taxonID	type	description	language	source
03DB2A41FFA5FF96FD9A24BB95F0FB04.taxon	materials_examined	Specimens collected. P 5 d: 1 ♂ (RER 23 _ LES 2; RER), 21 vi 2023; 2 ♂♂, RER 23 _ LES 3 – 4; BRIN), 21 vi 2023; 1 ♂ (RER 23 _ LES 5; UMRI), 21 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFA5FF96FD9A24BB95F0FB04.taxon	discussion	Remarks. Lestes decipiens is generally a locally occurring taxon, and this was certainly the case in the Kampar Peninsula where it was only found at one pond at the base of the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFA5FF96FDB7225F97BCF8E2.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFA5FF96FDB7225F97BCF8E2.taxon	materials_examined	Specimens collected. M 1 b: 1 ♀ (RER 20 _ LES 4; RER), 22 i 2020; M 1 e: 1 ♀ (RER 20 _ LES 5; BRIN), teneral, 26 i 2020; M 4 a: 1 ♂ (RER 20 _ LES 1; RER), 14 i 2020; 1 ♂ (RER 20 _ LES 2; BRIN), 14 i 2020; 1 ♂ (RER 20 _ LES 3; UMRI), 14 i 2020; M 4 b: 1 ♂ (RER 23 _ LES 1; UMRI )), 8 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFA5FF96FDB7225F97BCF8E2.taxon	discussion	Remarks. All individuals collected in the Kampar Peninsula were found in PSF. In the atypically dry conditions of January 2020 a number of males were found along the long trail in PSF at location M 4 despite the almost total lack of water at the time.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFA5FF89FD8B20F39753FDB1.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFA5FF89FD8B20F39753FDB1.taxon	materials_examined	Specimens collected. M 1 a: 1 ♂ (RER 20 _ EUP 1; RER), 19 i 2020; M 1 f: 1 ♂ (RER 23 _ EUP 1; BRIN), 16 ii 2023; 1 ♂ (RER 23 _ EUP 3; RER), 1 x 2023; 1 ♂ (RER 23 _ EUP 4; BRIN), 1 x 2023; M 1 l: 1 ♂ (RER 23 _ EUP 2; UMRI), 17 ii 2023; 1 ♂ (RER 23 _ EUP 5; UMRI), 4 x 2023; 1 ♀ (RER 23 _ EUP 6; RER), 4 x 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFA5FF89FD8B20F39753FDB1.taxon	discussion	Remarks. Dysphaea dimidiata is a forest stream species and the most generalist of its genus in Sundaland. In the Kampar Peninsula it was mostly found on Simpang Sianyir and close-to-closed canopy sections of Simpang Kanan, with a single individual found on Sungai Serkap itself, in a section with trees to the water’s edge. Although the Euphaeidae are largely absent from low pH waters, D. dimidiata seems capable of thriving in such waters, Dow & Reels (2024) report it in good numbers from the closed canopy part of Sungai Maludam in Maludam National Park, Sarawak.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBAFF88FDA12419959DFD60.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBAFF88FDA12419959DFD60.taxon	materials_examined	Specimens collected. M 1 e: 1 ♂ (RER 20 _ ARG 3; UMRI), 26 i 2020; M 1 f: 1 ♂ (RER 20 _ ARG 1; RER), in PSF near stream, 19 i 2020; M 1 h: 1 ♂ (RER 20 _ ARG 2; BRIN), 19 i 2020; M 1 k: 1 ♀ (RER 23 _ ARG 45; UMRI), 2 x 2023; M 1 n: 1 ♂ (RER 23 _ ARG 14; BRIN), 16 ii 2023; M 1 o: 1 ♂ (RER 23 _ ARG 43; RER), 1 x 2023; M 2 a: 1 ♂ (RER 23 _ ARG 44; BRIN), 2 x 2023; M 4 a: 2 ♂♂ (RER 23 _ ARG 21,24; UMRI), 5 vi 2023; 1 ♂ (RER 23 _ ARG 22; RER), 5 vi 2023; 1 ♂ (RER 23 _ ARG 23; BRIN), 5 vi 2023; 1 ♂, 1 ♀ (RER 23 _ ARG 35 – 36; BRIN, UMRI), 14 vi 2023; 2 ♂♂ (RER 23 _ ARG 38, 41; BRIN), 27 ix 2023; 1 ♂ (RER 23 _ ARG 39; UMRI), 27 ix 2023; 1 ♂ (RER 23 _ ARG 40; RER), 27 ix 2023; M 4 b: 1 ♂ (RER 23 _ ARG 5; BRIN), 8 ii 2023; 1 ♂ (RER 23 _ ARG 6; UMRI), 8 ii 2023; 2 ♂♂ (RER 23 _ ARG 10,13; RER), 13 ii 2023; 1 ♂ (RER 23 _ ARG 11; BRIN), 13 ii 2023; 1 ♂ (RER 23 _ ARG 12; UMRI), 13 ii 2023; 1 ♂, 1 ♀ (RER 23 _ ARG 31, 34; RER), 13 vi 2023; 1 ♂ (RER 23 _ ARG 32; BRIN), 13 vi 2023; 1 ♀ (RER 23 _ ARG 33; UMRI), 13 vi 2023; M 5 b: 1 ♂ (RER 23 _ ARG 7; RER), 9 ii 2023; 1 ♂ (RER 23 _ ARG 8; BRIN), 9 ii 2023; 1 ♂ (RER 23 _ ARG 9; UMRI), 9 ii 2023; 1 ♂ (RER _ ARG 37; RER), 17 vi 2023; M 5 e: 1 ♂ (RER 23 _ ARG 42; UMRI), 28 ix 2023; M 5 f: 1 ♂ (RER 23 _ ARG 25; RER), 6 vi 2023; M 10 a: 1 ♂, 1 ♀ (RER 23 _ ARG 1,4; RER), 7 ii 2023; 1 ♂ (RER 23 _ ARG 2; BRIN), 7 ii 2023; 1 ♂ (RER 23 _ ARG 3; UMRI), 7 ii 2023; 1 ♂ (RER 23 _ ARG 26; BRIN), 8 vi 2023; 1 ♂ (RER 23 _ ARG 27; UMRI), 8 vi 2023; 1 ♂ (RER 23 _ ARG 28; RER), 8 vi 2023; M 11 b: 1 ♂ (RER 23 _ ARG 29; BRIN), 11 vi 2023; 1 ♂ (RER 23 _ ARG 30; UMRI), 11 vi 2023; P 1 b: 2 ♂♂ (RER 23 _ ARG 15,18; UMRI), 31 v 2023; 1 ♂, 1 ♀ (RER 23 _ ARG 16, 19; RER), 31 v 2023; 1 ♂ (RER 23 _ ARG 17; BRIN), 31 v 2023; 1 ♂ (RER 23 _ ARG 20; BRIN), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBAFF88FDA12419959DFD60.taxon	discussion	Taxonomic remarks. Although described from a single male specimen from Riau Province (Lieftinck 1940 b), P. buwaldai had not been recorded from Sumatra again until now. It had remained known only from the holotype until Hämäläinen (2000) recorded it from Sabak Bernam in Selangor, Malaysia and all later records have been from Peninsular Malaysia. The Bornean P. chrysopus Selys, 1889 is closely allied with P. buwaldai, differing significantly only in markings of the thorax. Lieftinck (1940 b) states that the two species also differ “ very strikingly ” in the colour of the legs but this is not correct (an error arising from limited material and state of preservation of said material), in both species mature males develop luridly red legs, but this colour fades on preservation to a dull red or the “ salmon-orange ” described by Lieftinck (1940 b). Other remarks. Where the habitat has been recorded P. buwaldai has been found in swamp forest, in most cases (Dow et al. 2012, Choong 2013, Choong & Cheah 2013) explicitly PSF. Kalkman (2004: 26) recorded the species from Tasik Chini in Pahang, at “ a small, almost stagnant runnel, in a small piece of secondary swamp forest. ” It is not clear if this was PSF or freshwater swamp forest but PSF is (or was) present around Tasik Chini. Lieftinck (1940 b) does not state the habitat in which the holotype was collected in Riau. Hämäläinen (2000: 54) lists the species from a “ black-water stream ” in the Bernam swamp forest in Selangor; it can reasonably be assumed that this habitat was actually in PSF. The record in Choong, Orr & Dow (2008) is based on old student collections from the Universiti Kebangsaan Malaysia (UKM) campus or its vicinity in Selangor, in the UKM collection and without any information on habitat, but pockets of low pH swamp forest were likely present on or in the vicinity of the campus in the past but have since been lost to development. All records from our study in Riau are from PSF. It appears that P. buwaldai is a specialist species of low pH swamp forest, mainly PSF. In the Kampar Peninsula all individuals found were in the forest except one individual found close to its edge in a semi-open area. The species was generally abundant during all survey periods except January 2020 (when it was still present around Sungai Serkap at least), with far more individuals seen than were collected.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBBFF88FDBA2295938BF900.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBBFF88FDBA2295938BF900.taxon	materials_examined	Specimens collected. M 1 f: 1 ♂ (RER 23 _ CHL 3; BRIN), 17 ii 2023; M 1 m: 2 ♂♂ (RER 23 _ CHL 1 – 2; RER), 16 ii 2023; 1 ♂ (RER 23 _ CHL 19; UMRI), 1 x 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBBFF88FDBA2295938BF900.taxon	discussion	Remarks. As with Vestalis amoena, Libellago aurantiaca was only found at two tributaries of Sungai Serkap.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBBFF88FDD724499378FB3C.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBBFF88FDD724499378FB3C.taxon	materials_examined	Specimens collected. M 1 f: 1 ♂, (RER 23 _ CAL 3; UMRI), 17 ii 2023; 1 ♀ (RER 23 _ CAL 4; RER), 17 ii 2023; M 1 m: 1 ♂ (RER 23 _ CAL 1; RER), 16 ii 2023; 1 ♂ (RER 23 _ CAL 2; BRIN), 16 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBBFF88FDD724499378FB3C.taxon	discussion	Remarks. Like the Euphaeidae the Calopterygidae are not well represented in low pH waters but, like D. dimidiata, V. amoena can use them. In the Kampar Peninsula it was found only on two tributaries of Sungai Serkap. Curiously, it was only found in February 2023 with no individuals even seen at the same sites later in the year.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBBFF8BFD8F20199721FD4D.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBBFF8BFD8F20199721FD4D.taxon	materials_examined	Specimens collected. M 1 a: 2 ♂♂ (RER 20 _ CHL 2 – 3; BRIN), 15 i 2020; 1 ♂ (RER 23 _ CHL 14; UMRI), 14 vi 2023; M 1 b: 1 ♀ (RER 20 _ CHL 9; UMRI), 22 i 2020; M 1 d: 1 ♂ (RER 20 _ CHL 1; RER), 15 i 2020; M 1 e: 1 ♂ (RER 20 _ CHL 12; BRIN), 26 i 2020; M 1 f: 1 ♂ (RER 20 _ CHL 6; RER), 19 i 2020; 1 ♂ (RER 23 _ CHL 9; RER), 17 ii 2023; 1 ♂ (RER 23 _ CHL 25; UMRI), 4 x 2023; M 1 l: 1 ♂ (RER 23 _ CHL 8; UMRI), 16 i 2023; M 1 m: 1 ♂ (RER 23 _ CHL 7; BRIN), 16 ii 2023; 1 ♂ (RER 23 _ CHL 22; UMRI), 1 x 2023, coll. RER staff; 1 ♀ (RER 23 _ CHL 24; BRIN), 1 x 2023, coll. RER staff; 1 ♂ (RER 23 _ CHL 23; RER), 1 x 2023; M 2 a: 2 ♂♂ (RER 20 _ CHL 4 – 5; UMRI), 16 i 2020; M 3 b: 1 ♀ (RER 23 _ CHL 6; RER), 10 ii 2023; M 5 a: 2 ♂♂ (RER 20 _ CHL 7 – 8; BRIN), 20 i 2020; 2 ♂♂ (RER 20 _ CHL 10 – 11; RER), 25 i 2020; 2 ♂♂ (RER 23 _ CHL 4 – 5; UMRI), 9 ii 2023; M 5 f: 1 ♂ (RER 23 _ CHL 13; BRIN), 6 vi 2023; M 5 g: 1 ♂ (RER 23 _ CHL 20; RER), 28 ix 2023; M 7 e: 1 ♂ (RER 23 _ CHL 10; BRIN), 19 ii 2023; M 13 b: 1 ♀ (RER 23 _ CHL 15; RER), 18 vi 2023; 1 ♂ (RER 23 _ CHL 21; BRIN), 29 ix 2023; P 1 c: 1 ♂ (RER 23 _ CHL 12; RER), 2 vi 2023; P 2 a: 1 ♂ (RER 23 _ CHL 11; UMRI), 30 v 2023; P 5 c: 1 ♂ (RER 23 _ CHL 16; BRIN), 21 vi 2023; 1 ♀ (RER 23 _ CHL 17; UMRI), 21 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBBFF8BFD8F20199721FD4D.taxon	discussion	Remarks. Libellago hyalina thrives in low pH waters, although it is not confined to them. Generally abundant in the main part of the Kampar Peninsula and also present in the mineral to peat soils transition area at the base of the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB8FF8BFDA222249799F998.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB8FF8BFDA222249799F998.taxon	materials_examined	Specimens collected. P 5 c: 2 ♂♂, 1 ♀ (RER 23 _ PCD 49 – 51; RER), 21 vi 2023; P 6 d: 1 ♂ (RER 23 _ PCD 84; RER), 23 ix 2023; P 6 e: 1 ♂, 1 ♀ (RER 23 _ PCD 86 – 87; BRIN), in tandem, 25 ix 2023; P 6 f: 1 ♂ (RER 23 _ PCD 85; UMRI), 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB8FF8BFDA222249799F998.taxon	discussion	Remarks. Not found on the main, peat soil part of the Kampar Peninsula but occurring on mineral soils at the base of the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB8FF8BFDAF25D59706FBCC.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB8FF8BFDAF25D59706FBCC.taxon	materials_examined	Specimens collected. M 13 b: 1 ♂ (RER 23 _ CHL 18; RER), 18 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB8FF8BFDAF25D59706FBCC.taxon	discussion	Remarks. Recorded from Riau for the first time, a single, immature male in a shaded section of a small canal. Although small, this is a conspicuous species and it is evidently not common in the PSF habitats of the Kampar Peninsular.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB8FF8AFDBA21E0939CFC2C.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB8FF8AFDBA21E0939CFC2C.taxon	materials_examined	Specimens collected. M 1 b: 1 ♂ (RER 20 _ PCD 5; UMRI), 18 i 2020; 3 ♀♀ (RER 20 _ PCD 6 – 8; RER), 18 i 2020; 1 ♂, 1 ♀ (RER 20 _ PCD 9 – 10; BRIN), in tandem, 22 i 2020; M 1 d: 1 ♀ (RER 20 _ PCD 1; RER), 15 i 2020; M 1 e: 1 ♂ (RER 20 _ PCD 11; UMRI), 26 i 2020; M 2 a: 3 ♂♂ (RER 20 _ PCD 2 – 4; BRIN), 16 i 2020; 1 ♂ (RER 23 _ PCD 93; UMRI), 2 x 2023; M 4 a: 1 ♂, 1 ♀ (RER 23 _ PCD 8 – 9; BRIN), in tandem, 13 ii 2023; 2 ♂♂ (RER 23 _ PCD 54 – 55; UMRI), 5 vi 2023; 1 ♂, 1 ♀ (RER 23 _ PCD 63 – 64; RER), 14 vi 2023; 1 ♂, 1 ♀ (RER 23 _ PCD 89 – 90; UMRI), tandem pair, 27 ix 2023; 1 ♂ (RER 23 _ PCD 92; BRIN), 27 ix 2023; M 4 b: 2 ♂♂ (RER 23 _ PCD 2 – 3; BRIN), 8 ii 2023; 2 ♂♂ (RER 23 _ PCD 10 – 11; UMRI), 13 ii 2023; 1 ♀ (RER 23 _ PCD 12; RER), teneral, 13 ii 2023, coll. RER staff; 1 ♂, 1 ♀ (RER 23 _ PCD 58 – 59; BRIN), tandem pair, 13 vi 2023; 3 ♂♂ (RER 23 _ PCD 60 – 62; UMRI), 13 vi 2023; M 4 e: 1 ♀ (RER 23 _ PCD 7; RER), 13 ii 2023; 1 ♂ (RER 23 _ PCD 91; RER), 27 ix 2023; M 5 b: 2 ♂♂, 1 ♀ (RER 23 _ PCD 4 – 6; UMRI), 9 ii 2023; M 5 f: 1 ♂, 1 ♀ (RER 23 _ PCD 56 – 57; RER), tandem pair, 6 vi 2023; M 10 a: 1 ♂ (RER 23 _ PCD 1; RER), 7 ii 2023; M 12 d: 1 ♂ (RER 23 _ PCD 65; BRIN), 18 vi 2023; P 1 b: 1 ♂. 1 ♀ (RER 23 _ PCD 52 – 53; BRIN), 2 vi 2023; P 6 f: 1 ♂ (RER 23 _ PCD 88; RER), 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB8FF8AFDBA21E0939CFC2C.taxon	discussion	Taxonomic remarks. The variety of color forms (and general variability) of C. vittata has been commented on before (for example see Dow & Singa (2022) on forms found in Sarawak and Brunei) but no detailed study of C. vittata and its closest allies has been made since that of Lieftinck (1940 a). Two color forms were found at the Kampar Peninsula: (a) a largely dark form with yellow legs and a very variable (sometimes very reduced) pattern of pale markings on the synthorax, this form is very common in the low pH habitats of the main part of the peninsula; (b) the ‘ typical’ red-legged form, only a single individual of this form was collected, at a small stream in oil palm on mineral soils at the base of the peninsula. In Borneo typically a single form is found in a given area, for instance a yellow-legged form is found in the southwest of Sarawak (where it occurs in a variety of habitats including PSF), with an exception in Brunei and Limbang Division in northeast Sarawak where the ‘ typical’ red-legged form and a black form (Orr 2003, Dow & Singa 2022) both occur, but without clear partitioning by habitat. The situation in the Kampar Peninsula contrasts with that in Borneo with the two forms occurring in the Kampar Peninsula seemingly occupying different habitats. The yellow-legged form (which might be smaller, at least on average, than the ‘ typical’ form but detailed measurements are needed to confirm this) present in the peninsula might be a low pH phenotype but the possibility that it is a distinct taxon cannot be excluded and further work is needed on this entire complex.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB9FF8AFDAD234D97C1F860.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB9FF8AFDAD234D97C1F860.taxon	materials_examined	Specimens collected. M 1 b: 1 ♂ (RER 20 _ PCD 18; RER), 18 i 2020; 1 ♂, 1 ♀ (RER 20 _ PCD 25 – 26; BRIN), 22 i 2020; M 1 d: 3 ♂♂ (RER 20 _ PCD 12 – 14; RER), 15 i 2020; 1 ♂, 1 ♀ (RER 20 _ PCD 15 – 16; BRIN), tandem pair, 15 i 2020; M 1 e: 1 ♂ (RER 20 _ PCD 28; RER), 26 i 2020; M 1 f: 1 ♂ (RER 20 _ PCD 19; BRIN), 19 i 2020; 1 ♂, 1 ♀ (RER 23 _ PCD 22 – 23; BRIN, UMRI), tandem pair, 17 ii 2023; 1 ♂ (RER 23 _ PCD 24; UMRI), 17 ii 2023; M 1 m: 2 ♂♂ (RER 23 _ PCD 18 – 19; BRIN), 16 ii 2023; 1 ♂ (RER 23 _ PCD 97; BRIN), 1 x 2023; M 1 n: 1 ♂ (RER 23 _ PCD 20; UMRI), 16 ii 2023; 1 ♂ (RER 23 _ PCD 21; UMRI), 16 ii 2023; M 1 o: 1 ♂ (RER 23 _ PCD 98; UMRI), 1 x 2023; M 2 a: 1 ♂ (RER 20 _ PCD 17; UMRI), 16 i 2020; 1 ♂ (RER 23 _ PCD 99; RER), 2 x 2023; M 4 e: 1 ♂ (RER 23 _ PCD 17; RER), 13 ii 2023; 1 ♂ (RER 23 _ PCD 94; BRIN), 27 ix 2023; M 5 a: 3 ♂♂ (RER 20 _ PCD 20 – 22; UMRI), 20 i 2020; 2 ♂♂ (RER 20 _ PCD 23 – 24; RER), 20 i 2020; 1 ♂ (RER 20 _ PCD 27; UMRI), 25 i 2020; 2 ♂♂ (RER 23 _ PCD 13 – 14; BRIN), 9 ii 2023; 2 ♂♂ (RER 23 _ PCD 15 – 16; UMRI), 9 ii 2023; 2 ♂♂ (RER 23 _ PCD 72 – 73; BRIN), 17 vi 2023; 1 ♂ (RER 23 _ PCD 96; RER), 28 ix 2023; M 5 b: 1 ♀ (RER 23 _ PCD 74; UMRI), 17 vi 2023; M 5 e: 1 ♀ (RER 23 _ PCD 71; RER), 6 vi 2023; M 5 f: 1 ♂, 1 ♀ (RER 23 _ PCD 69 – 70; UMRI), tandem pair, 6 vi 2023; M 5 g: 1 ♂ (RER 23 _ PCD 95; UMRI), 28 ix 2023; M 12 a: 1 ♂ (RER 23 _ PCD 75; RER), 18 vi 2023; P 1 b: 1 ♂ (RER 23 _ PCD 66; RER), 31 v 2023; P 1 c: 2 ♂♂ (RER 23 _ PCD 67 – 68; BRIN), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB9FF8AFDAD234D97C1F860.taxon	discussion	Remarks. An abundant species on streams in low pH swamp forest, where it typically spreads out into flooded forest around the stream. Although no specimen was collected actually on the lower part of Sungai Serkap (M 1 a), it was frequently seen there.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8DFDD427B9959EFDC9.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8DFDD427B9959EFDC9.taxon	materials_examined	Specimens collected. M 1 a: 1 ♀ (RER 20 _ PCD 31; BRIN), 26 i 2020; 1 ♂ (RER 23 _ PCD 76; RER), 14 vi 2023; 3 ♂♂ (RER 23 _ PCD 100 – 102; BRIN), 4 x 2023; 1 ♀ (RER 23 _ PCD 103; UMRI), 4 x 2023; M 5 a: 2 ♂♂ (RER 20 _ PCD 29 – 30; RER), 25 i 2020; 1 ♂ (RER 23 _ PCD 25; UMRI), 9 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8DFDD427B9959EFDC9.taxon	discussion	Remarks. This is the first record of E. coomansi for mainland Sumatra. A few individuals were found on Sungai Serkap and Sungai Sangar but this is a dark colored species that tends to stay in vegetation at the stream bank, so it is easily missed and probably under-recorded as a result.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8DFDBA259197D2FBD0.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8DFDBA259197D2FBD0.taxon	materials_examined	Specimens collected. M 1 k: 1 ♂, 1 ♀ (RER 23 _ PCD 106 – 107; UMRI), tandem pair, 2 x 2023; M 4 a: 1 ♂, 1 ♀ (RER 23 _ PCD 104 – 105; BRIN), tandem pair, 27 ix 2023; M 5 b: 1 ♂, 1 ♀ (RER 23 _ PCD 26 – 27; RER), tandem pair, 9 ii 2023; 1 ♂ (RER 23 _ PCD 79; UMRI), 17 vi 2023; P 1 a: 1 ♂, 1 ♀ (RER 23 _ PCD 77 – 78; BRIN), tandem pair, 31 v 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8DFDBA259197D2FBD0.taxon	discussion	Remarks. Recorded from Riau Province for the first time. Onychargia atrocyana is quite often recorded in PSF and other low pH habitats.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8DFDD623EC9247F90C.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8DFDD623EC9247F90C.taxon	materials_examined	Specimens collected. M 1 f: 2 ♂♂ (RER 23 _ PCD 37 – 38; UMRI), 17 ii 2023; M 1 m: 4 ♂♂ (RER 23 _ PCD 29 – 30, 35 – 36; BRIN), 16 ii 2023; 2 ♂♂ (RER 23 _ PCD 31 – 32; UMRI), 16 ii 2023; 2 ♂♂ (RER 23 _ PCD 33 – 34; RER), 16 ii 2023; 2 ♂♂ (RER 23 _ PCD 111, 114; RER), 1 x 2023; 1 ♂, 1 ♀ (RER 23 _ PCD 111 – 112; UMRI), tandem pair, 1 x 2023; 1 ♂ (RER 23 _ PCD 118; BRIN), 1 x 2023; M 5 a: 2 ♂♂ (RER 20 _ PCD 32 – 33; RER), 20 i 2020; 2 ♂♂ (RER 20 _ PCD 34 – 35; BRIN), 20 i 2020; 3 ♂♂ (RER 20 _ PCD 36 – 38; UMRI), 25 i 2020; 1 ♂ (RER 23 _ PCD 28; RER), 9 ii 2023; 2 ♂♂ (RER 23 _ PCD 109 – 110; UMRI), 28 ix 2023; M 5 f: 1 ♂ (RER 23 _ PCD 80; RER), 6 vi 2023; 1 ♂ (RER 23 _ PCD 114; RER), 4 x 2023; M 5 g: 1 ♂ (RER 23 _ PCD 108; BRIN), 28 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8DFDD623EC9247F90C.taxon	discussion	Remarks. This is the true P. interrupta (as opposed to the Bornean taxon that has been treated as P. interrupta, see Note 27 in Dow et al. 2024). Although not confined to low pH waters it thrives in them and is generally present on forest streams and on some stream-like sections of canal in forest in the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8CFDBA2050925BFDDE.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8CFDBA2050925BFDDE.taxon	materials_examined	Specimens collected. M 1 a: 2 ♂♂ (RER 23 _ PCD 43 – 44; RER), 12 ii 2023; M 1 f: 1 ♂ (RER 23 _ PCD 47; RER), 17 ii 2023; M 1 r: 1 ♂ (RER 23 _ PCD 46; UMRI), 15 ii 2023; M 3 b: 2 ♂♂ (RER 20 _ PCD 39 – 40; RER), 17 i 2020; 1 ♂ (RER 23 _ PCD 40; BRIN), 10 ii 2023; 1 ♂, 1 ♀ (RER 23 _ PCD 41 – 42; UMRI), tandem pair, 10 ii 2023; M 3 e: 1 ♂ (RER 20 _ PCD 42; UMRI), 24 i 2020; M 4 c: 1 ♂ (RER 23 _ PCD 45; BRIN), 13 ii 2023; M 4 f: 1 ♂ (RER 23 _ PCD 48; BRIN), 19 ii 2023; M 7 c: 1 ♂ (RER 20 _ PCD 41; BRIN), 23 i 2020; M 10 c: 1 ♂ (RER 23 _ PCD 39; RER), 7 ii 2023; M 13 b: 1 ♂, 1 ♀ (RER 23 _ PCD 82 – 83; RER), 18 vi 2023; 1 ♀ (RER 23 _ PCD 117; RER), 29 ix 2023; P 3 c: 1 ♂ (RER 23 _ PCD 81; UMRI), 30 v 2023; P 5 e: 1 ♂ (RER 23 _ PCD 115; BRIN), 22 ix 2023; P 6 b: 1 ♂ (RER 23 _ PCD 116; UMRI), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBEFF8CFDBA2050925BFDDE.taxon	discussion	Remarks. Often found in vegetation around narrow channels on Sungai Serkap as well as in other habitats in the main part of the Kampar Peninsula, and also present at the base of the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBFFF8CFD812432971EFBA9.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBFFF8CFD812432971EFBA9.taxon	materials_examined	Specimens collected. P 5 d: 1 ♂ (RER 23 _ COE 167; RER), 21 vi 2023; 1 ♂ (RER 23 _ COE 168; BRIN), 21 vi 2023; 1 ♂ (RER 23 _ COE 169; UMRI), 21 vi 2023; 2 ♂♂ (RER 23 _ COE 280 – 281; BRIN), 22 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBFFF8CFD812432971EFBA9.taxon	discussion	Remark. Recorded from Riau Province for the first time, only found at one pond site at the base of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBFFF8CFDF623C997DDF90D.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBFFF8CFDF623C997DDF90D.taxon	materials_examined	Specimens collected. M 2 d: 1 ♂ (RER 20 _ COE 1; RER), 16 i 2020; M 2 e: 1 ♂ (RER 23 _ COE 1; RER), 6 ii 2023; 1 ♂ (RER 23 _ COE 2; BRIN), inside car, 9 ii 2023; M 7 a: 1 ♂, 1 ♀ (RER 20 _ COE 3 – 4; BRIN), 23 i 2020; M 7 g: 2 ♂♂ (RER 20 _ COE 5 – 6; UMRI), 23 i 2020; 3 ♂♂ (RER 23 _ COE 3 - 5; UMRI), 19 ii 2023; 1 ♂, 1 ♀ (RER 23 _ COE 6 - 7; RER), 19 ii 2023; 1 ♂ (RER 23 _ COE 170; BRIN), 13 vi 2023; P 5 a: 1 ♀ (RER 23 _ COE 171, 279; UMRI), 21 vi 2023; 1 ♀ (RER 23 _ COE 279; RER), 21 vi 2023; P 6 a: 1 ♂, 2 ♀♀ (RER 23 _ COE 282 – 284; BRIN), 22 ix 2023; P 6 b: 1 ♂ (RER 23 _ COE 286; RER), 23 ix 2023; P 6 e: 1 ♂ (RER 23 _ COE 285; UMRI), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBFFF8CFDF623C997DDF90D.taxon	discussion	Remarks. Less common (but not uncommon) than the next species on the peat soils of the main part of the Kampar Peninsula, with the relative abundancies of the two seemingly reversing at the peat-mineral soil transition at the base of the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBFFF8FFDB520159718FD61.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBFFF8FFDB520159718FD61.taxon	materials_examined	Specimens collected. M 1 c: 2 ♂♂ (RER 20 _ COE 7 – 8; RER), 15 i 2020; 2 ♀♀ (RER 20 _ COE 9 – 10; BRIN), 15 i 2020; M 1 g: 1 ♀ (RER 20 _ COE 2; RER), 15 i 2020; 1 ♂, 1 ♀ (RER 20 _ COE 11 – 12; UMRI), 15 i 2020; 1 ♀ (RER 20 _ COE 21; UMRI), 26 i 2020; M 1 i: 1 ♀ (RER 20 _ COE 13; BRIN), 15 i 2020; M 1 j: 1 ♀ (RER 23 _ COE 288; UMRI), 2 x 2023; M 2 c: 1 ♂, 1 ♀ (RER 20 _ COE 14 – 15; UMRI), 16 i 2020; 2 ♀♀ (RER 20 _ COE 16 – 17; RER), 16 i 2020; M 2 e: 1 ♂, 1 ♀ (RER 23 _ COE 8 – 9; RER), at dusk, 6 ii 2023; 1 ♂ (RER 23 _ COE 176; RER), 10 vi 2023; M 3 c: 1 ♀ (RER 23 _ COE 11; UMRI), 10 ii 2023; M 4 d: 1 ♂ (RER 23 _ COE 178; UMRI), perched on surface of dirt road, 14 vi 2023; M 5 c: 1 ♂ (RER 23 _ COE 10, BRIN), 9 ii 2023; M 5 d: 1 ♂ (RER 20 _ COE 20; BRIN), 23 i 2020; M 6: 1 ♀ (RER 20 _ COE 19; RER), 23 i 2020; M 7 a: 1 ♀ (RER 20 _ COE 166; BRIN), 23 i 2020; 2 ♀♀ (RER 23 _ COE 13 – 14; BRIN), 19 ii 2023; M 7 b: 1 ♀ (RER 20 _ COE 18; UMRI), 23 i 2020; M 7 d: 1 ♂ (RER 23 _ COE 12; RER), 19 ii 2023; M 7 g: 1 ♀ (RER 23 _ COE 15; UMRI), 19 ii 2023; M 11 a: 1 ♀ (RER 23 _ COE 177; BRIN), 11 vi 2023; P 1 a: 1 ♀ (RER 23 _ COE 287; BRIN), 24 ix 2023; P 3 a: 1 ♂, 1 ♀ (RER 23 _ COE 172 – 173; RER), 30 v 2023; P 4 a: 1 ♀ (RER 23 _ COE 175; UMRI), 2 vi 2023; P 4 c: 1 ♂ (RER 23 _ COE 174; BRIN), 30 v 2023; P 5 d: 1 ♀ (RER 23 _ COE 179; RER), 21 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBFFF8FFDB520159718FD61.taxon	discussion	Remarks. Abundant in low pH waters in the main part of the Kampar Peninsula, less so at the base of the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBCFF8FFDD025E1936FFBFD.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBCFF8FFDD025E1936FFBFD.taxon	materials_examined	Specimens collected. M 7 a: 1 ♀ (no reference code assigned; UMRI), 23 i 2020, leg. S. O. Zulya.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBCFF8FFDD025E1936FFBFD.taxon	discussion	Remarks. Only a single female was collected during the surveys in the Kampar Peninsula but A. pygmaea is likely to be more common than this single record suggests, grassy banks around drains (the habitat where the single specimen was collected) were not sampled much during the surveys but are good places to look for this species.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBCFF8FFDE523899451F8CC.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBCFF8FFDE523899451F8CC.taxon	materials_examined	Specimens collected. M 1 b: 5 ♂♂, 2 ♀♀ (RER 20 _ COE 22 – 28; RER), 18 i 2020; 7 ♂♂, 4 ♀♀ (RER 20 _ COE 29 – 39; RER), 22 i 2020; M 1 e: 1 ♂, 1 ♀ (RER 20 _ COE 40 – 41; RER), 26 i 2020; M 1 o: 1 ♀ (RER 23 _ COE 290; RER), 1 x 2023; M 4 a: 1 ♂ (RER 23 _ COE 33; RER), 13 ii 2023; M 4 b: 1 ♂ (RER 23 _ COE 34; RER), 13 ii 2023, coll. RER staff; M 5 b: 8 ♂♂, 5 ♀♀ (RER 23 _ COE 23 – 32, 45 – 46, 148; RER), 9 ii 2023; 2 ♂♂ (RER 23 _ COE 192 – 193; RER), 17 vi 2023; M 10 a: 6 ♂♂, 2 ♀♀ (RER 23 _ COE 16 – 22, 147; RER), 7 ii 2023; 3 ♂♂, 1 ♀ (RER 23 _ COE 183 – 186; RER), 8 vi 2023; M 11 b: 1 ♂, 4 ♀♀ (RER 23 _ COE 187 – 191; RER), 11 vi 2023; P 1 b: 1 ♂, 2 ♀♀ (RER 23 _ COE 180 – 182; RER), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBCFF8FFDE523899451F8CC.taxon	discussion	Remarks. This species had only been known from Peninsula Malaysia and Singapore until now. Moderately common in many parts of the PSF in the Kampar Peninsular when and where surface water is present. All specimens were retained in the RER collection pending the publication of Dow et al. (2025) and will be divided between BRIN, RER and UMRI at a later date.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBCFF8EFDEF205C92E5FD4C.taxon	materials_examined	Specimens collected. M 1 b: 1 ♂, 1 ♀ (RER 20 _ COE 43 – 44; RER), 18 i 2020; 9 ♂♂, 2 ♀♀ (RER 20 _ COE 45 – 55; RER), 22 i 2020; M 1 e: 8 ♂♂, 4 ♀♀ (RER 20 _ COE 56 – 67; RER), 26 i 2020; M 1 n: 1 ♂ (RER 23 _ COE 35; RER), 16 ii 2023; M 1 o: 1 ♀ (RER 23 _ COE 289; RER), 1 x 2023; M 2 a: 1 ♀ (RER 20 _ COE 42; RER), 16 i 2020.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBCFF8EFDEF205C92E5FD4C.taxon	discussion	Remarks. Until now this species was only known with certainty from the type series from Pahang. Amphicnemis hoisen might be considerably more widespread in Sumatra — the first author has seen a photograph (A. Alfarisyi personal communication 2019) of a male Amphicnemis from Bangka Island in which sufficient detail of the anal appendages can be seen to demonstrate that it is either A. hoisen or a closely allied but unnamed species. Despite the possible record from Bangka, the records from the Kampar Peninsular are the first confirmed and published records of this species from Indonesia. In the Kampar Peninsular Amphicnemis hoisen was only found in flooded PSF adjacent to the Serkap River and its tributary Simpang Sianyir, however at the type locality in Pahang it was found at pools in otherwise unflooded PSF apparently distant from any stream or river. All specimens were retained in the RER collection pending the publication of Dow et al. (2025) and will be divided between BRIN, RER and UMRI at a later date.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBDFF8EFE5925D595DAFB3B.taxon	materials_examined	Specimens collected. M 1 b: 1 ♂ (RER 20 _ COE 68; RER), 18 i 2020; M 1 n: 6 ♂♂ (RER 23 _ COE 47 – 52; RER), 16 ii 2023; M 4 b: 1 ♂, 1 ♀ (RER 23 _ COE 40 – 41; RER), 8 ii 2023; 1 ♂ (RER 23 _ COE 149; RER), 13 ii 2023; 1 ♀ (RER 23 _ COE 195; RER), 13 vi 2023; M 5 b: 3 ♂♂ (RER 23 _ COE 42 – 44; RER), 9 ii 2023; M 10 a: 3 ♂♂, 1 ♀ (RER 23 _ COE 36 – 39; RER), 7 ii 2023; 1 ♀ (RER 23 _ COE 194; RER), 8 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBDFF8EFE5925D595DAFB3B.taxon	discussion	Remarks. A clearly distinct new species of Amphicnemis, probably most closely allied to the Bornean Amphicnemis annae Lieftinck, 1940, was discovered during the surveys in the Kampar Peninsula and described as A. reri by Dow et al. (2025). In January 2020 only a single male was collected, in flooded PSF adjacent to the Serkap River. More individuals, of both sexes, were found at multiple sites in PSF in February 2023. No males of this species were found during surveys later in 2023, only a few females.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBDFF8EFDA522589361F942.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBDFF8EFDA522589361F942.taxon	materials_examined	Specimens collected. M 13 b: 1 ♂ (RER 23 _ COE 13; BRIN), 18 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBDFF8EFDA522589361F942.taxon	discussion	Remarks. A single male was found and collected on a blackwater canal flowing directly into the Kampar River. The habitat of this species is difficult to characterize, it is seldom abundant and generally local in occurrence, usually found at forest streams, often with low pH but equally often without and most often in disturbed forest.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBDFF81FDD721D69487FE48.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBDFF81FDD721D69487FE48.taxon	materials_examined	Specimens collected. M 1 b: 1 ♂, 1 ♀ (RER 20 _ COE 73 – 74; UMRI), 18 i 2020; M 1 d: 3 ♂♂ (RER 20 _ COE 69 – 71; RER), 15 i 2020; M 1 f: 1 ♂ (RER 23 _ COE 65; BRIN), 17 ii 2023; 1 ♂ (RER 23 _ COE 293; BRIN), 4 x 2023; M 1 h: 1 ♂ (RER 20 _ COE 75; RER), 19 i 2020; M 1 m: 2 ♂♂ (RER 23 _ COE 57 – 58; RER), 16 ii 2023; 2 ♂♂ (RER 23 _ COE 59 – 60; BRIN), 16 ii 2023; 1 ♂, 1 ♀ (RER 23 _ COE 61 – 62; UMRI), tandem pair, 16 ii 2023; 1 ♂ (RER 23 _ COE 292; RER), 1 x 2023; M 1 n: 2 ♂♂ (RER 23 _ COE 63 – 64; RER), 16 ii 2023; M 2 a: 1 ♂ (RER 20 _ COE 72; BRIN), 16 i 2020; M 4 a: 1 ♂ (RER 23 _ COE 197; UMRI), 5 vi 2023; M 4 b: 1 ♂ (RER 23 _ COE 53; BRIN), 8 ii 2023; M 4 e: 1 ♂ (RER 23 _ COE 291; UMRI), 27 ix 2023; M 5 a: 3 ♂♂ (RER 23 _ COE 54 – 56; UMRI), 9 ii 2023; 1 ♂, 1 ♀ (RER 23 _ COE 198 – 199; RER), tandem pair, 17 vi 2023; 3 ♂♂ (RER 23 _ COE 200 – 202; BRIN), 17 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFBDFF81FDD721D69487FE48.taxon	discussion	Remarks. This is the first record for Riau Province. This species is common on streams in PSF and is also frequently found on canals and drains in PSF.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB2FF81FDB126D99333FCA0.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB2FF81FDB126D99333FCA0.taxon	materials_examined	Specimens collected. M 4 b: 1 ♂ (RER 23 _ COE 66; RER), 8 ii 2023; M 10 a: 1 ♂ (RER 23 _ COE 203; BRIN), 8 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB2FF81FDB126D99333FCA0.taxon	discussion	Remark. A widely distributed species, but not recorded particularly often. On two occasions single males were collected along trails deep within PSF, we are not aware of any records from this habitat type until now.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB2FF81FD8924A19539F98C.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB2FF81FD8924A19539F98C.taxon	materials_examined	Specimens collected. M 1 b: 1 ♀ (RER 20 _ COE 84; RER), 22 i 2020; M 1 d: 1 ♂, 1 ♀ (RER 20 _ COE 76 – 77; RER), 15 i 2020; M 1 e: 1 ♂, 1 ♀ (RER 20 _ COE 85 – 86; BRIN), tandem pair, 26 i 2020; 1 ♂ (RER 20 _ COE 87; UMRI), 26 i 2020; 2 ♂♂ (RER 20 _ COE 88 – 89; RER), 26 i 2020; M 1 f: 1 ♂ (RER 20 _ COE 81; RER), 19 i 2020; M 1 g: 1 ♂ (RER 20 _ COE 78; BRIN), 15 i 2020; M 1 h: 1 ♂ (RER 20 _ COE 82; BRIN), 19 i 2020; M 1 p: 1 ♂ (RER 23 _ COE 72; BRIN), 12 ii 2023; M 3 b: 1 ♂, 1 ♀ (RER 20 _ COE 79 – 80; UMRI), 17 i 2020; 2 ♂♂, 1 ♀ (RER 23 _ COE 69 – 71; RER), 10 ii 2023; M 4 b: 1 ♀? (RER 23 _ COE 74; RER), teneral, 13 ii 2023; 2 ♀♀? (RER 23 _ COE 75 – 76; BRIN), 13 ii 2023, leg. RER staff; M 4 c: 1 ♂ (RER 23 _ COE 67; BRIN), 8 ii 2023; M 4 e: 1 ♂ (RER 23 _ COE 73; UMRI), 13 ii 2023; 1 ♂, 1 ♀ (RER 23 _ COE 294 – 295; BRIN), 27 ix 2023; 1 ♂ (RER 23 _ COE 296; UMRI), 27 ix 2023; M 5 a: 1 ♂ (RER 20 _ COE 83; UMRI), 20 i 2020; M 5 b: 1 ♀ (RER 23 _ COE 68; UMRI), 9 ii 2023; M 7 e: 1 ♂ (RER 23 _ COE 77, UMRI), 19 ii 2023; M 13 b: 1 ♂ (RER 23 _ COE 297; RER 0, 29 ix 2023; P 1 b: 1 ♀ (RER 23 _ COE 204, RER), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB2FF81FD8924A19539F98C.taxon	discussion	Remarks. The most common of the four Archibasis species recorded in the Kampar Peninsular, except on the tributaries of Sungai Kampar where A. melanocyana dominates. Females of this genus are difficult to identify to species unless found with males and at location M 4 b no males of A. viola were found but males of two other species (A. melanocyana and A. oscillans) were recorded, leaving some room for doubt over the identity of the females found there, they are listed with a ‘? ’ above for this reason.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB2FF80FE1621D89262FE34.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB2FF80FE1621D89262FE34.taxon	materials_examined	Specimens collected. M 1 i: 3 ♂♂ (RER 20 _ COE 90 – 92; RER), 15 i 2020; M 2 b: 1 ♂ (RER 20 _ COE 93; BRIN), 16 i 2020; M 2 c: 2 ♂♂ (RER 20 _ COE 94 – 95; UMRI), 16 i 2020; M 3 b: 1 ♂, 1 ♀ (RER 20 _ COE 96 – 97; RER), 17 i 2020; 1 ♂, 1 ♀ (RER 23 _ COE 78 – 79; BRIN), tandem pair, 10 ii 2023; 3 ♂♂ (RER 23 _ COE 81 – 83; RER), 10 ii 2023; M 3 c: 1 ♂ (RER 20 _ COE 126; BRIN), 17 i 2020, leg. Shinta Okka Zulya; 1 ♂ (RER 23 _ COE 80; UMRI), 10 ii 2023; 1 ♀ (RER 23 _ COE 84; BRIN), 10 ii 2023, leg. RER staff; M 7 a: 2 ♂♂ (RER 20 _ COE 98 – 99; UMRI), 23 i 2020; M 7 c: 2 ♂♂ (RER 20 _ COE 100 – 101; RER), 23 i 2020; M 7 g: 1 ♂, 1 ♀ (RER 23 _ COE 85, 145; UMRI), 19 ii 2023; M 13 d: 1 ♂ (RER 23 _ COE 301; BRIN), 29 ix 2023; P 3 d: 1 ♂ (RER 23 _ COEE 205; RER), 30 v 2023; P 4 b: 1 ♂ (RER 23 _ COE 206; BRIN), 31 v 2023; P 5 a: 1 ♂ (RER 23 _ COE 207; UMRI), 21 vi 2023; 1 ♂ (RER 23 _ COE 298; RER), 22 ix 2023; P 6 c: 1 ♂ (RER 23 _ COE 299; BRIN), 23 ix 2023; P 6 f: 1 ♂ (RER 23 _ COE 300; UMRI), 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB2FF80FE1621D89262FE34.taxon	discussion	Remarks. Moderately common throughout the Kampar Peninsula. The male collected at location P 3 d was in the male position in a same sex tandem with Ceriagrion cerinorubellum (Brauer, 1865).	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB3FF80FDCD250C9594FB18.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB3FF80FDCD250C9594FB18.taxon	materials_examined	Specimens collected. M 1 c: 1 ♂ (RER 20 _ COE 102; RER), 15 i 2020; M 1 h; 1 ♂ (RER 20 _ COE 107; RER), 19 i 2020; M 1 j: 1 ♂ (RER 23 _ COE 305; BRIN), 2 x 2023; M 1 p: 1 ♂ (RER 23 _ COE 89; UMRI), 12 ii 2023; M 1 r: 1 ♂ (RER 23 _ COE 92; BRIN), 15 ii 2023; M 2 a: 2 ♂♂, 1 ♀ (RER 20 _ COE 103 – 105; BRIN), 16 i 2020; M 3 c: 1 ♂ (RER 20 _ COE 106; UMRI), 17 i 2020; 1 ♂ (RER 88, BRIN), 10 ii 2023; M 3 e: 1 ♂ (RER 20 _ COE 110; RER), 24 i 2020; M 4 a: 1 ♀ (RER 23 _ COE 210; UMRI), 5 vi 2023; 1 ♂ (RER 23 _ COE 216, BRIN), 14 vi 2023; 1 ♂ (RE 23 _ COE 303; BRIN), 27 ix 2023; M 4 b: 1 ♀ (RER 23 _ COE 215; RER), 13 vi 2023; M 4 c: 1 ♂ (RER 23 _ COE 86; UMRI), 8 ii 2023; 1 ♂, 1 ♀ (RER 23 _ COE 90 – 91; RER), in tandem, 13 ii 2023; M 5 a: 1 ♂ (RER 20 _ COE 108; BRIN), 20 i 2020; 1 ♂ (RER 20 _ COE 111; BRIN), 20 i 2020; M 5 b: 1 ♂ (RER 23 _ COE 87; RER), 9 ii 2023; M 5 e: 1 ♂, 1 ♀ (RER 23 _ COE 211 – 212; BRIN), in tandem, 6 vi 2023; M 7 a: 1 ♂ (RER 20 _ COE 109; UMRI), 23 i 2020; M 11 a: 1 ♂ (RER 23 _ COE 214; BRIN), 11 vi 2023; M 11 b: 1 ♂ (RER 23 _ COE 213; UMRI), 11 vi 2023; M 13 d: 1 ♂ (RER 23 _ COE 304; UMRI), 29 ix 2023; P 1 b: 1 ♂ (RER 23 _ COE 209; BRIN), 31 v 2023; P 3 d: 1 ♂ (RER 23 _ COE 208; UMRI), 30 v 2023; P 5 b: 1 ♂ (RER 23 _ COE 217; UMRI), 21 vi 2023; P 5 d: 1 ♂ (RER 23 _ COE 218; BRIN), 21 vi 2023; P 6 c: 1 ♂ (RER 23 _ COE 302; UMRI), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB3FF80FDCD250C9594FB18.taxon	discussion	Remarks. Very common throughout the Kampar Peninsula. The male collected at location P 3 d was in the female position in a same sex tandem with Ceriagrion auranticum.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB3FF80FDCC226095ECF8B3.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB3FF80FDCC226095ECF8B3.taxon	materials_examined	Specimens collected. M 1 i: 1 ♂ (RER 20 _ COE 112; RER), 15 i 2020; 1 ♀ (RER 20 _ COE 113; BRIN), 15 i 2020; M 2 a: 3 ♂♂ (RER 20 _ COE 114 – 116; UMRI), 16 i 2020; 2 ♀♀ (RER 20 _ COE 117 – 118; RER), 16 i 2020; 2 ♀♀ (RER 20 _ COE 119 – 120; BRIN), 16 i 2020; M 2 b: 2 ♂♂ (RER 20 _ COE 121 – 122; UMRI), 16 i 2020; M 3 c: 2 ♀♀ (RER 23 _ COE 93 – 94; RER), 10 ii 2023; 2 ♂♂ (RER 23 _ COE 95 – 96; BRIN), 10 ii 2023; M 4 a: 1 ♀ (RER 23 _ COE 219; RER), 5 vi 2023; M 5 b: 1 ♂, 1 ♀ (RER 23 _ COE 221 - 222; UMRI), 17 vi 2023; M 7 a: 2 ♂♂ (RER 23 _ COE 123 – 124; RER), 23 i 2020; M 7 b: 1 ♂ (RER 23 _ COE 125; BRIN), 23 i 2020; 1 ♂ (RER 20 _ COE 167; UMRI), 23 i 2020; M 7 f: 1 ♂ (RER 23 _ COE 97; UMRI), 19 ii 2023; M 10 a: 1 ♀ (RER 23 _ COE 220; BRIN), 8 vi 2023; M 13 d: 1 ♂ (RER 23 _ COE 306; RER), 29 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB3FF80FDCC226095ECF8B3.taxon	discussion	Remarks. This is the first record of C. praetermissum from Riau Province. It was found quite frequently but at low densities at well vegetated drains but also sometimes on dry trails in PSF.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB0FF83FDD127F19449FCE1.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB0FF83FDD127F19449FCE1.taxon	materials_examined	Specimens collected. M 1 i: 1 ♀ (RER 20 _ COE 127; RER), 15 i 2020; M 2 e: 1 ♂ (RER 23 _ COE 98; UMRI), at dusk, 6 ii 2023; 1 ♂ (RER 23 _ COE 225; UMRI), 10 vi 2023; M 7 a: 1 ♂ (RER 23 _ COE 99, RER), 19 ii 2023; M 7 g: 1 ♂, 1 ♀ (RER 23 _ COE 100 – 101; BRIN), 19 ii 2023; 1 ♂ (RER 23 _ COE 102; UMRI), 19 ii 2023; 1 ♂ (RER 23 _ COE 226; RER), 15 vi 2023; M 8 a: 1 ♂, 1 ♀ (RER 20 _ COE 128 – 129; BRIN), in tandem, 16 i 2020; P 4 a: 1 ♂ (RER 23 _ COE 223; RER), 31 v 2023; 1 ♂ (RER 23 _ COE 224; BRIN), 2 vi 2023; P 6 a: 1 ♀ (RER 23 _ COE 307; BRIN), 22 ix 2023; 1 ♂, 1 ♀ (RER 23 _ COE 308 – 309; UMRI), in tandem, 22 ix 2023; P 6 b: 1 ♂ (RER 23 _ COE 312; BRIN), 23 ix 2023; P 6 e: 1 ♂, 1 ♀ (RER 23 _ COE 310 - 311; RER), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB0FF83FDD127F19449FCE1.taxon	discussion	Remarks. An (immature) male collected at M 7 g on 15 vi 2023 was perched in bright sunlight in the obelisk position with its wings half opened, several other males of the same species were perched around the same shallow pool in a normal manner for the species.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB1FF82FE1923DC959AF84C.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB1FF82FE1923DC959AF84C.taxon	materials_examined	Specimens collected. M 7 h: 1 ♂ (RER 23 _ COE 110; RER), 19 ii 2023; 1 ♂, 1 ♀ (RER 23 _ COE 161 – 162; RER), 13 vi 2023; 1 ♂, 3 ♀♀ (RER 23 _ COE 163 – 166; RER), 15 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB1FF82FE1923DC959AF84C.taxon	discussion	Taxonomic remarks. Until now M. megabinluyog had only been known from the type locality in freshwater mangrove in Brunei (Dow & Choong 2015). In February 2023 a single male Mortonagrion was collected at a pool just inside a stand of what appears to be mangrove near the Kampar River and was found to agree well with M. megabinluyog. In June 2023 the same stand of mangrove was entered at a different spot and a few individuals of both sexes were collected. Males agree in their anal appendages with M. megabinluyog but show some small differences in markings. Females have the posterior lobe of the prothorax more rounded in dorsal view than that shown for M. megabinluyog in fig. 6 in Dow & Choong (2015) and two of the females have blue dorsal markings dorsally on abdominal S 8, absent in the female paratype. None of these differences seem particularly significant, especially since variation even in the Brunei population can hardly be assessed with a type series of three males and a single female from one small site and we have little hesitation in assigning the Kampar Peninsula population to M. megabinluyog. This record is however surprising, and this species should be looked for not only in coastal areas in other parts of Borneo and Sumatra but also in Peninsular Malaysia.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB6FF85FE1127B99452FAFE.taxon	description	Selected references for Mortonagrion forficulatum	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB6FF85FE1127B99452FAFE.taxon	materials_examined	Specimens collected. M 4 a: 3 ♂♂ (RER 23 _ COE 107 – 109; RER), 13 ii 2023; 4 ♂♂ (RER 23 _ COE 151 – 154; RER), 5 vi 2023; 2 ♂♂ (RER 23 _ COE 158 – 159; RER), 14 vi 2023; M 4 b: 4 ♂♂ (RER 23 _ COE 155 – 157, 280; RER), one semi-teneral, 13 vi 2023; M 5 b: 1 ♂ (RER 23 _ COE 160; RER), 17 vi 2023; M 10 a: 1 ♂ (RER 23 _ COE 106; RER), 7 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB6FF85FE1127B99452FAFE.taxon	discussion	Taxonomic remarks. A series of a Mortonagrion (males only) collected in PSF in the Kampar Peninsula strongly resembles M. appendiculatum Lieftinck, 1937 (only known from Belitung) in general colouration but differs in the anal appendages. Mortonagrion forficulatum was described from present day Central Kalimantan (Lieftinck 1953 b) and all of the few later records are from Central or East Kalimantan. The Kampar Peninsula taxon has very similar anal appendages to those of M. forficulatum but differs strongly in markings, in particular males from confirmed populations of M. forficulatum have only a basal blue ring on abdominal S 7, with S 8 blue in the apical quarter and S 9 – 10 “ wholly deep black, but 9 occasionally with a pair of small, sky-blue baso-dorsal spots ” Lieftinck (1953 b: 394), in contrast the Kampar taxon has larger, irregular blue marks on S 7, and the dorsum of S 8 – 9 entirely blue. The Kampar taxon was initially thought likely to be a new species, but doubts developed because of the similarity of the anal appendages with those of M. forficulatum. Recently the first author was shown photographs (inaturalist. org / observations / 197961299) of an unidentified Mortonagrion from Kapuas Hulu Regency in West Kalimantan that resembles the Kampar taxon, little can be seen of the anal appendages in the photographs but what can be seen is at least similar to M. forficulatum and the Kampar taxon. There is no certain way of determining the identity of the species in the photographs without specimens, but one possibility is that it is M. forficulatum (another is that it is M. appendiculatum) and that the blue markings of the terminal abdominal segments become more extensive in the north and west of its range, in which case the Kampar population would be less atypical than it currently appears to be. In view of the uncertainty involved we have decided to leave the identity of the Kampar population open for the time being, rather than describing a new species or subspecies. Other remarks. All specimens of this taxon were collected in PSF, either at discrete pools or (most often) in extensively but shallowly flooded sections when the water table was high. Although present in February 2023 it was more abundant in June 2023, although it was not found at high densities at any time and no specimens were collected in September / October 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB6FF85FDFE229A9532F8ED.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB6FF85FDFE229A9532F8ED.taxon	materials_examined	Specimens collected. P 5 d: 2 ♂♂ (RER 23 _ COE 230 – 231; RER), 21 vi 2023; 2 ♂♂ (RER 23 _ COE 232 – 233; BRIN), 21 vi 2023; 1 ♂ (RER 23 _ COE 317; UMRI), 22 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB6FF85FDFE229A9532F8ED.taxon	discussion	Remarks. This is the first record of this taxon from mainland Sumatra. It was found at a large pond at the peat-mineral soil transition at the base of the Kampar Peninsula, where males (rather typically for the species) perched on lily pads or other vegetation close to the water surface.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB6FF84FDC6208D9442FE64.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB6FF84FDC6208D9442FE64.taxon	materials_examined	Specimens collected. M 4 b: 1 ♀ (RER 23 _ COE 111; BRIN), 8 ii 2023; M 5 f: 1 ♂ (RER 23 _ COE 234; RER), 6 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB6FF84FDC6208D9442FE64.taxon	discussion	Remarks. This is the first record from Riau Province. Solitary individuals of P. stictica were found in PSF at two locations in the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB7FF84FDAB25349784FCC2.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB7FF84FDAB25349784FCC2.taxon	materials_examined	Specimens collected. P 5 d: 1 ♂ (RER 23 _ COE 235; BRIN), 21 vi 2023; 1 ♂ (RER 23 _ COE 318; RER), 22 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB7FF84FDAB25349784FCC2.taxon	discussion	Remarks. Only found on the same large pond as P. calamorum dyeri and evidently not a common species in the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB7FF84FDFB2457924FF9B6.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB7FF84FDFB2457924FF9B6.taxon	materials_examined	Specimens collected. M 2 c: 2 ♂♂ (RER 20 _ COE 132 – 133; RER), 16 i 2020; M 2 e: 1 ♂ (RER 23 _ COE 112; RER), just before dusk, 6 ii 2023; M 3 a: 1 ♂ (RER 20 _ COE 137; UMRI), 17 i 2020; M 3 b: 3 ♂♂ (RER 20 _ COE 134 – 136; BRIN), 17 i 2020; M 3 e: 1 ♂ (RER 20 _ COE 146; UMRI), 24 i 2020; M 5 a: 2 ♂♂ (RER 20 _ COE 138 – 139; RER), 20 i 2020; 2 ♂♂ (RER 20 _ COE 140 – 141; BRIN), 20 i 2020; M 6: 2 ♂♂ (RER 20 _ COE 144 – 145; BRIN), 23 i 2020; M 7 e: 3 ♂♂ (RER 23 _ COE 113 – 115; BRIN), 19 ii 2023; M 7 g: 1 ♂ (RER 20 _ COE 143; RER), 23 i 2020; 3 ♂♂ (RER 23 _ COE 117 – 119; RER), 19 ii 2023; M 7 j: 1 ♂ (RER 23 _ COE 116; UMRI), 19 ii 2023; M 9 a: 1 ♂ (RER 20 _ COE 142; UMRI), 21 i 2020; M 11 a: 1 ♂, 1 ♀ (RER 23 _ COE 245 – 246; UMRI), in tandem, 11 vi 2023; 1 ♂ (RER 23 _ COE 247; BRIN), 11 vi 2023; M 13 a: 3 ♂♂ (RER 23 _ COE 248 – 250; UMRI), 18 vi 2023; M 13 b: 1 ♂ (RER 23 _ COE 251; BRIN), 18 vi 2023; 2 ♂♂ (RER 23 _ COE 322 – 323; BRIN), 29 ix 2023; P 2 a: 1 ♂ (RER 23 _ COE 236; BRIN), 30 v 2023; P 4 a: 3 ♂♂ (RER 23 _ COE 237 – 239; UMRI), 31 v 2023; 2 ♂♂ (RER 23 _ COE 240 – 241; BRIN), 31 v 2023; 1 ♂, 1 ♀ (RER 23 _ COE 242 – 243; UMRI), in tandem, 2 vi 2023; 1 ♂ (RER 23 _ COE 244; BRIN), 2 vi 2023; P 5 a: 2 ♂♂ (RER 23 _ COE 252 – 253; UMRI), 21 vi 2023; P 5 b: 1 ♂ (RER 23 _ COE 254, BRIN), 21 vi 2023; P 5 d: 2 ♂♂ (RER 23 _ COE 255 – 256; UMRI), 21 vi 2023; P 6 a: 1 ♂ (RER 23 _ COE 319; BRIN), 22 ix 2023; P 6 b: 1 ♂, 1 ♀ (RER 23 _ COE 320 – 321; UMRI), in tandem, 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB7FF84FDFB2457924FF9B6.taxon	discussion	Remarks. This widespread species is common throughout the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB7FF84FDF5218B928AF82A.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB7FF84FDF5218B928AF82A.taxon	materials_examined	Specimens collected. P 6 e: 1 ♂ (RER 23 _ COE 324; BRIN), 23 ix 2023; 1 ♂, 1 ♀ (RER 23 _ COE 325 – 326; UMRI), in tandem, 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB7FF84FDF5218B928AF82A.taxon	discussion	Remarks. Only found at a stream in mineral soils at the base of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB4FF87FDF127B99713FDEC.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB4FF87FDF127B99713FDEC.taxon	materials_examined	Specimens collected. P 5 a: 1 ♂ (RER 23 _ COE 257; BRIN), 21 vi 2023; P 6 a: 3 ♂♂ (RER 23 _ COE 327 – 329; UMRI), 22 ix 2023; P 6 b: 1 ♂, 1 ♀ (RER 23 _ COE 330 – 331; RER), in tandem, 23 ix 2023; P 6 e: 1 ♂, 1 ♀ (RER 23 _ COE 332 – 333; BRIN), in tandem, 23 ix 2023; 1 ♀ (RER 23 _ COE 334; UMRI), 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB4FF87FDF127B99713FDEC.taxon	discussion	Remarks. Only found at the base of the Kampar Peninsula, where it appears to be at least moderately common.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB4FF87FDD425759796FC13.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB4FF87FDD425759796FC13.taxon	materials_examined	Specimens collected. M 1 g: 1 ♂, 1 ♀ (RER 20 _ COE 147 – 148; RER), in tandem, 26 i 2020; 2 ♂♂ (RER 23 _ COE 149 – 150; BRIN), 26 i 2020; 2 ♂♂ (RER 20 _ COE 151 – 152; UMRI), 26 i 2020; M 1 p: 1 ♂ (RER 23 _ COE 120; RER), 12 ii 2023; 1 ♂, 1 ♀ (RER 23 _ COE 121 – 122, BRIN), 12 ii 2023; M 12 a: 1 ♂ (RER 23 _ COE 262; BRIN), 18 vi 2023; M 13 a: 1 ♂, 1 ♀ (RER 23 _ COE 258 – 259; UMRI), 18 vi 2023; 2 ♂♂ (RER 23 _ COE 260 – 261; RER), 18 vi 2023; M 13 b: 1 ♂ (RER 23 _ COE 335; UMRI), 29 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB4FF87FDD425759796FC13.taxon	discussion	Remarks. Fairly common among vegetation in the open lake areas on Sungai Serkap and also along the banks of the Kampar River.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB4FF86FDBE23A09721FE34.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB4FF86FDBE23A09721FE34.taxon	materials_examined	Specimens collected. M 1 b: 1 ♂ (RER 20 _ COE 160; RER), 18 i 2020; 2 ♂♂ (RER 20 _ COE 161 – 162; BRIN), 22 i 2020; 1 ♂ (RER 20 _ COE 163; UMRI), 22 i 2020; M 1 e: 1 ♂ (RER 20 _ COE 164; RER), 26 i 2020; M 2 a: 3 ♂♂ (RER 20 _ COE 153 – 155; RER), 16 i 2020; M 3 c: 2 ♂♂ (RER 20 _ COE 156 – 157; BRIN), 17 i 2020; 2 ♂♂ (RER 20 _ COE 158 – 159; UMRI), 17 i 2020; M 4 a: 1 ♂, 1 ♀ (RER 23 _ COE 133 – 134; RER), in tandem, 13 ii 2023; 1 ♂ (RER 23 _ COE 267; BRIN), 5 vi 2023; M 4 b: 3 ♂♂ (RER 23 _ COE 124 – 126; UMRI), 8 ii 2023; 2 ♂♂ (RER 23 _ COE 127 – 128; RER), 8 ii 2023; 3 ♂♂ (RER 23 _ COE 135 – 137; BRIN), 13 ii 2023, RER staff; 1 ♂ (RER 23 _ COE 270; BRIN), 13 vi 2023; M 5 b: 1 ♂, 1 ♀ (RER 23 _ COE 129 – 130; BRIN), in tandem, 9 ii 2023; 2 ♂♂ (RER 23 _ COE 131 – 132; UMRI), 9 ii 2023; M 7 i: 1 ♂ (RER 23 _ COE 138; UMRI), 19 ii 2023; 2 ♂♂ (RER 23 _ COE; UMRI), 15 vi 2023; M 10 a: 2 ♂♂ (RER 23 _ COE 123, 146; BRIN), 7 ii 2023; 2 ♂♂ (RER 23 _ COE 268 – 269; UMRI), 8 vi 2023; P 1 b: 3 ♂♂ (RER 23 _ COE 263 – 265; BRIN), 31 v 2023; 1 ♂ (RER 23 _ COE 266; UMRI), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB4FF86FDBE23A09721FE34.taxon	discussion	Taxonomic Remarks. Dow (2010) provided the first description of the female of T. ruficollis, based on a female from Sarawak, noting that the posterior pronotal lobe had a “ deep and narrow (ca 1 / 16 of width) central rectangular excision ”, illustrating this character in his Fig. 3 d. Later the first author examined a red-marked female Teinobasis in the collections of the Lee Kong Chian Natural History Museum in Singapore, collected in Singapore where the only red-marked Teinobasis recorded is T. ruficollis; this specimen lacked the excision in the posterior pronotal lobe, raising doubts over the value of this character for identification. More recently (2021 – 2022) the first author collected three additional females of T. ruficollis from Sarawak, two in tandem with males, all have the same excision in the posterior pronotal lobe as the female described in Dow (2010). The series from the Kampar Peninsula includes two females, both collected in tandem, one of these females lacks the pronotal lobe excision, the other has an excision, but shallower than that of the Sarawak females. It appears that the posterior pronotal lobe excision provides a diagnostic character for females of T. ruficollis in Sarawak (and quite possibly throughout Borneo) but is variably present and developed in elsewhere in its range and therefore not reliably diagnostic in general. Remarks. Almost all Sumatran records of T. ruficollis are from satellite islands, only the record in Ris (1927) is from the mainland, but the species has also been recorded from Belitung (firstly by Lieftinck 1954), the Lingga Islands (St. Quentin 1968, as T. rajah Laidlaw, 1912 but with the male anal appendages illustrated and clearly those of T. ruficollis) and Durian Island (firstly by Lieftinck 1935 a). During the surveys in the Kampar Peninsula T. ruficollis was recorded from Riau Province for the first time and proved to be moderately common in the PSF of the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB5FF86FDDB253595FDFB6C.taxon	description	Selected relevant references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB5FF86FDDB253595FDFB6C.taxon	materials_examined	Specimens collected. M 1 e: 1 ♀ (RER 20 _ COE 165; RER), 26 i 2020; M 4 a: 1 ♀ (RER 23 _ COE 276; UMRI), 14 vi 2023; M 4 b: 2 ♀♀ (RER 23 _ COE 140 – 141; BRIN), 8 ii 2023; 3 ♀♀ (RER 23 _ COE 142 – 144; RER), 13 ii 2023; 3 ♀♀ (RER 23 _ COE 273 – 275; RER), 13 vi 2023; 2 ♀♀ (RER 276 – 277; UMRI), 13 vi 2023; M 10 a: 1 ♀ (RER 23 _ COE 139; BRIN), 7 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB5FF86FDDB253595FDFB6C.taxon	discussion	Taxonomic remarks. During the surveys in the Kampar Peninsula a number of female Teinobasis were collected, all in PSF, which are at least very closely related to the poorly known T. suavis from Central Kalimantan. As with a very similar female recorded from Sarawak by Dow (2010), these females might actually be T. suavis (significantly increasingly the known range of that species) but until males are found the possibility that they belong to some other, undescribed, species closely allied to T. suavis cannot be ruled out. Other remarks. The female collected in January 2020 was perched at chest height in PSF flooded by the Serkap River. Those collected in February and June 2023 were perched low and inconspicuously in vegetation to the sides of trail pools. Despite deliberate searches no males could be found.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB5FFBBFDBE224D92FFF81F.taxon	description	Figs 20, 21, 24 Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB5FFBBFDBE224D92FFF81F.taxon	materials_examined	Specimens collected. M 3 b: 1 ♂ (RER 20 _ AES 1; RER), 2 ♀♀ (RER 20 _ AES 2 – 3; RER), all immature, one female semi-teneral 17 i 2020; M 10 a: 1 ♂ (RER 23 _ AES 11; RER), 8 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FFB5FFBBFDBE224D92FFF81F.taxon	discussion	Taxonomic remarks. Eleven species of Gynacantha are known from Sundaland of which seven have been recorded from Sumatra (Dow et al. 2024). Two of these, G. dohrni and G. maclachlani, difer from the others in details of the male anal appendages and / or the absence of a well-defined T-spot on the frons. The differences between G. dohrni and G. maclachlani are subtle and have not been adequately documented. The two were both described in the same paper (Krüger 1899), G. dohrni from a male from Sumatra with an additional male from Java and three males and two females from “ Nord-Borneo ” (very likely referring to present day Sabah, note that for simplicity we state Sabah for these specimens hereafter), G. maclachlani from two males from Borneo: one from Sabah and the other from Brunei. The type series of both are in the Museum and Institute of Zoology, Polish Academy of Science, Warsaw, Poland (but see below). Krüger (1899) includes small, inadequate sketches of the male anal appendages of the two species in dorsal view (figs 9 – 10 on page 280 of Krüger 1899). Martin (1909) gave illustrations of the male anal appendages of both species, that of G. dohrni from a specimen in the Selys collection and that of G. maclachlani stated to be from a specimen in the Krüger collection so presumably one of the types; Lieftinck (1954: 101) states “ ♂ app., type ” in reference to this illustration. For G. dohrni Krüger (1899: 286) states “ Stirn sehr schmal, vorn gelbbraun, oben vorn mit breitem, braunen rand, aber ohne T-fleck. ” [“ Forehead very narrow, yellow-brown at the front, with a broad brown border at the top, but without a T-spot. ”]. Martin (1911: Plate 5, fig. 5) includes a colour illustration of G. dohrni, but this shows a T-spot on the frons and superior anal appendages different from Krüger’s illustration. Martin (1911) also includes separate illustrations of the anal appendages of dohrni in his Plate 5, reproduced from Martin (1909: fig. 204, and also those of G. dohrni are shown as those of G. basiguttata in fig. 197 in error, see Lieftinck (1934) under G. dohrni), curiously these do agree with Krüger’s illustration. To our knowledge there are no other published illustrations of G. maclachlani whereas the anal appendages of G. dohrni have been illustrated at least twice more: Ris (1911: fig. 11) gives a dorsal view alongside his G. demeter (Ris, 1911) and Lieftinck (1953 a: fig. 10 left side images) gives both a dorsal and a lateral view. Gynacantha demeter (not recorded from Sumatra) is very similar to both G. dohrni and G. maclachlani but especially the former, and Dow (2021, note 51) raised doubts over its status as a species separate from G. dohrni but this issue has not been resolved and cannot be resolved here. All three species lack a distinct T-spot on the frons (for G. machlachlani Krüger (1899: 319) only mentions this indirectly: “ Von hyalina Selys unterscheidet sie sich durch geringere gröbe, das fehlen des T-flecks ” [“ It differs from hyalina Selys by its smaller size and the absence of the T-spot ”]), easily separating them from G. subinterrupta Rambur, 1842 which has similar anal appendages and whose range overlaps broadly with theirs; this raises severe doubts over the colour illustration purporting to be G. dohrni in Martin (1911); also see below. The first author has been able to obtain images of almost the entire type series of both species held in the Krüger collection in Warsaw. This collection contains both males of G. maclachlani listed by Krüger and all of the material listed for G. dohrni with the exception of two males from Sabah. Both of the females of G. dohrni from Sabah (one of which is teneral, with one cercus still attached, the other mature with both cerci broken) are mislabeled as male, this error appears to have originated with Krüger (on labels stating: “ L. Krüger determ. 1926 ”), however this is almost certainly just careless label preparation not reflecting an error in the original paper. More problematically the type series might be composite, the male from Sabah and the mature female from Sabah both appear to have T-spots on the frons (this is based on images of the entire specimen rather than specifically of the head, so not taken from an ideal angle, and the apparent T-spots could be shadows or some other artefact) and if that is the case maybe are not the same species as the specimen from Sumatra, but this might explain Martin’s (1911) illustration. On the other hand, given that the determination labels were added to the type series more than two decades after the description of G. dohrni was published they might have simply been added to the wrong specimens (the incorrect sex on two of these labels might support this hypothesis). Although seemingly not present in Warsaw the other two males from the type series of G. dornhi from Sabah might still be in existence, either mislabeled in the main Krüger collection or in another museum. Of Krüger’s material at least the holotype of Heliaeschna crassa Krüger, 1899 is in the Naturalis Biodiversity Center, Leiden, so the missing G. dohrni specimens could be looked for there. Further to this, Lieftinck (1953 a: 258) indicates that in 1938 he had two specimens from the type series of G. dohrni on loan, the male from Sumatra and one specimen from Borneo (referred to as the type and a paratype, but this is incorrect, see below); the male from Sumatra was clearly returned but that from Borneo might not have been. Although Krüger (1899) made no explicit type designation in his description of G. dohrni he lists the male from Sumatra (shown in Fig. 20) first and the other material in parentheses, however this seems to indicate nothing more than that the latter specimens were not collected by Dohrn; the description is clearly composite. A label indicating “ Type ” is present with the male from Sumatra, but it is not clear when this was added, also a label stating that the specimen is G. dohrni determined by L. Krüger in 1926. Nothing that can be considered as a holotype designation is included in Krüger’s description of G. maclachlani but the type specimen from Sabah (Fig. 22) has a handwritten label stating “ type ” as well as a partly typed, partly handwritten label seemingly added by Lieftinck in 1938 and stating “ HOLOTYPE ” as well as a typed label stating “ Type ” and label indicating determination by L. Krüger in 1926. We are not aware of any published designation as holo- or lectotype for either species so in fact all the type specimens are syntypes (the first author asked the opinion of Jan van Tol, an authority on the International Code of Zoological Nomenclature on this issue and the latter agreed). There is no need to select a lectotype for G. maclachlani but if the type series of G. dohrni is indeed composite then there are grounds to select a lectotype, in which case the male from Sumatra already labelled as the type would be the obvious and best choice. Using the images of the type series of the two species and material from the Kampar Peninsula and Sarawak the following remarks can be made. The length of superior anal appendages (illustrated in dorsal view in Figs 24 – 26) falls into the same range in both species but the inferior appendage is shorter in G. dohrni compared to G. maclachlani so that the ratio of superior to inferior anal appendage is always greater in G. dohrni than G. maclachlani. Caution must be exercised in quantifying the previous sentence, the exact ratio in any given specimen will vary according to the method used to measure it, e. g. on how the base of the appendages is defined, which can appear different in lateral or dorsal view and in dorsal view can vary according to the exact angle that the specimen is viewed from, and the accuracy of the measurement, for instance an ordinary ruler will clearly give less accurate results than using a microscope lens with a scale. With non-pinned specimens the lateral view is easier to achieve, and the first author takes the measurement of the superior anal appendage from its lowest basal corner to the tip and that of the inferior anal appendage from the same point on the base of the superior anal appendage to the tip of the inferior anal appendage. Note that taking the ratio from measurements made in dorsal view of the same specimens can give a higher value because of the shape of abdominal segment 10. In the specimens from the Kampar Peninsula and Sarawak examined the (lateral) ratio in G. dohrni is consistently greater than 3.6, in one individual (from Sarawak) well in excess of 4, while that in G. maclachlani is consistently below 3.4, and only greater than 3.3 in two out of eight individuals measured. Since the stacked images of the anal appendages of the type material only include dorsal views, ratios for these were calculated (using Photoshop) by measuring the left superior anal appendage (except that shown in Fig. 26 where the left is deformed near the tip, and shorter than the right, in this case the right was used) from its outer lateral base (to be as close as possible to the method used in lateral view) to the level of the apex and the inferior appendage was measured from the a horizontal line at the level of the starting point for the superior anal appendage measurement to the tip; this gives ratios of ca. 3.6 for the syntype of G. dohrni from Sumatra and ca. 3.3 for the two G. maclachlani syntypes. Aside from the differences in the ratio of superior to inferior anal appendage length, the superior anal appendages of G. dohrni bear a small but well defined, rounded subbasal interior-ventral expansion, visible in both dorsal and lateral view (and visible even in the illustration in Krüger (1899 )). In contrast those of G. maclachlani lack such an expansion, although often a slight thickening is visible in the same position in dorsal view. The superior anal appendages of both species bear many inward directed, hair-like setae on their inner margins, easily visible in dorsal view. In G. dohrni there is an increase in the density and length of these setae starting at less than the length of the inferior anal appendage from the tip of that appendage while in G. maclachlani although there is an increase in density it begins more apically. Note that the illustration in Martin (1909) shows setae rather evenly distributed on the cerci of G. maclachlani, in contrast to the actual condition (compare fig. 205 in Martin (1909) with Figs 25 – 26 here). The last two differences are the most easily checked characters to differentiate males of the two species, no clear difference in, for instance, body or wing length has been found. There might however be differences in the anal appendages in lateral view but since the first author only requested dorsal views of the anal appendages in the type series of both species it has not yet been possible to check these in the types. In specimens from Sarawak and the Kampar Peninsula identified using the characters mentioned already, the superior anal appendage of G. dohrni viewed laterally is almost completely straight (which is in agreement with Martin’s illustrations of the anal appendages), but that of G. maclachlani is slightly curved over its length (in the same manner as illustrated for G. saltatrix Martin, 1909 by Asahina (1986: 90, fig. 67 )); the images of the specimen labelled as ‘ type’ of G. maclachlani, which is not pinned, include lateral views of the entire specimen inside a clear plastic envelope, it appears that the condition of the superior anal appendage is the same but certainty is not possible without a high magnification lateral view of the anal appendages (the G. dohrni holotype is pinned and no lateral view at all is currently available). This curved appearance to the cercus in lateral view is not shown in fig. 205 in Martin (1909), but considering there is at least one other problem with Martin’s illustrations of these two species, it cannot necessarily be taken as accurately depicting either of the syntypes of G. maclachlani. There also appears to be a small difference in the apex of the inferior anal appendages viewed laterally but this could not be checked at all in the type material. In the key in Krüger (1899: 19) the length of the pterostigma is used as a distinguishing character for G. dohrni and G. maclachlani, with that of the former stated to be in the range 3.5 – 4 mm long and that of the latter to be 3 mm long. Unfortunately, the pterostigma length does not serve to separate the two species in general, in specimens from Borneo both the length of, and the number of underlying cells to, the pterostigma broadly overlaps. Identification of females of both these Gynacantha species remains somewhat problematic. For G. dohrni Krüger (1899) had little to say on the two female syntypes (which might not be conspecific with the male in any case) beyond “ Ganz änlich dem ♂. Ein reifes ♀ mit auϐerordentlich dunkel angeräucherten flügeln. Appendices lang und dünn, hinten blattartig erweitert. Letzes Ventralsegment mit 2 Spitzen. ” [“ Very similar to the ♂. A mature ♀ with exceptionally dark, smoked wings. Appendices long and thin, expanding leaf-like posteriorly. Last ventral segment with two points. ”] Latter authors have little or nothing useful to add on the females and that of G. maclachlani has never been described (Martin (1909) includes a short description of the supposed female, but this is based on specimens from New Guinea that are undoubtedly not G. maclachlani), whilst in all cases the females appear to be associated with the males by supposition only. The males of both species have prominent auricles, Martin (1909: 199) describes those of G. dohrni as “ énormément grandes ” [“ enormously large ”] and states that “ ♀ comme le mâle, meis le thorax marron, l’abdomen bien moins gonflé à la base, beaucoup moins étranglé au 3 ”, ... ” [“ ♀ like the male, but the thorax brown, the abdomen much less swollen at the base, much less constricted at 3 ”, ... ”] perhaps implying that the female also has large auricles. More concretely, Lieftinck (1953 a) clearly implies that what he considered to be female G. dohrni have prominent auricles (unusual in females) when, writing about an atypical female specimen, he states “ … the auricles somewhat larger than usual … ” The first author of this paper has generally taken the presence of prominent (clearly visible with the unaided eye, as large or almost as large as those of the male) auricles on female Gynacantha from Sundaland that otherwise resemble male G. dohrni as diagnostic for G. dohrni and those listed above were identified on this basis. Female Gynacantha collected by the first author while feeding in numbers with male G. maclachlani (and no other species caught in the same ‘ flock’) during rain at a location in southwest Sarawak lack prominent auricles (not easily visible without magnification), so the presence of large, clearly visible without magnification auricles appears to distinguish female G. dohrni from female G. maclachlani but is not guaranteed to be diagnostic for female G. maclachlani. In summary, the males of G. dohrni and G. maclachlani can be separated from each other based on the ratio of the superior to the inferior anal appendages, the presence of a small but well defined rounded subbasal expansion on the superior appendages of G. dohrni and differences in the density of setae along the inner margins of the superior appendages. As far as is known females can be separated based on the size of the auricles, with females of G. dohrni having auricles that are larger than usually found in females Gynacantha. At present G. maclachlani is known from Borneo and Sumatra while G. dohrni is known from Peninsular Malaysia, Borneo, Sumatra and Java (Dow et al. 2024) and there is at least one record from Indochina. Given the problemtic identification of these species it is however not unlikely that some published records of G. dohrni are actually of G. maclachlani in whole or part, but such records will need to be individually checked by examination of the specimens concerned. When publishing further records of these two species it is advisable to mention the characters used for identification.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF89FFBDFDDF27B997A1FF30.taxon	description	Figs 22, 23, 25, 26 Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF89FFBDFDDF27B997A1FF30.taxon	materials_examined	Specimens collected. M 1 b: 1 ♂ (RER 20 _ AES 4, RER), 22 i 2020; M 2 f: 1 ♂ (RER 23 _ AES 2; RER), at lights at 6: 30 – 7 PM, 17 ii 2023; M 5 a: 1 ♂ (RER 23 _ AES 1; RER), 9 ii 2023; M 10 a: 1 ♂ (RER 23 _ AES 13; UMRI), 8 vi 2023; P 1 d: 1 ♂ (RER 23 _ AES 12; BRIN), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF89FFBDFDDF27B997A1FF30.taxon	discussion	Remarks. Gynacantha maclachlani is recorded from Riau Province for the first time. See the taxonomic remarks under G. dohrni.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBDFD56262D94EAFDD8.taxon	materials_examined	Specimens collected. M 2 f: 1 ♀ (RER 23 _ AES 4; RER), at lights at ca. 6: 45 PM, 14 ii 2023; 1 ♀ (RER 23 _ AES 5; RER), at lights after 7 PM, 14 ii 2023; 1 ♀ (RER 23 _ AES 6; RER), at lights after 7 PM, 18 ii 2023, leg. Eco Camp canteen staff; M 5 a: 1 ♀ (RER 23 _ AES 27; RER), 28 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBDFD56262D94EAFDD8.taxon	discussion	Remarks. None of these females are G. dohrni, most or all of them are probably G. maclachlani but since there could be additional Gynacantha species present in the area, with females difficult to distinguish from those of G. maclachlani, their identity is best left open for now. Note that these undetermined specimens are not included in the count of species recorded from the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBDFDB125F99583FB36.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBDFDB125F99583FB36.taxon	materials_examined	Specimens collected. M 2 f: 1 ♂ (RER 23 _ AES 9; RER), at lights at 6: 30 – 7 PM, 12 ii 2023; 1 ♂ (RER 23 _ AES 14; BRIN), at lights at ca. 7: 15 PM, 4 vi 2023; 1 ♂ (RER 23 _ AES 15), at lights at ca. 6: 45 PM, 7 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBDFDB125F99583FB36.taxon	discussion	Taxonomic remark. The identification of specimens as either H. crassa or H. idae (Brauer, 1865) here is consistent with other such determinations made by the first author. The reader is referred to note 52 in Dow (2021) for a brief discussion of the issues involved; a thorough study of the types and much other material as well as accurate and consistent translations of original descriptions into English is needed and such a study might result in changes to published identifications and even the introduction of an additional name. Other remark. Recorded from Riau Province for the first time.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBDFDB5220A9285F97D.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBDFDB5220A9285F97D.taxon	materials_examined	Specimens collected. M 2 f: 1 ♂ (RER 23 _ AES 17; BRIN), in toilet at Eco Camp, 8 vi 2023, Eco Camp housekeeping staff; 1 ♂ (RER 23 _ AES 28; UMRI), at lights at 7 – 7: 30 PM, 2 x 2023; P 1 d: 1 ♂ (RER 23 _ AES 16; RER), 31 v 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBDFDB5220A9285F97D.taxon	discussion	Remarks. There has been no record from mainland Sumatra since the vague “ Sumatra ” in Lieftinck (1954). Recorded from Riau Province for the first time. See the taxonomic remark under H. crassa.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBCFD5421C6942FFE80.taxon	materials_examined	Specimens collected. M 2 f: 1 ♀ (RER 23 _ AES 7; RER), at lights at ca. 7: 30 PM, 9 ii 2023; 1 ♀ (RER 23 _ AES 8; RER), at lights 6: 30 – 7 PM, 9 ii 2023; 1 ♀ (RER 23 _ AES 18; RER), at lights at ca. 7 PM, 3 vi 2023; 1 ♀ (RER 23 _ AES 19; RER), at lights at ca. 7 PM, 4 vi 2023; 1 ♀ (RER 23 _ AES 22; RER), at lights at ca. 6: 40 PM, 10 vi 2023; 1 ♀ (RER 23 _ AES 23; RER), at lights at ca. 6: 30 PM, 11 vi 2023; 1 ♀ (RER 23 _ AES 24; RER), found in laboratory, 15 vi 2023; 1 ♀ (RER 23 _ AES 25; RER), found in building, 18 vi 2023, leg Eco Camp housekeeping staff; 1 ♀ (RER 23 _ AES 26; RER), at lights at ca. 6: 30 PM, 19 vi 2023, leg Eco Camp canteen staff; M 4 d: 2 ♀♀ (RER 23 _ AES 20 – 21; RER), flying over road at dusk, 9 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8EFFBCFD5421C6942FFE80.taxon	discussion	Remarks. In the opinion of the first author, females of H. idae and H. crassa cannot be reliably separated at this time, with the situation complicated by (possible) polymorphism in wing markings. It is therefore better to leave the females identified only to genus. Note that these undetermined specimens are not included in the count of species recorded from the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8FFFBCFD2C25699760FB98.taxon	discussion	Remarks. A single male Gomphidia was seen perched at the side of a closed canopy PSF stream (Simpang Sianyir, M 1 m) during suboptimal weather conditions on 1 x 2023, but flew away before the boat could be brought into a position from which capture was possible. The individual was very darkly coloured, more so than is typical for any species of Gomphidia known from Sundaland. Two species of Gomphidia have been recorded from Sumatra — G. abbotti Williamson, 1907 and G. machlachlani Selys, 1873 (but see below) — both of these species have been recorded in low pH waters elsewhere (e. g. Dow et al. (2012) for G. abbotti and Dow & Reels (2024) for G. machlachlani) but G. abbotti can be ruled out by the coloration of the Simpang Sianyir male. Gomphidia machlachlani is darker than G. abbotti and although still not typically as dark as the Simpang Sianyir male, the first author has collected one male almost as dark as that individual in Sarawak. Moreover, G. machlachlani has been found in numbers in very similar conditions at Maludam National Park in Sarawak (Dow & Reels 2024). Although the Simpang Sianyir male might be G. machlachlani, Lieftinck (1954) questioned the identity of Sumatran material supposedly of that species. Specimens will need to be collected before identification to species can be made. Whichever species of Gomphidia is present in the Kampar Peninsula, the genus has not been recorded in Riau Province until now.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8FFFBCFDD022399764F814.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8FFFBCFDD022399764F814.taxon	materials_examined	Specimens collected. M 1 a: 2 ♂♂ (RER 20 _ GOM 1, 4; RER), 15 i 2020; 1 ♂ (RER 20 _ GOM 2; BRIN), 15 i 2020; 1 ♂ (RER 20 _ GOM 3; UMRI), 15 i 2020; 1 ♂ (RER 20 _ GOM 5; BRIN), 18 i 2020; 1 ♂ (RER 20 _ GOM 6; UMRI), 19 i 2020; 1 ♂ (RER 23 _ GOM 2; UMRI), 12 ii 2023; 1 ♂ (RER 23 _ GOM 13; UMRI), 14 vi 2023; 1 ♀ (RER 23 _ GOM 26; RER), 4 x 2023; M 1 f: 1 ♂ (RER 23 _ GOM 6; BRIN), 17 ii 2023; 1 ♂ (RER 23 _ GOM 27; BRIN), 4 x 2023; M 1 j: 1 ♂ (RER 23 _ GOM 25; BRIN), 2 x 2023; M 1 l: 1 ♂ (RER 23 _ GOM 3; RER), 15 ii 2023; 1 ♂ (RER 23 _ GOM 4; BRIN), 15 ii 2023; M 1 m: 1 ♂ (RER 23 _ GOM 5; UMRI), 16 ii 2023; M 5 a: 1 ♂ (RER 20 _ GOM 7; RER), 20 i 2020; 1 ♂ (RER 20 _ GOM 8; BRIN), 20 i 2020; 1 ♂ (RER 20 _ GOM 9; UMRI), 20 i 2020; 1 ♂ (RER 20 _ GOM 10; RER), 25 i 2020; 1 ♂ (RER 23 _ GOM 1; BRIN), 9 ii 2023; 1 ♂ (RER 23 _ GOM 12; BRIN), 6 vi 2023; 1 ♂ (RER 23 _ GOM 23; BRIN), 28 ix 2023; 1 ♀ (RER 23 _ GOM 24; UMRI), 28 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8FFFBCFDD022399764F814.taxon	discussion	Remarks. Recorded from the mainland of Sumatra for the first time. Aside from the record from Belitung and a few records from Pahang (e. g. Norma-Rashid et al. 2001, Dow et al. 2012, Choong et al. 2016), most records of I. acutus are from Borneo. When the habitat has been recorded the species is invariably associated with low pH waters; its preferred habitat is low gradient streams in PSF or other forest formations with low pH and it can sometimes be found at drains with flow at the edge of PSF (although it has not been found in this situation in the Kampar Peninsula).	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8CFFBFFE6427B9926AFC8E.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8CFFBFFE6427B9926AFC8E.taxon	materials_examined	Specimens collected. M 1 l: 1 ♂ (RER 23 _ GOM 8; BRIN), 15 ii 2023; M 3 a: 1 ♀ (RER 20 _ GOM 12; BRIN), 17 i 2020; M 3 b: 1 ♂ (RER 20 _ GOM 11; RER), 17 i 2020; M 3 c: 1 ♀ (RER 23 _ GOM 7; RER), 10 ii 2023; M 7 b: 1 ♂ (RER 20 _ GOM 13; UMRI), 23 i 2020; M 11 a: 2 ♂♂ (RER 23 _ GOM 17 – 18; RER), 11 vi 2023; M 12 a: 1 ♀ (RER 23 _ GOM 20; UMRI), 18 vi 2023; M 13 a: 1 ♂ (RER 23 _ GOM 19; BRIN), 18 vi 2023; M 13 b: 1 ♂ (RER 23 _ GOM 21; RER), 18 vi 2023; P 3 b: 1 ♂ (RER 23 _ GOM 14; UMRI), 30 v 2023; P 4 a: 1 ♂ (RER 23 _ GOM 15; RER), 31 v 2023; 1 ♂ (RER 23 _ GOM 16; BRIN), 2 vi 2023; P 5 d: 1 ♂ (RER 23 _ GOM 22; BRIN), 21 vi 2023; P 5 e: 1 ♂ (RER 23 _ GOM 28; UMRI), 22 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8CFFBFFE6427B9926AFC8E.taxon	discussion	Remarks. Generally common in the Kampar Peninsula. Although no specimens were collected there, this species is abundant on the lower part of Sungai Serkap (M 1 a). It is also far more abundant along the banks of the Kampar River than the single specimen collected there suggests. On the Kampar River many individuals were also seen on large clumps of floating grass and other vegetation in the river, detached from the banks and slowly floating downstream. Dow et al. (2018) listed I. decoratus melaenops from Riau Province as a first record, but at the time were unaware of the record in Gesriantuti et al. (2018) that had been published shortly before.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8CFFBFFDCD232A9772F9AA.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8CFFBFFDCD232A9772F9AA.taxon	materials_examined	Specimens collected. M 1 f: 1 ♂ (RER 23 _ GOM 29; RER), 4 x 2023; 1 ♂ (RER 23 _ GOM 30; BRIN), 4 x 2023; M 1 m: 1 ♂ (RER 23 _ GOM 9; RER), 16 ii 2023; 1 ♂ (RER 23 _ GOM 10; BRIN), 16 ii 2023; 1 ♂ (RER 23 _ GOM 11; UMRI), 16 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8CFFBFFDCD232A9772F9AA.taxon	discussion	Remarks. Macrogomphus decemlineatus is a relatively seldomly recorded species. Although certainly not confined to PSF and low pH waters, a significant percentage of the available records are from such habitats. It has also sometimes been found at streams in oil palm or at the oil palm — forest boundary. In the Kampar Peninsula it has so far only been found at two tributary streams of the Serkap River. In February 2023 in sunny conditions in the afternoon single males were found perched in small clearings on Sungai Sianyir, but in overcast conditions in October 2023 no individuals definitely of this species were seen at the same stream. Also in October 2023, but in sunny conditions, males were relatively common on Simpang Kanan, although the species had not been found there in February 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8CFFBEFDE9201E9752FEA5.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8CFFBEFDE9201E9752FEA5.taxon	materials_examined	Specimens collected. M 1 a: 1 ♂ (RER 20 _ MAC 1; RER), 18 i 2020; 1 ♂ (RER 20 _ MAC 2; BRIN); 26 i 2020; 1 ♂ (RER 23 _ MAC 2; BRIN), 12 ii 2023; 1 ♂ (RER 23 _ MAC 4; UMRI), 4 x 2023; M 5 a: 1 ♂ (RER 23 _ MAC 1; UMRI), 9 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8CFFBEFDE9201E9752FEA5.taxon	discussion	Remarks. Frequently found flying over broad, open areas of Sungai Serkap. Less common on the smaller Sungai Sangar.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8DFFBEFDAF214C92A3F81C.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8DFFBEFDAF214C92A3F81C.taxon	materials_examined	Specimens collected. M 1 i: 1 ♀ (RER 20 _ LIB 1; RER), 15 i 2020; M 2 e: 1 ♀ (RER 23 _ LIB 1; BRIN), near dusk, 6 ii 2023; 1 ♂ (RER 23 _ LIB 117; UMRI), 10 vi 2023; 1 ♀ (RER 23 _ LIB 118; BRIN), 10 vi 2023; M 7 a: 1 ♂ (RER 20 _ LIB 3; UMRI), 23 i 2020; M 7 b: 1 ♂ (RER 20 _ LIB 5; BRIN), 23 i 2020; M 7 c: 1 ♂ (RER 20 _ LIB 6; UMRI), 23 i 2020; M 7 d: 1 ♂ (RER 23 _ LIB 2; UMRI), 19 ii 2023; M 7 g: 1 ♂ (RER 20 _ LIB 4; RER), 23 i 2020; 1 ♂ (RER 23 _ LIB 3; BRIN), 19 ii 2023; M 8 a: 1 ♀ (RER 20 _ LIB 2; BRIN), 16 i 2020.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8DFFBEFDAF214C92A3F81C.taxon	discussion	Remarks. Moderately common at open waters in the main part of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8DFFBEFDAC23159578FA68.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8DFFBEFDAC23159578FA68.taxon	materials_examined	Specimens collected. M 5 f: 1 ♀ (RER 23 _ COR 1; BRIN), 6 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8DFFBEFDAC23159578FA68.taxon	discussion	Remarks. Until now the vague records in Lieftinck (1954) have been the only ones of H. tenera from Sumatra. Although ‘ east’ Sumatra might refer to Riau Province, it might not and in the absence of concrete evidence to the contrary we consider the record from the Kampar Peninsula to be the first from Riau Province. A single female was collected while it was flying over wet peat near a canal in forest.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8DFFBEFD8926E197F2FC85.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8DFFBEFD8926E197F2FC85.taxon	materials_examined	Specimens collected. M 1 a: 1 ♂ (RER 20 _ MAC 3; RER), 15 i 2020; 1 ♂ (RER 20 _ MAC 4; BRIN), 18 i 2020; 2 ♂♂ (RER 20 _ MAC 5, 7; UMRI), 19 i 2020; 1 ♂ (RER 20 _ MAC 6; BRIN), 19 i 2020; 1 ♀ (RER 23 _ MAC 3; RER), 12 ii 2023; 1 ♂ (RER 23 _ MAC 5; BRIN), 4 x 2023; 1 ♂ (RER 23 _ MAC 6; UMRI), 4 x 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF8DFFBEFD8926E197F2FC85.taxon	discussion	Remarks. Recorded from Riau Province for the first time, abundant on Sungai Serkap. No specimen has been collected on Sungai Sangar but Macromia of the same size have been seen flying on that stream and are, in all probability, the same species.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB1FD8E27F192B2FDB2.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB1FD8E27F192B2FDB2.taxon	materials_examined	Specimens collected. M 7 a: 2 ♂♂ (RER 20 _ LIB 7, 10; RER), 23 i 2020; 2 ♂♂ (RER 20 _ LIB 8, 11; BRIN), 23 i 2020; 1 ♂ (RER 20 _ LIB 9; UMRI), 23 i 2020; 1 ♂ (RER 23 _ LIB 4; BRIN), 19 ii 2023; M 7 b: 1 ♂ (RER 20 _ LIB 12; UMRI), 23 i 2020; 1 ♂ (RER 20 _ LIB 13; RER), 23 i 2020.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB1FD8E27F192B2FDB2.taxon	discussion	Remarks. The vague record in Lieftinck (1954) was the only one of A. aethra from Sumatra until now, as in similar cases we treat our record as the first from Riau Province. This is generally a locally occurring species and in the Kampar Peninsula it was only found on acacia edge drains in one part of Meranti Estate.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB1FE35258894FAFC1E.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB1FE35258894FAFC1E.taxon	materials_examined	Specimens collected. M 2 b: 1 ♂ (RER 20 _ LIB 14; RER), 16 i 2020; 1 ♂ (RER 20 _ LIB 15; BRIN), 16 i 2020; 1 ♂ (RER 20 _ LIB 16; UMRI), 16 i 2020; M 2 e: 1 ♂ (RER 23 _ LIB 119; BRIN), 10 vi 2023; 1 ♂ (RER 23 _ LIB 120; UMRI), 10 vi 2023; M 7 a: 1 ♂ (RER 20 _ LIB 17; RER), 23 i 2020.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB1FE35258894FAFC1E.taxon	discussion	Remarks. Recorded from Riau Province for the first time. Found at drains and ponds with plentiful floating vegetation in the main part of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB1FDAB2362972BF9B0.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB1FDAB2362972BF9B0.taxon	materials_examined	Specimens collected. M 1 g: 1 ♂ (RER 23 _ LIB 281; RER), 2 x 2023; M 1 p: 2 ♂♂ (RER 23 _ LIB 5 (teneral), 8; BRIN), 12 ii 2023; 2 ♂♂ (RER 23 _ LIB 6 – 7; RER), 12 ii 2023; M 2 e: 1 ♂ (RER 23 _ LIB 121; BRIN), 10 vi 2023; M 5 a: 1 ♂ (RER 20 _ LIB 20; UMRI), 25 i 2020; 1 ♂ (RER 20 _ LIB 21; RER), 25 i 2020; M 7 a: 1 ♂ (RER 20 _ LIB 18; RER), 23 i 2020; 1 ♂ (RER 20 _ LIB 19; BRIN), 23 i 2020; 1 ♂ (RER 23 _ LIB 10; RER), 19 ii 2023; M 7 d: 1 ♂ (RER 23 _ LIB 9; UMRI), 19 ii 2023; M 11 a: 1 ♂ (RER 23 _ LIB 122; UMRI), 11 vi 2023; 1 ♂ (RER 23 _ LIB 123; RER), 11 vi 2023; 1 ♂ (RER 23 _ LIB 124; BRIN), 11 vi 2023; M 12 a: 1 ♀ (RER 23 _ LIB 280; UMRI), 29 ix 2023; P 5 d: 1 ♂ (RER 23 _ LIB 125; UMRI), 21 vi 2023; 1 ♂ (RER 23 _ LIB 278; RER), 22 ix 2023; P 6 b: 1 ♂ (RER 23 _ LIB 279; BRIN), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB1FDAB2362972BF9B0.taxon	discussion	Remarks. Recorded from Riau Province for the first time. Moderately common in open habitats in the Kampar Peninsula. The teneral male from Tasik Tengah (M 1 p) was found perched low in sparse emergent vegetation with its wings shut.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB0FDE321C09533FF1D.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB0FDE321C09533FF1D.taxon	materials_examined	Specimens collected. M 12 d: 1 ♂ (RER 23 _ LIB 126; RER), 18 vi 2023; 1 ♂ (RER 23 _ LIB 127; BRIN), 18 vi 2023; P 6 f: 1 ♀ (RER 23 _ LIB 282; UMRI), 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF82FFB0FDE321C09533FF1D.taxon	discussion	Remarks. Seemingly uncommon in the Kampar Peninsula, perhaps surprisingly so for a species that had been considered to be “ A littoral species, very abundant in the mangrove scrub and coastal swamps ” Lieftinck (1954: 131) but see the comments on this species in Dow (2022).	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF83FFB0FDA326659574FD7D.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF83FFB0FDA326659574FD7D.taxon	materials_examined	Specimens collected. M 10 a: 1 ♂ (RER 23 _ LIB 11; RER), 7 ii 2023; 1 ♀ (RER 23 _ LIB 12; BRIN), 7 ii 2023; 1 ♂ (RER 23 _ LIB 130; BRIN), 8 vi 2023; M 12 c: 1 ♀ (RER 23 _ LIB 132; BRIN), 18 vi 2023; M 12 d: 1 ♂ (RER 23 _ LIB 131; UMRI), 18 vi 2023; P 1 b: 1 ♀ (RER 23 _ LIB 128; UMRI), 31 v 2023; P 1 d: 1 ♂ (RER 23 _ LIB 129; RER), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF83FFB0FDA326659574FD7D.taxon	discussion	Remarks. Not uncommon in PSF in the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF83FFB0FDE724099597FA3D.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF83FFB0FDE724099597FA3D.taxon	materials_examined	Specimens collected. M 1 a: 1 ♂ (RER 23 _ LIB 17; UMRI), 12 ii 2023; M 1 i: 1 ♂ (RER 20 _ LIB 23; RER), 15 i 2020; 1 ♂ (RER 20 _ LIB 24; BRIN), 15 i 2020; M 2 b: 1 ♂ (RER 20 _ LIB 25; UMRI), 16 i 2020; M 2 c: 1 ♂ (RER 20 _ LIB 26; RER), 16 i 2020; M 2 e: 1 ♂ (RER 23 _ LIB 135; UMRI), 10 vi 2023; 1 ♂ (RER 23 _ LIB 136; RER), 10 vi 2023; M 3 b: 1 ♂ (RER 20 _ LIB 27; BRIN), 17 i 2020; 1 ♂ (RER 23 _ LIB 15; UMRI), 10 ii 2023; M 3 c: 1 ♂ (RER 23 _ LIB 13; UMRI), 10 ii 2023; 2 ♂♂ (RER 23 _ LIB 14, 16; BRIN), 10 ii 2023; M 6: 1 ♂ (RER 20 _ LIB 32; BRIN), 23 i 2020; M 7 a: 1 ♂ (RER 20 _ LIB 29; RER), 23 i 2020; M 7 b: 1 ♂ (RER 20 _ LIB 30; BRIN), 23 i 2020; 1 ♂ (RER 20 _ LIB 31; UMRI), 23 i 2020; M 7 d: 1 ♂ (RER 23 _ LIB 18; BRIN), 19 ii 2023; M 7 g: 1 ♂ (RER 23 _ LIB 19; UMRI), 19 ii 2023; M 9 a: 1 ♂ (RER 20 _ LIB 28; UMRI), 21 i 2020; M 11 a: 1 ♂ (RER 23 _ LIB 137; BRIN), 11 vi 2023; M 12 d: 1 ♂ (RER 23 _ LIB 138; UMRI), 18 vi 2023; M 13 b: 1 ♂ (RER 23 _ LIB 287; UMRI), 29 ix 2023; M 13 c: 1 ♂ (RER 23 _ LIB 288; BRIN), 29 ix 2023; P 1 a: 1 ♂ (RER 23 _ LIB 286; BRIN), 24 ix 2023; P 3 c: 1 ♀ (RER 23 _ LIB 133; RER), 30 v 2023; P 4 c: 1 ♂ (RER 23 _ LIB 134; BRIN), 30 v 2023; P 5 a: 1 ♂ (RER 23 _ LIB 139; RER), 21 vi 2023; P 5 d: 1 ♂ (RER 23 _ LIB 140; BRIN), 21 vi 2023; P 6 a: 1 ♂ (RER 23 _ LIB 283; UMRI), 22 ix 2023; P 6 b: 1 ♂ (RER 23 _ LIB 285; UMRI), 23 ix 2023; P 6 c 1 ♂ (RER 23 _ LIB 284; BRIN), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF83FFB0FDE724099597FA3D.taxon	discussion	Remarks. Common in open habitats in the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF83FFB3FDAE21499292FD60.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF83FFB3FDAE21499292FD60.taxon	materials_examined	Specimens collected. M 1 b: 1 ♂ (RER 20 _ LIB 39; RER), 22 i 2020; 1 ♂ (RER 20 _ LIB 40; BRIN), 22 i 2020; M 1 e: 1 ♂ (RER 20 _ LIB 41; UMRI), 26 i 2020; 1 ♂ (RER 20 _ LIB 42; BRIN), 26 i 2020; M 1 f: 1 ♀ (RER 20 _ LIB 34; BRIN), 19 i 2020; M 1 h: 2 ♂♂ (RER 20 _ LIB 35, 38; UMRI), 19 i 2020; 1 ♂ (RER 20 _ LIB 36, RER), 19 i 2020; 1 ♂ (RER 20 _ LIB 37; BRIN), 19 i 2020; M 1 j: 1 ♂ (RER 23 _ LIB 295; RER), 2 x 2023; M 1 m: 1 ♂ (RER 23 _ LIB 293; BRIN), 1 x 2023; M 1 o: 1 ♂ (RER 23 _ LIB 294; UMRI), 1 x 2023; M 1 q: 1 ♀ (RER 23 _ LIB 32; UMRI), 16 ii 2023; 1 ♂ (RER 23 _ LIB 46; RER), 16 ii 2023; M 2 a: 1 ♀ (RER 20 _ LIB 33; RER), 16 i 2020; M 4 a: 1 ♂, 1 ♀ (RER 23 _ LIB 26 – 27; UMRI), in tandem, 13 ii 2023; 1 ♂ (RER 23 _ LIB 145; RER), 5 vi 2023; 1 ♂ (RER 23 _ LIB 153; BRIN), 14 vi 2023; 1 ♂ (RER 23 _ LIB 291; BRIN), 27 ix 2023; M 4 b: 2 ♂♂ (RER 23 _ LIB 22 – 23; BRIN), 8 ii 2023; 1 ♂ (RER 23 _ LIB 28; BRIN), 13 ii 2023; 2 ♂♂ (RER 23 _ LIB 29 – 30; RER), 13 ii 2023, leg. RER staff; 1 ♂ (RER 23 _ LIB 31; BRIN), 13 ii 2023, leg. RER staff; 1 ♂ (RER 23 _ LIB 152; UMRI), 13 vi 2023; M 4 c: 1 ♂ (RER 23 _ LIB 24; UMRI), 8 ii 2023; M 4 e: 1 ♂ (RER 23 _ LIB 32; BRIN), 19 ii 2023; M 5 b: 1 ♂ (RER 23 _ LIB 25; BRIN), 9 ii 2023; 1 ♂ (RER 23 _ LIB 154; UMRI), 17 vi 2023; M 5 e: 1 ♀ (RER 23 _ LIB 148; UMRI), 6 vi 2023; M 5 f: 1 ♂, 1 ♀ (RER 23 _ LIB 146 – 147; BRIN), in tandem, 6 vi 2023; M 5 g: 1 ♂ (RER 23 _ LIB 292; UMRI), 28 ix 2023; M 10 a: 2 ♂♂ (RER 23 _ LIB 20 – 21; UMRI), 7 ii 2023; 1 ♂ (RER 23 _ LIB 149; BRIN), 8 vi 2023; M 11 b: 1 ♂ (RER 23 _ LIB 150; UMRI), 11 vi 2023; M 11 c: 1 ♂ (RER 23 _ LIB 151; BRIN), 11 vi 2023; M 12 b: 1 ♂ (RER 23 _ LIB 156; UMRI), 18 vi 2023; M 12 d: 1 ♂ (RER 25 _ LIB 155; BRIN), 18 vi 2023; P 1 b: 1 ♂ (RER 23 _ LIB 142; RER), 31 v 2023; 1 ♂ (RER 23 _ LIB 143; BRIN), 31 v 2023; 1 ♂ (RER 23 _ LIB 144; UMRI), 2 vi 2023; P 1 c: 1 ♂ (RER 23 _ LIB 289; BRIN), 24 ix 2023; P 3 d: 1 ♂ (RER 23 _ LIB 141; UMRI), 30 v 2023; P 6 f: 1 ♂ (RER 23 _ LIB 290; UMRI), 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF83FFB3FDAE21499292FD60.taxon	discussion	Remarks. Although quite regularly found in other habitats, B. oculata is primarily a swamp forest species, usually common in PSF and abundant in much of the main part of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF80FFB3FDA225E1975DFBA2.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF80FFB3FDA225E1975DFBA2.taxon	materials_examined	Specimens collected. M 1 r: 1 ♂ (RER 23 _ LIB 34; RER), 15 ii 2023; 2 ♂♂ (RER 23 _ LIB 35, 37; BRIN), 15 ii 2023; 1 ♂ (RER 23 _ LIB 36; UMRI), 15 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF80FFB3FDA225E1975DFBA2.taxon	discussion	Remarks. Recorded from mainland Sumatra for the first time, previously only known from Belitung and Borneo. A population of B. puella was found at a single section of flooded but rather open forest at the upper part of Sungai Serkap. Searches were made for the species at apparently similar sites in the same part of Sungai Serkap without success.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF80FFB3FDFC23F89211F9C3.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF80FFB3FDFC23F89211F9C3.taxon	materials_examined	Specimens collected. P 4 a: 1 ♂ (RER 23 _ LIB 157; RER), 31 v 2023; P 5 d: 1 ♂ (RER 23 _ LIB 158; BRIN), 21 vi 2023; 1 ♂ (RER 23 _ LIB 296; UMRI), immature, 22 ix 2023; 1 ♀ (RER 23 _ LIB 297; BRIN), immature, 22 ix 2023; P 6 b: 1 ♂ (RER 23 _ LIB 298; UMRI), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF80FFB3FDFC23F89211F9C3.taxon	discussion	Remarks. Only found around ponds at the base of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF80FFB3FDA521EF94DBF80A.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF80FFB3FDA521EF94DBF80A.taxon	materials_examined	Specimens collected. M 5 a: 1 ♀ (RER 23 _ LIB 299; BRIN), 28 ix 2023;	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF80FFB3FDA521EF94DBF80A.taxon	discussion	Remarks. Recorded from Riau Province for the first time. In addition to the female collected, two males were seen at location P 6 c on 23 ix 2023. The female was collected while flying over the main channel of Sungai Sangar and might have been far from its breeding habitat.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF81FFB2FE1427F19784FD7C.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF81FFB2FE1427F19784FD7C.taxon	materials_examined	Specimens collected. M 12 a: 1 ♂ (RER 23 _ LIB 159; RER), 18 vi 2023; 1 ♂ (RER 23 _ LIB 301; UMRI), 29 ix 2023; 1 ♂ (RER 23 _ LIB 302; RER), 29 ix 2023; P 6 b: 1 ♂ (RER 23 _ LIB 300; BRIN), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF81FFB2FE1427F19784FD7C.taxon	discussion	Remarks. Recorded from Indonesia for the first time, until now C. chini had only been known from a few sites in Peninsular Malaysia, with records from Pahang (Dow et al. 2007), Kelantan (Choong et al. 2017), Selangor (Dow & Ng 2020) and Terengganu (Choong et al. 2020). Found in the largely open lower part of Sungai Kutup where it occurred together with Chalybeothemis fluviatilis Lieftinck, 1933, a single male was found at a pond at the base of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF81FFB2FDCF25C5973CF9E2.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF81FFB2FDCF25C5973CF9E2.taxon	materials_examined	Specimens collected. M 1 a: 1 ♂ (RER 20 _ LIB 45; UMRI), 15 i 2020; 1 ♂ (RER 20 _ LIB 46; RER), 19 i 2020; 1 ♂ (RER 23 _ LIB 47; UMRI), 12 ii 2023; M 1 c: 1 ♂ (RER 20 _ LIB 43; RER), 15 i 2020; 1 ♂ (RER 20 _ LIB 44; BRIN), 15 i 2020; M 1 g: 1 ♂ (RER 23 _ LIB 54; UMRI), 26 i 2020; 1 ♂ (RER 20 _ LIB 55; RER), 26 i 2020; 1 ♂ (RER 23 _ LIB 310; UMRI), 2 x 2023; M 1 j: 1 ♂ (RER 23 _ LIB 309; BRIN), 2 x 2023; M 1 l: 1 ♂ (RER 23 _ LIB 48; BRIN), 15 ii 2023; M 1 p: 1 ♂ (RER 23 _ LIB 42; RER), 12 ii 2023; 2 ♂♂ (RER 23 _ LIB 43, 45; BRIN), 12 ii 2023; 1 ♂ (RER 23 _ LIB 44; UMRI), 12 ii 2023; M 5 a: 3 ♂♂ (RER 20 _ LIB 47, 49, 51; BRIN), 20 i 2020; 2 ♂♂ (RER 20 _ LIB 48, 50; UMRI), 20 i 2020; 1 ♂ (RER 20 _ LIB 52; UMRI), 25 i 2020; 1 ♀ (RER 20 _ LIB 53; BRIN), 25 i 2020; 1 ♂ (RER 23 _ LIB 38; RER), 9 ii 2023; 2 ♂♂ (RER 23 _ LIB 39, 41; BRIN), 9 ii 2023; 1 ♂ (RER 23 _ LIB 40; UMRI), 9 ii 2023; 1 ♂ (RER 23 _ LIB 160; RER), 6 vi 2023; M 12 a: 3 ♂♂ (RER 23 _ LIB 306 – 308; RER), 29 ix 2023; P 5 d: 1 ♂ (RER 23 _ LIB 303; RER), 22 ix 2023; 1 ♂ (RER 23 _ LIB 304; BRIN), 22 ix 2023; 1 ♂ (RER 23 _ LIB 305; UMRI), 22 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF81FFB2FDCF25C5973CF9E2.taxon	discussion	Remarks. Recorded from Riau Province for the first time. There is no previously published record from mainland Sumatra of C. fluviatilis but Pujiastuti et al. (2017) list Diplacodes trivialis (Rambur, 1842) with a photograph (their Fig. 3 g) that clearly shows C. fluviatilis rather than the Diplacodes, so at least part of their records are actually of C. fluviatilis. Common on open parts of the streams and rivers in the main part of the Kampar Peninsula, as it often is in low pH waters. Quite abundant on the large pond at location P 5 d in September 2023 but not even seen there in June 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF81FFB2FDD7218E9240F82E.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF81FFB2FDD7218E9240F82E.taxon	materials_examined	Specimens collected. P 6 c: 1 ♀ (RER 23 _ LIB 311; UMRI), 23 ix 2023; P 6 g: 1 ♂ (RER 23 _ LIB 312; RER), 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF81FFB2FDD7218E9240F82E.taxon	discussion	Remarks. Only found at the base of the Kampar Peninsula, not on the peat soils.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDC827F192F9FDC8.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDC827F192F9FDC8.taxon	materials_examined	Specimens collected. M 7 g: 1 ♂ (RER 20 _ LIB 56; RER), 23 i 2020; 1 ♂ (RER 23 _ LIB 49; BRIN), 19 ii 2023; 1 ♂ (RER 23 _ LIB 161; UMRI), 15 vi 2023; P 6 a: 1 ♂ (RER 23 _ LIB 313; RER), 22 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDC827F192F9FDC8.taxon	discussion	Remarks. Found at a rain-fed pond (lacking the black water seen almost everywhere else on the peat soils) beside the Kampar River in the main part of the peninsula and one site at the base of the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDD325E99465FC37.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDD325E99465FC37.taxon	materials_examined	Specimens collected. P 5 d: 1 ♂ (RER 23 _ LIB 314; UMRI), 22 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDD325E99465FC37.taxon	discussion	Remarks. A few individuals were seen at the large pond at location P 5 d on both occasions it was visited, with a single male collected on the second visit.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDA3230C9511FA97.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDA3230C9511FA97.taxon	materials_examined	Specimens collected. M 7 g: 1 ♂ (RER 20 _ LIB 57; RER), 23 i 2020; 1 ♂ (RER 23 _ LIB 50; BRIN), 19 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDA3230C9511FA97.taxon	discussion	Remarks. As with I. decoratus melaenops Dow et al. (2018) listed D. trivialis from Riau Province as a first record, in ignorance of Gesriantuti et al. (2018) that had been published shortly before. Apparently not common in the Kampar Peninsula, with records from only one site.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDD4212497EFF8D3.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDD4212497EFF8D3.taxon	materials_examined	Specimens collected. M 8 b: 1 ♀ (RER 20 _ LIB 58; UMRI), attracted to lights in building, 24 i 2020.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB5FDD4212497EFF8D3.taxon	discussion	Remark. More common in the Kampar Peninsula than the single record above suggests, seen flying over large, open drains on many occasions.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB4FDF920E09516FDCE.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB4FDF920E09516FDCE.taxon	materials_examined	Specimens collected. M 3 c: 1 ♂, 1 ♀ (RER 20 _ LIB 59, 62; RER), 17 i 2020; 1 ♂, 1 ♀ (RER 20 _ LIB 60, 63; BRIN), 17 i 2020; 1 ♂ (RER 20 _ LIB 61; UMRI), 17 i 2020; 1 ♂ (RER 23 _ LIB 51; UMRI), 10 ii 2023; 1 ♂ (RER 23 _ LIB 52; BRIN), 10 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF86FFB4FDF920E09516FDCE.taxon	discussion	Remarks. Selys (1889) lists L. difficilis from Banca (Bangka), this name is now considered a junior synonym of the neo-tropical Anatya guttata (Erichson, 1848). The record from Bangka obviously cannot refer to A. guttata and given that Selys placed it in Lathrecista we assume here that it refers to L. asiatica. Quite common under rubber trees at a site in the main part of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF87FFB4FD2825EB975AFB6A.taxon	materials_examined	Specimens collected. M 4 b: 1 ♀ (RER 23 _ LIB 162; RER), 13 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF87FFB4FD2825EB975AFB6A.taxon	discussion	Remarks. Although four species of Lyriothemis have been recorded in Sumatra (L. biappendiculata (Selys, 1878), L. cleis Brauer, 1868, L. magnificata (Selys, 1878) and L. salva Ris, 1927), none had been recorded in Riau Province until now. The single female collected is certainly not L. biappendiculata, is atypically coloured for L. cleis and is probably L. magnificata. We are not aware of any previous records of this genus definitely from PSF but, for instance, there is an old female specimen of L. magnificata from the Matang Road in Sarawak in the collection of the Sarawak Museum (Dow & Morris 2021), much of the surrounds of the Matang Road near to Kuching was originally PSF; there are also records of L. cleis from the surrounds of Tasik Bera (Norma-Rashid et al. 2001) and Tasik Chini (Kalkman 2004) in Pahang that could be from PSF. The female from the Kampar Peninsula was collected whilst it was flying around in a small clearing made by a treefall (L. cleis is quite often found in such situations in other types of forest).	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF87FFB4FDAE2207921AF95A.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF87FFB4FDAE2207921AF95A.taxon	materials_examined	Specimens collected. M 1 g: 1 ♂ (RER 23 _ LIB 317; BRIM), 2 x 2023; M 1 j: 1 ♂ (RER 23 _ LIB 316; RER), 2 x 2023; M 1 p: 1 ♂ (RER 23 _ LIB 54; RER), 12 ii 2023; M 1 r: 1 ♂, 1 ♀ (RER 23 _ LIB 56, 57; UMRI), 15 ii 2023; M 4 d: 1 ♂ (RER 23 _ LIB 53; UMRI), 8 ii 2023; M 4 e: 1 ♂ (RER 23 _ LIB 55; BRIN), 13 ii 2023; 1 ♂ (RER 23 _ LIB 315; UMRI), 27 ix 2023; M 5 a: 1 ♂ (RER 20 _ LIB 64; RER), 20 i 2020; M 5 d: 2 ♂♂ (RER 20 _ LIB 65 – 66), 23 i 2020; M 5 e: 1 ♂ (RER 23 _ LIB 163; RER), 6 vi 2023; M 11 a: 1 ♂ (RER 23 _ LIB 164; BRIN), 11 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF87FFB4FDAE2207921AF95A.taxon	discussion	Remarks. Moderately common in the main part of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF87FFB7FDF22077947CFEF8.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF87FFB7FDF22077947CFEF8.taxon	materials_examined	Specimens collected. P 1 c: 1 ♀ (RER 23 _ LIB 165; BRIN), teneral, 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF87FFB7FDF22077947CFEF8.taxon	discussion	Remarks. This is the first record of Nannophyopsis chalcosoma from mainland Sumatra, until now all records of N. chalcosoma had been from Belitung or Borneo. A single teneral female was collected while perched at head height over the ‘ stream’ at Tasik Anggrek.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF84FFB7FD8D269892B6FCC6.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF84FFB7FD8D269892B6FCC6.taxon	materials_examined	Specimens collected. M 3 d: 1 ♂ (RER 20 _ LIB 22; BRIN), 17 i 2020; M 4 b: 1 ♂ (RER 23 _ LIB 58; UMRI), 13 ii 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF84FFB7FD8D269892B6FCC6.taxon	discussion	Remarks. Only recorded twice in the Kampar Peninsula but likely to be more common there than these records suggest since it always occurs at low densities and can be wary so it is often under-recorded.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF84FFB7FDC524929232F8C2.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF84FFB7FDC524929232F8C2.taxon	materials_examined	Specimens collected. M 1 g: 1 ♀ (RER 20 _ LIB 75; BRIN), 26 i 2020; M 1 i: 1 ♂ (RER 20 _ LIB 67; RER), 15 i 2020; M 2 a: 1 ♂ (RER 20 _ LIB 68; BRIN), 16 i 2020; 1 ♂ (RER 20 _ LIB 69; UMRI), 16 i 2020; M 2 e: 1 ♂ (RER 23 _ LIB 59; UMRI), just before dusk, 6 ii 2023; 1 ♂ (RER 23 _ LIB 60; RER), just before dusk, 6 ii 2023; 2 ♂♂ (RER 23 _ LIB 171, 174; UMRI), 10 vi 2023; 1 ♂ (RER 23 _ LIB 172; RER), 10 vi 2023; 2 ♂♂ (RER 23 _ LIB 173, 175; BRIN), 10 vi 2023; M 2 f: 1 ♀ (RER 23 _ LIB 77; BRIN), at lights after 7 PM, 14 ii 2023, leg. Eco Camp canteen staff; 1 ♀ (RER 23 _ LIB 176; UMRI), at lights at ca. 7 PM, 14 vi 2023, leg. Eco Camp canteen staff; M 3 b: 1 ♂ (RER 20 _ LIB 70; RER), 17 i 2020; 1 ♂ (RER 23 _ LIB 63; BRIN), 10 ii 2023; 1 ♀ (RER 23 _ LIB 64; UMRI), 10 ii 2023; M 3 c: 1 ♂, 1 ♀ (RER 20 _ LIB 71 – 72; BRIN), in tandem, 17 i 2020; 3 ♂♂, 3 ♀♀ (♂♂: RER 23 _ LIB 65, 71, 73, ♀♀: RER 23 _ LIB 67, 69, 75; BRIN), 10 ii 2023, leg. RER staff; 2 ♂♂, 3 ♀♀ (♂♂: RER 23 _ LIB 66, 72, ♀♀: RER 23 _ LIB 68, 70, 74; UMRI), 10 ii 2023, leg. RER staff; M 4 e: 1 ♂ (RER 23 _ LIB 76; UMRI), 13 ii 2023; M 4 f: 1 ♂ (RER 23 _ LIB 170; BRIN), 5 vi 2023; M 5 d: 1 ♀ (RER 20 _ LIB 74; RER), 23 i 2020; M 7 b: 1 ♂ (RER 20 _ LIB 73; UMRI), 23 i 2020; M 7 g: 1 ♂ (RER 23 _ LIB 78; UMRI), 19 ii 2023; M 10 b: 1 ♀ (RER 23 _ LIB 62; UMRI), 7 ii 2023; M 10 c: 1 ♂ (RER 23 _ LIB 61; BRIN), 7 ii 2023; M 12 a: 1 ♂ (RER 23 _ LIB 178; UMRI), 18 vi 2023; M 12 c: 1 ♂ (RER 23 _ LIB 177; BRIN), 18 vi 2023; P 1 e: 1 ♂ (RER 23 _ LIB 167; BRIN), 2 vi 2023; P 4 a: 1 ♂ (RER 23 _ LIB 168; UMRI), 2 vi 2023; 1 ♂ (RER 23 _ LIB 169; RER), 2 vi 2023; P 4 b: 1 ♂ (RER 23 _ LIB 166; RER), 31 v 2023; P 5 a: 1 ♂ (RER 23 _ LIB 179; BRIN), 21 vi 2023; 1 ♂ (RER 23 _ LIB 180; UMRI), 21 vi 2023; 1 ♂ (RER 23 _ LIB 320; RER), 22 ix 2023; P 5 b: 1 ♂ (RER 23 _ LIB 181; BRIN), 21 vi 2023; P 5 d: 1 ♂ (RER _ LIB 319; RER), 22 ix 2023; P 5 e: 1 ♂ (RER 23 _ LIB 318; UMRI), 22 ix 2023; P 6 c: 1 ♂ (RER 23 _ LIB 322; RER), 23 ix 2023; P 6 e: 1 ♂ (RER 23 _ LIB 321; RER), 23 ix 2023;	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF84FFB7FDC524929232F8C2.taxon	discussion	Remarks. Very common in open habitats throughout the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF84FFB6FDBF20569428FE80.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF84FFB6FDBF20569428FE80.taxon	materials_examined	Specimens collected. P 3 c: 1 ♂ (RER 23 _ LIB 182; UMRI), 30 v 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF84FFB6FDBF20569428FE80.taxon	discussion	Remarks. Recorded from Riau Province for the first time, the single male collected was the only individual of this species seen. There are only a few records of N. fulvia from Sumatra, apart from the first record listed above (of three males), Lieftinck (1935 a) listed five specimens from Jambi and South Sumatra and Seehausen & Dow (2016) listed a single old specimen from Lampung; in total only 10 specimens of the species have been recorded from Sumatra, all from the mainland.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF85FFB6FDA526D19761FC07.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF85FFB6FDA526D19761FC07.taxon	materials_examined	Specimens collected. M 1 f: 1 ♂ (RER 23 _ 326; RER), 4 x 2023; M 1 m: 1 ♂ (RER 23 _ LIB 79; RER), 16 ii 2023; 1 ♂ (RER 23 _ LIB 80; BRIN), 16 ii 2023; 1 ♂ (RER 23 _ LIB 81; UMRI), 16 ii 2023; 1 ♂ (RER 23 _ LIB 324; BRIN), 1 x 2023; 1 ♂ (RER 23 _ LIB 325; UMRI), 1 x 2023; M 5 a: 1 ♂ (RER 23 _ LIB 323; RER), 28 ix 2023; M 5 f: 1 ♂ (RER 23 _ LIB 185; UMRI), 6 vi 2023; P 1 b: 1 ♂ (RER 23 _ LIB 183; RER), 31 v 2023; P 1 c: 1 ♂ (RER 23 _ LIB 184; BRIN), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF85FFB6FDA526D19761FC07.taxon	discussion	Remarks. Recorded from Riau Province for the first time, almost all Sumatran records of O. pruinans have been from Bangka-Belitung. Found at and around smaller forest streams and stream-like canals in PSF in the Kampar Peninsula. This is another species that, while certainly not confined to PSF, thrives in PSF where suitable microhabitats exist.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF85FFB6FDDE231C92E7F9F7.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF85FFB6FDDE231C92E7F9F7.taxon	materials_examined	Specimens collected. M 4 a: 1 ♂ (RER 23 _ LIB 192; RER), 5 vi 2023; M 5 f: 2 ♂♂ (RER 23 _ LIB 193, 196; BRIN), 6 vi 2023; 1 ♂ (RER 23 _ LIB 194; UMRI), 6 vi 2023; 1 ♂ (RER 23 _ LIB 195; RER), 6 vi 2023; M 5 g: 1 ♂ (RER 23 _ LIB 327; UMRI), 28 ix 2023; P 1 b: 1 ♂ (RER 23 _ LIB 188; UMRI), 31 v 2023; 1 ♂ (RER 23 _ LIB 189; RER), 31 v 2023; 1 ♂ (RER 2 _ LIB 190; BRIN), 2 vi 2023; P 1 c: 1 ♂ (RER 23 _ LIB 191; UMRI), 2 vi 2023; P 1 d: 1 ♀ (RER 23 _ LIB 186; RER), 30 v 2023; 1 ♀ (RER 23 _ LIB 187; BRIN), 30 v 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF85FFB6FDDE231C92E7F9F7.taxon	discussion	Remarks. Although eventually recorded from six locations in the Kampar Peninsula, O. pulcherrima was not found at all during the surveys in January 2020 and February 2023. This is generally a common species in swamp forest so its apparent absence early in the year is curious but could be an artefact of sampling effort.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF85FFA9FD8B21849208FEF8.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF85FFA9FD8B21849208FEF8.taxon	materials_examined	Specimens collected. M 2 c: 1 ♂ (RER 20 _ LIB 76; RER), 16 i 2020; M 3 b: 1 ♀ (RER 20 _ LIB 77; BRIN), 17 i 2020; 1 ♂ (RER 23 _ LIB 83; UMRI), 10 ii 2023; M 3 e: 1 ♂ (RER 20 _ LIB 79; RER), 24 i 2020; M 4 d: 1 ♂ (RER 23 _ LIB 82; BRIN), 8 ii 2023; M 7 b: 1 ♂ (RER 20 _ LIB 78; UMRI), 23 i 2020; M 12 a: 1 ♂ (RER 23 _ LIB 200; BRIN), 18 vi 2023; M 13 b: 1 ♂ (RER 23 _ LIB 330; RER), 29 ix 2023; P 1 a: 1 ♂ (RER 23 _ LIB 328; UMRI), 24 ix 2023; P 1 c: 1 ♂ (RER 23 _ LIB 199; UMRI), 2 vi 2023; P 1 e: 1 ♂ (RER 23 _ LIB 197; RER), immature, 30 v 2023; 1 ♂ (RER 23 _ LIB 198; BRIN), 30 v 2023; P 6 f: 1 ♂ (RER 23 _ LIB 329; RER), 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF85FFA9FD8B21849208FEF8.taxon	discussion	Remarks. Fairly common throughout the Kampar Peninsula. The immature male from location P 1 e was being consumed by a female of Orthetrum sabina when it was collected on 30 v 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA9FDA4264994B5FD74.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA9FDA4264994B5FD74.taxon	materials_examined	Specimens collected. P 6 f: 1 ♂ (RER 23 _ LIB 331; UMRI), 25 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA9FDA4264994B5FD74.taxon	discussion	Remarks. Seemingly absent from the main part of the Kampar Peninsula, with just a single record from the base of the peninsula made during our surveys.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA9FDAA25F59220FBCB.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA9FDAA25F59220FBCB.taxon	materials_examined	Specimens collected. P 5 a: 1 ♂ (RER 23 _ LIB 201; BRIN), 21 vi 2023; 1 ♂ (RER 23 _ LIB 202; UMRI), 21 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA9FDAA25F59220FBCB.taxon	discussion	Remarks. Orthetrum luzonicum had not been recorded from Riau Province prior to this. A few individuals were present in marshy sections with long emergent grasses at the margins of the stream in the afternoon in June 2023; no individuals were recorded on a morning visit to this site in September 2023, but higher water levels made the exact parts of the stream where it was found in June very difficult to access on that visit.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA9FDDC235D9776F8FA.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA9FDDC235D9776F8FA.taxon	materials_examined	Specimens collected. M 2 b: 1 ♂ (RER 20 _ LIB 81; BRIN), 16 i 2020; M 2 e: 1 ♂ (RER 23 _ LIB 208; UMRI), 10 vi 2023; M 3 a: 1 ♀ (RER 20 _ LIB 82; UMRI), 17 i 2020; M 3 e: 1 ♀ (RER 20 _ LIB 84; BRIN), 24 i 2020; M 7 d: 1 ♂ (RER 23 _ LIB 85; BRIN), 19 ii 2023; M 7 g: 1 ♂ (RER 20 _ LIB 83; RER), 23 i 2020; 1 ♂ (RER 23 _ LIB 86; UMRI), 19 ii 2023; M 8 a: 1 ♂ (RER 20 _ LIB 80; RER), 16 i 2020; M 10 b: 1 ♂ (RER 23 _ LIB 84; UMRI), 7 ii 2023; M 13 c: 1 ♂ (RER 23 _ LIB 334; UMRI), 29 ix 2023; P 1 e: 1 ♀ (RER 23 _ LIB 205; BRIN), 30 v 2023; P 3 b: 1 ♂, 1 ♀ (RER 23 _ LIB 203 – 204; RER), in tandem, 30 v 2023; P 3 c: 1 ♂ (RER 23 _ LIB 206; UMRI), 30 v 2023; P 4 a: 1 ♂ (RER 23 _ LIB 207; BRIN), 31 v 2023; P 5 a: 1 ♂ (RER 23 _ LIB 209; RER), 29 vi 2023; P 5 d: 1 ♂ (RER 23 _ LIB 332; UMRI), 22 ix 2023; P 6 e: 1 ♂ (RER 23 _ LIB 333; BRIN), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA9FDDC235D9776F8FA.taxon	discussion	Remarks. In addition to the specimen records above, O. sabina was very frequently seen at open drains around the plantations in the Kampar Peninsula, and around muddy pools on and beside roads, often in company with Pantala flavescens (Fabricius, 1798) in the latter situation. On at least two occasions, while sitting in a vehicle behind a lorry loaded with acacia logs, the first author observed O. sabina attempting to perch on the logs while the lorry was moving.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA8FE19204C92C7FE86.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA8FE19204C92C7FE86.taxon	materials_examined	Specimens collected. P 4 b: 1 ♂ (RER 23 _ LIB 210; RER), 31 v 2023; P 5 a: 1 ♂ (RER 23 _ LIB 211; BRIN), 21 vi 2023; P 6 e: 1 ♂ (RER 23 _ LIB 335; UMRI), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9AFFA8FE19204C92C7FE86.taxon	discussion	Remarks. Probably much more common in the Kampar Peninsula than the records above suggest, this species was seen in passing at muddy puddles on roads and at drains in acacia on a number of occasions.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9BFFA8FDAC26D39439FC8A.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9BFFA8FDAC26D39439FC8A.taxon	materials_examined	Specimens collected. M 1 i: 1 ♀ (RER 20 _ LIB 85; BRIN), flying over road, 18 i 2020; M 12 c: 1 ♂ (RER 23 _ LIB 212; UMRI), 18 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9BFFA8FDAC26D39439FC8A.taxon	discussion	Remarks. Pantala flavescens is much more common in the Kampar Peninsula than the specimen records suggest, during the surveys it was seen flying over roads and large drainage ditches during sunny weather almost everywhere visited in the peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9BFFABFDA3219196EDFEA4.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9BFFABFDA3219196EDFEA4.taxon	materials_examined	Specimens collected. M 7 h: 1 ♂ (RER 23 _ LIB 228; RER), 15 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9BFFABFDA3219196EDFEA4.taxon	discussion	Remarks. Recorded from Riau Province for the first time. One individual was collected and another seen in the same stand of freshwater mangrove beside the Kampar River where M. megabinluyog was found. Although described from South Sumatra the only later Sumatran records had been from Belitung (Alfarisyi 2018, 2019). Although fairly widely distributed in Sundaland, this species has not often been recorded but might well be more common than is currently obvious. It appears to prefer mangrove and nipa palm habitats that are difficult to work in and is often found at the banks of rivers or around creeks with deep water in these habitats, so that a boat is required to find it.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9BFFA8FDC5232795D2F989.taxon	description	Selected references for Pornothemis serrata Krüger, 1902:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9BFFA8FDC5232795D2F989.taxon	materials_examined	Specimens collected. M 2 a: 1 ♂ (RER 20 _ LIB 86; RER), 16 i 2020; M 4 a: 5 ♂♂ (RER 23 _ LIB 216 – 220; RER), 5 vi 2023; M 4 b: 4 ♂♂ (RER 23 _ LIB 87 – 90; RER), 13 ii 2023; 1 ♂, 1 ♀ (RER 23 _ LIB 221 – 222; RER), 5 vi 2023; 2 ♂♂ (RER 23 _ LIB 225 – 226; RER), 13 vi 2023; 1 ♂, 1 ♀ (RER 23 _ LIB 338 – 339; RER), 27 ix 2023; M 5 b: 1 ♂ (RER 23 _ LIB 227; RER), 17 vi 2023; M 5 e: 1 ♀ (RER 23 _ LIB 223; RER), 6 vi 2023; M 5 f: 1 ♂ (RER 23 _ LIB 340; RER), 28 ix 2023; M 10 a: 1 ♂ (RER 23 _ LIB 224; RER), 8 vi 2023; P 1 b: 1 ♂ (RER 23 _ LIB 213; RER), 31 v 2023; 2 ♂♂ (RER 23 _ LIB 214 – 215; RER), 2 vi 2023; P 1 c: 2 ♂♂ (RER 23 _ LIB 336 – 337; RER), 24 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9BFFA8FDC5232795D2F989.taxon	discussion	Remarks. This is the taxon listed as P. serrata A in, for instance, Dow (2021). Until recently it was not possible to tell which of three similar taxa was the true P. serrata. At least in the lowlands this is a PSF species while the true P. serrata appears to be primarily a species of freshwater swamp forest. The first author is working on a revision of Pornothemis and it would be premature to make further comments here.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF98FFABFDD026F594B7FC94.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF98FFABFDD026F594B7FC94.taxon	materials_examined	Specimens collected. P 4 a: 1 ♂ (RER 23 _ LIB 229; RER), 31 v 2023; 1 ♂ (RER 23 _ LIB 230; BRIN), 31 v 2023; 1 ♂ (RER 23 _ LIB 231; UMRI), 31 v 2023; P 5 a: 1 ♂ (RER 23 _ LIB 342; UMRI), at the shallow pool beside the stream, 22 ix 2023; P 5 f: 1 ♂ (RER 23 _ LIB 341; BRIN), 22 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF98FFABFDD026F594B7FC94.taxon	discussion	Remarks. Seemingly fairly common on the mineral soils at the base of the Kampar Peninsula but almost entirely absent from the peat soil part of the peninsula, where the only record is from a rain-fed artificial pond not in peat and presumably not with low pH water.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF98FFABFDB9232595F7F9D4.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF98FFABFDB9232595F7F9D4.taxon	materials_examined	Specimens collected. M 1 i: 1 ♂ (RER 20 _ LIB 87; RER), 15 i 2020; 1 ♀ (RER 20 _ LIB 88; BRIN), found in spider’s web, 18 i 2020; M 1 j: 1 ♂ (RER 23 _ LIB 343; RER), 2 x 2023; M 2 e: 1 ♂ (RER 23 _ LIB 91; BRIN), just before dusk, 6 ii 2023; 1 ♂ (RER 23 _ LIB 233; BRIN), 10 vi 2023; 1 ♀ (RER 23 _ LIB 234; UMRI), 10 vi 2023; M 3 c: 1 ♀ (RER 23 _ LIB 92; UMRI), 10 ii 2023, leg. RER staff; 1 ♀ (RER 23 _ LIB 116; BRIN), 10 ii 2023, leg. RER staff; M 5 a: 1 ♂ (RER 20 _ LIB 89; UMRI), 20 i 2020; 1 ♂ (RER 20 _ LIB 94; UMRI), 25 i 2020; M 7 a: 1 ♂ (RER 20 _ LIB 91; BRIN), 23 i 2020; 1 ♀ (RER 20 _ LIB 92; UMRI), perched in dry hole just inside acacia, 23 i 2020; 1 ♂ (RER 23 _ LIB 93; UMRI), 19 ii 2023; 1 ♂ (RER 23 _ LIB 94; BRIN), 19 ii 2023; M 7 c: 1 ♂ (RER 20 _ LIB 93; BRIN), 23 i 2020; M 9 a: 1 ♂ (RER 20 _ LIB 90; RER), 21 i 2020; M 11 a: 1 ♂ (RER 23 _ LIB 235; BRIN), 11 vi 2023; P 3 a: 1 ♂ (RER 23 _ LIB 232; UMRI), 30 v 2023; P 5 d: 1 ♀ (RER 23 _ LIB 236; RER), 21 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF98FFABFDB9232595F7F9D4.taxon	discussion	Remarks. Generally common at open waters in the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF98FFAAFDB221E595D3FE48.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF98FFAAFDB221E595D3FE48.taxon	materials_examined	Specimens collected. M 1 a: 2 ♂♂ (RER 20 _ LIB 96, 99; BRIN), 19 i 2020; 1 ♂ (RER 20 _ LIB 97; UMRI), 19 i 2020; 1 ♂ (RER 20 _ LIB 98; RER), 19 i 2020; 1 ♂ (RER 23 _ LIB 95; BRIN), semi-teneral, 12 ii 2023; M 1 c: 1 ♂ (RER 20 _ LIB 95; RER), 15 i 2020; M 1 j: 1 ♂ (RER 23 _ LIB 345; BRIN), 2 x 2023; 1 ♂ (RER 23 _ LIB 346; UMRI), 2 x 2023; 1 ♂ (RER 23 _ LIB 347; RER), 2 x 2023; M 2 e: 1 ♂ (RER 23 _ LIB 238; UMRI), 10 vi 2023; M 4 d: 1 ♂ (RER 23 _ LIB 344; UMRI), 27 ix 2023; M 5 a: 1 ♂ (RER 20 _ LIB 100; UMRI), 20 i 2020; 1 ♂ (RER 20 _ LIB 101; RER), 20 i 2020; 1 ♂ (RER 20 _ LIB 102; BRIN), 20 i 2020; M 7 a: 1 ♂ (RER 20 _ LIB 103; UMRI), 23 i 2020; 1 ♂ (RER 23 _ LIB 96; UMRI), 19 ii 2023; M 11 a: 2 ♂♂ (RER 23 _ LIB 239, 241; BRIN), 11 vi 2023; 1 ♂ (RER 23 _ LIB 240; UMRI), 11 vi 2023; P 1 c: 1 ♂ (RER 23 _ LIB 237; BRIN), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF98FFAAFDB221E595D3FE48.taxon	discussion	Remarks. Rhyothemis aterrima thrives in low pH waters (although it is not confined to them) and this is very evident in the Kampar Peninsula where the species is common on open sections of PSF streams with emergent vegetation and even occurs on some sections of drain at the acacia edge.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF99FFAAFDAC26D192CAFBC0.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF99FFAAFDAC26D192CAFBC0.taxon	materials_examined	Specimens collected. M 1 f: 1 ♂ (RER 20 _ LIB 104; RER), 19 i 2020; M 1 h: 1 ♂ (RER 20 _ LIB 105; BRIN), 19 i 2020; M 1 j: 1 ♂ (RER 23 _ LIB 351; UMRI), 2 x 2023; M 1 m: 1 ♂ (RER 23 _ LIB 99; UMRI), 16 ii 2023; M 4 c: 1 ♂ (RER 23 _ LIB 245; UMRI), 5 vi 2023; M 4 d: 1 ♂ (RER 23 _ LIB 97; RER), 8 ii 2023; M 5 a: 2 ♂♂ (RER 20 _ LIB 106 – 107; UMRI), 20 i 2020; 2 ♂♂ (RER 20 _ LIB 108 – 109; RER), 20 i 2020; 1 ♂, 1 ♀ (RER 20 _ LIB 110 – 111; BRIN), 20 i 2020; 1 ♂ (RER 20 _ LIB 112; UMRI), 25 i 2020; 1 ♂ (RER 23 _ LIB 98; BRIN), 9 ii 2023; M 5 e: 1 ♂ (RER 23 _ 246; BRIN), 6 vi 2023; M 12 a: 1 ♀ (RER 23 _ LIB 247; RER), 18 vi 2023; 1 ♂ (RER 23 _ LIB 350; BRIN), 29 ix 2023; P 1 a: 1 ♂ (RER 23 _ LIB 242; BRIN), 31 v 2023; P 1 c: 1 ♂ (RER 23 _ LIB 243; UMRI), 2 vi 2023; 1 ♂ (RER 23 _ LIB 244; BRIN), 2 vi 2023; P 1 e: 1 ♀ (RER 23 _ LIB 349; UMRI), 24 ix 2023; P 6 c: 1 ♂ (RER 23 _ LIB 348; RER), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF99FFAAFDAC26D192CAFBC0.taxon	discussion	Remarks. Rhyothemis obsolescens is common in swampy habitats, forest edge ponds and slow, open and semi-open streams in swamp; very common in such habitats in the main part of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF99FFAAFDC42359959AF9E8.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF99FFAAFDC42359959AF9E8.taxon	materials_examined	Specimens collected. M 2 e: 1 ♂ (RER 23 _ LIB 249; UMRI), 10 vi 2023; M 3 a: 1 ♀ (RER 20 _ LIB 113; RER), 17 i 2020; 1 ♀ (RER 23 _ LIB 100; BRIN), 10 ii 2023, leg. RER staff; M 7 d: 1 ♂ (RER 23 _ LIB 101; UMRI), 19 ii 2023; M 7 g: 1 ♂ (RER 23 _ LIB 102; RER), 19 ii 2023; P 3 a: 1 ♂ (RER 23 _ LIB 248; BRIN), 30 v 2023; P 5 d: 1 ♂ (RER 23 _ LIB 250; RER), 21 vi 2023; P 6 c: 1 ♀ (RER 23 _ LIB 352; BRIN), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF99FFAAFDC42359959AF9E8.taxon	discussion	Remarks. Another species that is much more common in the Kampar Peninsula than the records above suggest, with individuals frequently seen flying at open drains in the acacia.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF99FFAAFDA5218992EAF810.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF99FFAAFDA5218992EAF810.taxon	materials_examined	Specimens collected. M 5 b: 1 ♂ (RER 23 _ LIB 103; RER), 9 ii 2023; 1 ♀ (RER 23 _ LIB 253; RER), 17 vi 2023; P 1 b: 1 ♂ (RER 23 _ LIB 251; BRIN), 2 vi 2023; 1 ♂ (RER 23 _ LIB 252; UMRI), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF99FFAAFDA5218992EAF810.taxon	discussion	Remarks. Risiophlebia dohrni is most often found in swamp forest but is usually local in occurrence even in such habitat and this is the case in the Kampar Peninsula where it was found at only two sites so far.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFADFDA727F19702FDD7.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFADFDA727F19702FDD7.taxon	materials_examined	Specimens collected. M 5 b: 1 ♀ (RER 23 _ LIB 254; RER), 17 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFADFDA727F19702FDD7.taxon	discussion	Remarks. Recorded from Riau Province for the first time. The single female collected was placing eggs in a mass on hair-like aerial roots hanging from horizonal shoot ca 20 cm above shaded water between a log and the densely vegetated margin of the pool. Ngiam et al. (2024) describe similar oviposition behavior of this species in Singapore.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFADFDA725919395FBE8.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFADFDA725919395FBE8.taxon	materials_examined	Specimens collected. M 2 f: 1 ♀ (RER 23 _ LIB 256: BRIN), 6 vi 2023; M 3 d: 1 ♀ (RER 23 _ LIB 115; BRIN), 17 i 2020; M 8 b: 1 ♀ (RER 20 _ LIB 114; RER), 13 i 2020; P 3 a: 1 ♀ (RER 23 _ LIB 255; UMRI), 30 v 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFADFDA725919395FBE8.taxon	discussion	Remarks. Another species that is more common in the Kampar Peninsula than the number of specimens collected suggests, frequently seen in the late afternoon and at dusk around drains and flooded areas in clearings.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFADFDCF23B494BEF99B.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFADFDCF23B494BEF99B.taxon	materials_examined	Specimens collected. M 3 a: 1 ♀ (RER 20 _ LIB 116; RER), 17 i 2020; M 3 e: 1 ♂ (RER 20 _ LIB 118; UMRI), 24 i 2020; M 7 g: 1 ♂ (RER 20 _ LIB 117; BRIN), 23 i 2020; 1 ♂ (RER 23 _ LIB 257; RER), 15 vi 2023; 1 ♂, 1 ♀ (RER 23 _ LIB 258 – 259; BRIN), in tandem, 15 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFADFDCF23B494BEF99B.taxon	discussion	Remarks. Common at the rain-fed pond beside the Kampar River at location M 7 and probably common at drains in the acacia as well, where Tramea were quite often seen but much less often captured. It cannot however simply be assumed that all red-marked Tramea seen in the Kampar Peninsula are T. transmarina euryale since other similar looking Tramea might occur there, in particular T. phaeoneura Lieftinck, 1953, which has been recorded in Belitung and might well occur in mainland Sumatra and which is most often found at low pH waters in Borneo. Recorded from Riau Province for the first time.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFACFDD12020977FFE34.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFACFDD12020977FFE34.taxon	materials_examined	Specimens collected. M 1 m: 1 ♂ (RER 23 _ LIB 356; RER), 1 x 2023; M 1 o: 1 ♂ (RER 23 _ LIB 357; RER), 1 x 2023; M 4 a: 1 ♂ (RER 23 _ LIB 109; UMRI), 13 ii 2023; 2 ♂♂ (RER 23 _ LIB 262, 264; BRIN), 5 vi 2023; 1 ♂ (RER 23 _ LIB 263; UMRI), 5 vi 2023; 1 ♂ (RER 23 _ LIB 354; BRIN), 27 ix 2023; 1 ♂ (RER 23 _ LIB 355; UMRI), 27 ix 2023; M 4 b: 1 ♂ (RER 23 _ LIB 104; RER), 8 ii 2023; 1 ♂ (RER 23 _ LIB 267; UMRI), 13 vi 2023; M 4 e: 1 ♂ (RER 23 _ LIB 353; UMRI), 27 ix 2023; M 5 b: 2 ♂♂ (RER 23 _ LIB 105, 108; BRIN), 9 ii 2023; 1 ♂ (RER 23 _ LIB 106; UMRI), 9 ii 2023; 1 ♂ (RER 23 _ LIB 107; RER), 9 ii 2023; 1 ♂ (RER 23 _ LIB 268; BRIN), 17 vi 2023; M 5 f: 1 ♂ (RER 23 _ LIB 265; UMRI), 6 vi 2023; M 10 a: 1 ♂ (RER 23 _ LIB 266; BRIN), 8 vi 2023; P 1 b: 1 ♂ (RER 23 _ LIB 260; BRIN), 31 v 2023; 1 ♂ (RER 23 _ LIB 261; UMRI), 31 v 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9EFFACFDD12020977FFE34.taxon	discussion	Remarks. Not found during the unusually dry conditions of January 2020, but when surface water was present in the PSF during the surveys from February 2023 onwards, T. kuekenthali was generally common in the PSF of the Kampar Peninsula.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9FFFACFDC125459713FAA0.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9FFFACFDC125459713FAA0.taxon	materials_examined	Specimens collected. M 1 a: 1 ♂ (RER 20 _ LIB 120; BRIN), 15 i 2020; 1 ♂ (RER 20 _ LIB 123; BRIN), 19 i 2020; 1 ♀ (RER 20 _ LIB 124; UMRI), 19 i 2020; M 1 c: 1 ♂ (RER 20 _ LIB 119; RER), 15 i 2020; M 1 l: 1 ♂ (RER 23 _ LIB 112; BRIN), 15 ii 2023; 1 ♂ (RER 23 _ LIB 361; UMRI), 1 x 2023; M 1 p: 1 ♂ (RER 23 _ LIB 111; UMRI), 12 ii 2023; M 2 e: 1 ♂ (RER 23 _ LIB 209; BRIN), 10 vi 2023; M 3 a: 1 ♂ (RER 20 _ LIB 121; UMRI), 17 i 2020; 1 ♀ (RER 20 _ LIB 122; RER), 17 i 2020; M 5 a: 1 ♂ (RER 20 _ LIB 125; BRIN), 20 i 2020; 1 ♂ (RER 20 _ LIB 128; UMRI), 25 i 2020; 1 ♂ (RER 23 _ LIB 110; BRIN), 9 ii 2023; M 7 a: 1 ♂ (RER 20 _ LIB 126; UMRI), 23 i 2020; 1 ♂ (RER 23 _ LIB 113; UMRI), 19 ii 2023; M 7 b: 1 ♀ (RER 20 _ LIB 127; BRIN), immature, 23 i 2020; M 7 g: 1 ♂ (RER 23 _ LIB 271; BRIN), 15 vi 2023; M 11 a: 1 ♂ (RER 23 _ LIB 270; UMRI), 11 vi 2023; M 12 a: 1 ♂ (RER 23 _ LIB 360; BRIN), 29 ix 2023; M 13 a: 1 ♂ (RER 23 _ LIB 272; RER), 18 vi 2023; P 5 d: 1 ♂ (RER 23 _ LIB 358; RER), 22 ix 2023; P 6 b: 1 ♂ (RER 23 _ LIB 359; RER), 23 ix 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9FFFACFDC125459713FAA0.taxon	discussion	Remarks. The first record of this species from Sumatra was from “ Soeroegoen [Surugun] ”, collected by J. F. Snelleman (Selys 1889) during an expedition to central Sumatra that passed through several of the present day provinces, it is not clear where exactly this location was, but where the locations for other odonate material from the same source can be traced, they are all in present day West Sumatra, so “ Soeroegoen ” might well be in that province.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9FFFACFDAE22F097AFF89F.taxon	description	Selected references:	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9FFFACFDAE22F097AFF89F.taxon	materials_examined	Specimens collected. M 2 f: 1 ♂ (RER 23 _ LIB 114; UMRI), 18 ii 2023; 1 ♂ (RER 23 _ LIB 277; UMRI), at lights at ca. 7 PM, 4 vi 2023, leg. Eco Camp canteen staff; M 3 b: 1 ♂ (RER 20 _ LIB 129; RER), 17 i 2020; M 3 d: 1 ♂ (RER 20 _ LIB 130; BRIN), 17 i 2020; M 7 f: 1 ♀ (RER 23 _ LIB 115; RER), 19 ii 2023; P 2 a: 1 ♂ (RER 23 _ LIB 273; BRIN), 30 v 2023; 1 ♂ (RER 23 _ LIB 274; UMRI), 30 v 2023; P 4 a: 1 ♀ (RER 23 _ LIB 275; RER), found dead in spider web, 31 v 2023; 1 ♀ (RER 23 _ LIB 276; BRIN), 2 vi 2023.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
03DB2A41FF9FFFACFDAE22F097AFF89F.taxon	discussion	Remarks. At least moderately common in the Kampar Peninsula and probably under-recorded because of its mostly crepuscular habits.	en	Dow, Rory A., Gesriantuti, Novia, Iqbal, Muhammad, Lupiyan-Ingdyah, Pungki, Periera, John F., Zulya, Shinta O., Sanders, Bradford M. (2025): Odonata of the Kampar Peninsula: the first systematic survey of Odonata in Peat Swamp Forest in Sumatra. Zootaxa 5732 (1): 1-70, DOI: 10.11646/zootaxa.5732.1.1, URL: https://doi.org/10.11646/zootaxa.5732.1.1
