: Spermacoceae) from the ʻcampo rupestreʼ of Brazil. European Journal of Taxonomy 1075: 1-22, DOI: 10.5852/ejt.2026.1075.3302, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3302/14539
03E787B22417FF918A40CB3B1D8F1047.text	03E787B22417FF918A40CB3B1D8F1047.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexasepalum motaviananum Cabana Fader, J. A. M. Carmo & R. M. Salas 2026	<div><p>Hexasepalum motaviananum Cabaña Fader, J.A.M.Carmo &amp; R.M.Salas sp. nov.</p><p>urn:lsid:ipni.org:names:77382789-1</p><p>Figs 5–7</p><p>Diagnosis</p><p>Hexasepalum motaviananum sp. nov. is similar to H. zappiae E.L.Cabral &amp; Cabaña Fader, but differs by the ovate leaves (vs linear in H. zappiae), glabrous fimbriae (vs scabrid margins), 6–7(8)-lobed calyx (vs 4–5-lobed), calyx lobes equal in size (vs 2 major and 2–3 minor), corolla 3.5–4 mm long, white to lilac with pink lobes (vs 7–9 mm long, white), stigma with lamellate lobes (vs with globose lobes), ventral surface of the seed with a Y-shaped groove, delimited by an apical fold, without apical grooves (vs without a Y-shaped groove nor apical fold, with 3–5 apical grooves).</p><p>Etymology</p><p>The specific epithet refers to two esteemed botanists from Minas Gerais. The initial segment, ʻmotaʼ, refers to Nara Mota, a Xyridaceae specialist, while the latter part, ʻviananumʼ, is in recognition of Pedro Lage Viana, Bambusoideae ( Poaceae) specialist. With a dedication spanning over a decade, their exploration of the ʻParque Estadual do Rio Pretoʼ culminated in an estimated assemblage of more than 1600 species thriving within this conservation bastion, underscoring its pivotal significance for conservation efforts. Furthermore, they collected the type specimen of the new species described here.</p><p>Material examined</p><p>Type</p><p>BRAZIL – Minas Gerais • São Gonçalo do Rio Preto, Parque Estadual do Rio Preto; Mota &amp; Viana 2661; holotype: BHCB; isotypes: CTES, SI .</p><p>Paratypes</p><p>BRAZIL – Minas Gerais • São Gonçalo do Rio Preto, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.356388&amp;materialsCitation.latitude=-18.083889" title="Search Plazi for locations around (long -43.356388/lat -18.083889)">Parque Estadual do Rio Preto</a>; 18°05′02″ S, 43°21′23″ W; 5 Jul. 2014; Carmo, Scatigna &amp; Piassa 212; UEC [UEC 181477] • São Gonçalo do Rio Preto, Parque Estadual do Rio Preto, Areial, próximo à portaria do parque; 8 Oct. 2006; Marino et al. 86; BHCB [BHCB 105413] • São Gonçalo do Rio Preto, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.35583&amp;materialsCitation.latitude=-18.080278" title="Search Plazi for locations around (long -43.35583/lat -18.080278)">Trilha para a lapa do Crescêncio, área após a cerca do Parque Estadual do Rio Preto</a>; 18°04′49″ S, 43°21′21″ W; 11 Jan. 2015; Carmo &amp; Scatigna 279; UEC [UEC 181475] .</p><p>Description</p><p>Subshrubs, 5–30 cm tall, much-branched, underground branches covered with foliar remains, well-developed main root, up to 1.5 m deep. Stems erect or decumbent, internodes 1–3 mm long, subcylindrical to tetragonal, glabrous or glabrescent. Stipular sheath 1–1.2 mm long, slightly exceeding the insertion of the pair of leaves, glabrous, 3–5-fimbriate, fimbriae 0.8–1.5 mm long, narrowly triangular, slightly succulent, glabrous. Leaves 3–6 × 0.8–1.1 mm, strongly overlapping, ovate, apex acuminate, base truncate, margins slightly revolute, with an epidermal thickening, slightly succulent, glabrous, inconspicuous secondary veins. Inflorescences axillary, 3–5 per flowering branch, fascicles 2–5-flowered, bracts 2, foliaceous. Flowers sessile to subsessile; hypanthium obovate, hexagonal in section; calyx 6–7(8)-lobed, tube absent, lobes 1–1.4 mm long, narrowly triangular, glabrous, margin scabrous, with glandular intercalary teeth; corolla 3.5–4 mm long, 4(5)-lobed, infundibuliform, white to lilac with pink lobes, externally glabrous, internally with a ring of moniliform trichomes near the base of the tube; stamens exserted, filament 0.4–0.6 mm, anthers shorter than the lobes; pollen grains 11–12-zonocolporate, suboblate to oblate-spheroidal, endoaperture forming an endocingulum, spines and small granules irregularly distributed over the grain; style 4–4.5 mm long, stigma bilobate, lobes lamellate; nectariferous disk entire. Schizocarps 2.8–3 mm long, obovoid, pubescent, mericarps with dorsal face convex, conspicuously 3-carinate, ventral face with 2 depressions on both sides of the midline. Seeds 1.3–1.4 mm long, obovate in outline, dorsal face convex, without grooves, ventral face with a Y-shaped groove delimited by an apical fold and the longitudinal groove, forked apical grooves not reaching the dorsal face; testa reticulate-foveate, cells isodiametric or elongated; embryo with curved apex.</p><p>Distribution, habitat, and phenology</p><p>Hexasepalum motaviananum sp. nov. is endemic to Minas Gerais, Brazil. It has been recorded in the ʻParque Estadual do Rio Pretoʼ and its surroundings, in the municipality of São Gonçalo do Rio Preto (Fig. 4). This species grows on white sand soils surrounded by rocky outcrops in ʻcampo rupestreʼ vegetation (Fig. 7A–B). In this environment, only the apical floriferous branches protrude from the soil, while the older branches with foliar remains are buried by the continuous deposition of sand from the erosion of the surrounding rocks (Figs 5A, 7C). This species is also one of the few with such a developed root system, whose taproot can reach down to 1.5 m deep. Specimens were collected with flowers and fruits in April.</p><p>Preliminary conservation status</p><p>Hexasepalum motaviananum sp. nov. is a rare species, known only from four herbarium specimens representing one location. This species is known to occur within the limits of the ʻParque Estadual do Rio Pretoʼ, a conservation unit of integral protection located in the municipality of São Gonçalo do Rio Preto (Fig. 4). Nevertheless, the IUCN Standards and Petitions Committee (2022) states that the criteria for the threatened categories are to be applied to a taxon, whatever the level of conservation action affecting it.</p><p>The impacts on the ʻcampo rupestreʼ, the vegetation where Hexasepalum motaviananum sp. nov. occurs, in the region of the ʻParque Estadual do Rio Pretoʼ, are usually associated with the expansion of eucalyptus plantations, as noticed by us during fieldwork.</p><p>Hexasepalum motaviananum sp. nov. has EOO and AOO equal to 0 km 2 and 4 km 2, respectively (kml file available in Supp. file 2). According to the IUCN Standards and Petitions Committee (2022), if EOO is less than AOO, it should be changed to make them equal and ensure consistency with the definition of AOO, which is an area defined within the EOO. Therefore, the one occurrence represents one location, and the species is preliminary assessed as Critically Endangered CR B1ab(iii)+B2ab(iii), based on its EOO (less than 100 km 2) and AOO (less than 10 km 2), number of locations (less than five), and an inferred continuing decline of the quality of the habitat.</p></div>	https://treatment.plazi.org/id/03E787B22417FF918A40CB3B1D8F1047	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	do Carmo, João A. M.;Salas, Roberto M.;Cabral, Elsa L.;Cabaña Fader, Andrea A.	do Carmo, João A. M., Salas, Roberto M., Cabral, Elsa L., Cabaña Fader, Andrea A. (2026): Two new species of Hexasepalum (Rubiaceae: Spermacoceae) from the ʻcampo rupestreʼ of Brazil. European Journal of Taxonomy 1075: 1-22, DOI: 10.5852/ejt.2026.1075.3302, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3302/14539
03E787B22413FF938BC6CEB81A1C163F.text	03E787B22413FF938BC6CEB81A1C163F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hexasepalum DC.	<div><p>Key to the species of Hexasepalum in the Americas</p><p>1. Plants up to 3–4 m in diam., with long creeping branches, rooting at the nodes, with short erect flowering shoots ................................................................................................................................ 2</p><p>– Plants less than 1 m in diam., erect or decumbent, rarely creeping, normally without roots at the nodes ................................................................................................................................................. 3</p><p>2. Leaf blade linear, 1.5–4 mm wide, concolorous, secondary veins inconspicuous on abaxial face; stipular sheath 4.2–7 mm long, tubular, fimbriae 5–6 mm long; flowers partially covered by the stipular sheath; calyx (4)5–6-lobed; corolla 12–18 mm long, purple or lilac; mericarps with 2 ventral depressions (as in Fig. 9A, H, N, Q) scarcely excavated on both sides of the midline; seed 3–3.1 mm long; Mexico ............................................................. Hexasepalum angustifolium (Benth.) Borhidi</p><p>– Leaf blade elliptic, 3.2–6 mm wide, discolorous, with 3–5 secondary veins visible on abaxial face; stipular sheath 2.5 mm long, with truncate edge, not tubular, fimbriae ca. 2.5 mm long; flowers not covered by the stipular sheath; calyx 4-lobed; corolla 4–7 mm long, white; ventral face of mericarps flat, without depressions (as in Fig. 9B, D, F–G, I–J, L); seed 2–2.1 mm long; Central America and tropical Africa .................................... Hexasepalum serrulatum (P.Beauv.) J.H.Kirkbr. &amp; Delprete</p><p>3. Ventral face of mericarps with two depressions, one on each side of the midline (Fig. 9A, H, N, Q) ........................................................................................................................................................... 4</p><p>– Ventral face of mericarps flat, without depressions (Fig. 9B, D, F–G, I–J, L) ................................. 8</p><p>4. Subshrubs up to 30 cm tall, with well-developed taproots, much-branched below and above the substrate ............................................................................................................................................ 5</p><p>– Annual herbs, subshrubs or shrubs up to 1 m tall, slightly branched above the substrate ............... 6</p><p>5. Leaf blade ovate; fimbriae glabrous; calyx 6–7(8)-lobed, lobes equal in size; corolla 3.5–4 mm long, white to lilac with pink lobes; stigma with lamellate lobes; ventral surface of the seed with a Y-shaped groove, delimited by an apical fold, without apical grooves; Minas Gerais ...................................... ........................... Hexasepalum motaviananum Cabaña Fader, J.A.M.Carmo &amp; R.M.Salas sp. nov.</p><p>– Leaf blade linear; fimbriae with scabrid margins; calyx 4–5-lobed, with 2 major and 2–3 minor lobes; corolla 7–9 mm long, white; stigma with globose lobes; ventral surface of the seed without a Y-shaped groove nor apical fold, with 3–5 apical grooves; Bahia ..................................................................... ............................................................................ Hexasepalum zappiae E.L.Cabral &amp; Cabaña Fader</p><p>6. Fruits dorsally smooth, mericarps with two slight ventral depressions; ventral surface of seed with an apical invagination surpassing half of the seed length, with a T-shaped groove; Dominican Republic ............................................ Hexasepalum domingense (DC.) Cabaña Fader &amp; R.M.Salas</p><p>– Fruits dorsally 1–5-carinate (Fig. 8B, H), mericarps with two prominent ventral depressions (Fig. 9A, N–O); ventral surface of seed with an apical invagination equal to ⅓ of seed length, with Y-shaped groove (Figs 10B–C, 11I); America and elsewhere .......................................................................... 7</p><p>7. Annual plants, rarely woody at the base; calyx lobes usually equal; corolla 3–7 mm long ............... ............................................................................................. Hexasepalum teres (Walter) J.H.Kirkbr.</p><p>– Perennial, suffruticose plants, sometimes shrubs; calyx usually with two large and two short lobes; corolla 7–20 mm long; Americas, introduced in Africa and Asia ....................................................... ................................................................ Hexasepalum apiculatum (Willd.) Delprete &amp; J.H.Kirkbr.</p><p>8. Leaf blade linear or narrowly elliptic, secondary veins inconspicuous ............................................ 9</p><p>– Leaf blade obovate, ovate, elliptic or lanceolate, 3–5 secondary veins, plicate-nervose visible on both faces or veins visible only on the abaxial face ............................................................................... 10</p><p>9. Plants scandent with graceful stems, leaf blade linear, puberulous, chartaceous; stipular sheath with 3–5 fimbriae, fimbriae glabrous; corolla 5–6 mm long; Dominican Republic ................................... ....................................................................... Hexasepalum scandens (Sw.) J.H.Kirkbr. &amp; Delprete</p><p>– Erect subshrub with robust stems, leaf blade narrowly elliptic, glabrous, fleshy, with thickened margin, coriaceous when dry; stipular sheath with 5–7 fimbriae, fimbriae pilose; corolla 7–8 mm long ....... .......................................................... Hexasepalum mello-barretoi (Standl.) J.H.Kirkbr. &amp; Delprete</p><p>10. Calyx lobes 5–9 mm long; seed dorsally ruminate (Fig. 10G) ........................................................... .............................................................. Hexasepalum gardneri (K.Schum.) J.H.Kirkbr. &amp; Delprete</p><p>– Calyx lobes 0.8–3 mm long; seed dorsally smooth (Fig. 11F) or with 1–2 apical furrows extending from the ventral face (Figs 10D–G, 11C–D), not ruminate .............................................................11</p><p>11. Fruit width equal to or greater than its length (Fig. 8C, F), seed subtriangular or triangular in outline, dorsally with 1–2 apical furrows (Figs 10D–G, 11C–D) ................................................................ 12</p><p>– Fruit width shorter than its length, seed obovate or ellipsoidal in outline, dorsally smooth .......... 13</p><p>12. Leaves plicato-nervose, scabridulous, papery, not succulent, stipular sheath 6–12-fimbriate, fimbriae pilose, dorsal face of the mericarps smooth (Fig. 9K), pollen grains 9–11-zonocolporate, exine covered by microspines, fruits pilose .......................... Hexasepalum radula (Willd.) Delprete &amp; J.H.Kirkbr.</p><p>– Leaves non plicato-nervose, glabrous, succulent, coriaceous when dry, stipular sheath 3–5-fimbriate, fimbriae glabrous, dorsal face of the mericarps with 3–4 soft longitudinal grooves (Fig. 9C), pollen grains 19–20-zonocolporate, exine covered by spines and small granules, fruit glabrous ................. ..................................................... Hexasepalum bacigalupoae Cabaña Fader &amp; E.L.Cabral sp. nov.</p><p>13. Subshrub scandent up to 2–10 m long; leaves 25–50 mm long; stipular sheath with 7–11 fimbriae, fimbriae 2–8 mm long; bracts elliptic, pseudopetiolate; ventral face of seed with short apical furrows (Fig. 11E); tropical regions of America, Africa, and Asia .................................................................. ................................................................ Hexasepalum sarmentosum (Sw.) Delprete &amp; J.H.Kirkbr.</p><p>– Subshrub erect 0.5–0.8 m or with scandent branches up to 1 m tall; leaves 10–24 mm long; stipular sheath with 3–5 fimbriae, fimbriae 0.6–1.5 mm long; bracts cordate, sessile; ventral face of seed with 2–5 apical furrows that do not reach the dorsal face (Fig. 11B, with 2 furrows); Piauí and Ceará, Brazil .......................................................... Hexasepalum nordestinum Cabaña Fader &amp; E.B.Souza</p></div>	https://treatment.plazi.org/id/03E787B22413FF938BC6CEB81A1C163F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	do Carmo, João A. M.;Salas, Roberto M.;Cabral, Elsa L.;Cabaña Fader, Andrea A.	do Carmo, João A. M., Salas, Roberto M., Cabral, Elsa L., Cabaña Fader, Andrea A. (2026): Two new species of Hexasepalum (Rubiaceae: Spermacoceae) from the ʻcampo rupestreʼ of Brazil. European Journal of Taxonomy 1075: 1-22, DOI: 10.5852/ejt.2026.1075.3302, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3302/14539
