taxonID	type	description	language	source
03DBDE753342FFB1769D6BF2DE46FCB9.taxon	description	Resurrection of genera within the Goniodes - complex Historically, Goniodes has been the repository of many heterogeneous species of lice parasitic on gamefowl (Galliformes) around the world (e. g., Hopkins & Clay 1952; Price et al. 2003). However, Kéler (1940) regarded the genus as more narrowly defined, comprising only three species, while he placed all other species in different genera. Moreover, as argued by e. g., Gustafsson et al. (2021), there are no morphological characters to reliably separate Goniodes from Goniocotes as defined by Hopkins & Clay (1952) and Price et al. (2003). Clay (1951) had already acknowledged this fact, stating that the two genera “ seem to grade into each other ”, implying that it may be best to synonymise them. Based on a detailed examination of a great number of Goniodes and Goniocotes species, we agree with Kéler’s (1940) assessment, and disagree with Clay’s (1951). The morphological diversity among the species placed in Goniocotes is less extensive and may not justify its division into different genera, although it would make the genus more homogeneous (Gustafsson et al. 2021, 2023, 2024 a, b). However, the same rationale cannot be applied to Goniodes, which comprises around 30 distinct morphological groups of species that can be recognised as genera or subgenera. In our opinion, the following genera should be resurrected from synonymy, largely following Mey (2009):	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753342FFB1769D6BF2DE46FCB9.taxon	description	Resurrecting these genera contributes to establishing a clearer morphological classification of the species of Goniodidae. In addition, several other morphologically distinct groups need to be regarded as genera, and most species need redescription. Hence, we propose six new genera (see below) and give the morphological rationale for resurrecting three other genera (Gonotyles, Oulocrepis, Solenodes). As most species placed in these three genera were previously considered part of Goniodes, we provide comparisons between these genera with Goniodes, to highlight the differences. Detailed morphological arguments for resurrecting the other genera listed above will be published elsewhere (Gustafsson & Johnson, in prep.) but a list of the species we place in the resurrected genera is given in Appendix I. One consequence of this proposal is that of the 100 species listed by Price et al. (2003) under Goniodes, this genus now will comprise only two species, the type species and a new species described here. This is because G. pavonis — the type species of Goniodes designated by Johnston & Harrison (1911) — is a morphologically distinct species. A similar result is the recent removal of a large number of species from the genus Brueelia Kéler, 1936 to other genera (Gustafsson & Bush 2017).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753343FFB1769D6BF8D88EFA9D.taxon	description	Clay (1940) placed the type species of Goniodes in Species Group A, and a small number of species which she claimed are morphologically similar to the type species of Goniodes in Species Group B. Species Group B is here considered to comprise three distinct, new genera: Rhopaloncus, Sulciferodes and Tragoniodes. These genera share some morphological characteristics, including: 1) inverse V- or U-shaped internal sclerite in the female genitalia (not found in Rhopaloncus, but see below for a discussion on the identity of females of this genus); 2) narrow fusion of tergopleurite II and pteronotum medianly at least in males; 3) squamous or reticulate horn on posterior margin of male scape. Besides the fusion of the tergopleurite and pteronotum, those characters are also shared with Goniodes sensu stricto (the one species of Clay’s (1940) species group A), suggesting that these genera may be closely related. However, the antennal morphology, head chaetotaxy, abdominal chaetotaxy, and the male and female genitalia differ markedly among Goniodes sensu stricto and the three new genera, as detailed below.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753343FFB2769D6D9CD998FA79.taxon	type_taxon	Type species: Pediculus pavonis Linnaeus, 1758: 613. By subsequent designation (Johnston & Harrison, 1911: 326).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753343FFB2769D6D9CD998FA79.taxon	description	Description Both sexes. Large chewing lice of the Goniodes body louse ecomorph. Head broader than long, frons gently rounded to somewhat flattened. Marginal carina uninterrupted, terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni prominent, bent strongly posterior to overlap with base of antennae. Antennae sexually dimorphic: in male, scape much swollen and elongated compared to that of female, with one distally squamous process on posterior margin; male pedicel similar in size to that of female, but noticeably curved; male flagellomere I much enlarged compared to that of female and to male flagellomeres II – III, extended distally into curved horn with squamous inner margin. Eyes large. Temples rounded laterally; posterior margin of head with narrowly pointed occipital process. Head chaetotaxy: dsms, ads, pns, pts, avs 2 all mesosetae or longer; mts 1, mts 3 macrosetae; mts 4 – 5 situated on occipital process; male postantennal head with row of sensilla from pns to pts on each side; head sensilla s 1 – 2, s 6 present. Rhombic sclerite small, clearly separated from pronotum. Pronotum flaring posteriorly, with ppss on postero-lateral corner. Pteronotum roughly pentagonal, but with posterior margin more rounded in type species. Postero-lateral corner of pteronotum modified so that both lpts and ipts are roughly lateral, and ipts are more or less directly posterior to lpts; smns with associated sensillum; mpts situated in hyaline tegument posterior to pteronotum. Mesosternum present, metasternum absent; mets and mss present. Proepimera large, curling around coxae II, but not fused medianly. Metepisterna not sclerotised medianly. Abdomen rounded in male, more elongate in female; tergopleurite II not fused to pteronotum in either sex, and tergopleurites II – VIII medianly separated, with section median to spiracular openings conspicuously narrowed in more posterior segments. Male tergopleurite IX fused to central plate, medianly continuous, and extending laterally to reach tergopleurite VIII; central plate overhanging genital opening such that small inner plate and setal row are completely obscured by tergopleurite. Female tergopleurites IX – XI fused together, and medianly continuous. Central sternal plates absent, but abdominal segments II – VI in both sexes with lateral accessory sternal plates. Male abdominal chaetotaxy: tergopleurites II – VI with more than four tcs on each side; tergopleurites VII – VIII with one tcs on each side; tergopleurites II – VIII with multiple tps microsetae on each side, the medial of which may be intercalated with tcs; tergopleurites II – VIII with 2 – 3 psc setae on each side; ventral abdominal segments II – VI with dense rows of sts; on segments VII – VIII only one sts on each side. Female abdominal chaetotaxy: tergopleurites II – VI with more than four tcs on each side; tergopleurites VII – VII with one tcs on each side; tcs microsetae only present on tergopleurites VI – VII, not intercalated; tergopleurites II – VIII with 2 – 3 psc setae on each side; ventral abdominal segments II – VI with median rows of sts, on segment VII only two sts on each side. Neither sex with sternal microsetae. Male subgenital plate large, expanded laterally; female subgenital plate absent, but subvulval plates may be present and postvulval plates present. Male genitalia large, reaching anteriorly to or almost to posterior margin of pteronotum when non-everted. Basal apodeme less than half the length of genitalia, trapezoidal with distally bifurcated median thickening ventrally; postero-lateral corners of basal apodeme extended into large ventral plates. Mesosome large, roughly rectangular or trapezoidal with lateral lobes in proximal end and deeply bifurcated distal end; ventrally with proximal rounded structure (here presumed to be gonopore) and fleshy distal lobes with rugose lateral margins. Parameres long, approximately half the length of genitalia, widening distally with pst 1 – 2 on lateral or postero-lateral margins. Female genitalia with bifid structure (sensu Clay 1940) internally, but without triangular processes laterally. Vulval margin mildly concave to mildly convex, without lateral lobes. Vulval chaetotaxy: mesosetae (presumably vms and vss) clustered laterally, with multiple microsetae (vos and possibly vss) forming vague rows more anteriorly.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753340FFB2769D6E60DEF5F891.taxon	biology_ecology	Type host: Pavo cristatus Linnaeus, 1758 — peacock.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753341FFB3769D68C1DF8FFDF0.taxon	biology_ecology	Type host: Pavo muticus Linnaeus, 1758. Host distribution: Goniodes sensu stricto is known from two species of peacocks (Galliformes: Phasianidae: Pavo Linnaeus, 1758).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753341FFB3769D68C1DF8FFDF0.taxon	distribution	Geographical range: Although Goniodes pavonis has been reported from across the world from captive populations of peacocks (e. g., Adam 2007; González-Acuña et al. 2009; Nasser et al. 2015; Gustafsson et al. 2019 a), the natural range of Goniodes sensu stricto is southern Asia.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753341FFA8769D6A0ED893FCB9.taxon	description	(Figs 1 – 8)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753341FFA8769D6A0ED893FCB9.taxon	materials_examined	Type host: Pavo muticus imperator Delacour, 1949 — green peafowl. Type locality: Jingdong Yi Autonomous County, Yunnan Province, China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753341FFA8769D6A0ED893FCB9.taxon	diagnosis	Diagnosis: Goniodes claytonbushorum can be separated from G. pavonis by the following combination of characters [see Gustafsson et al. (2023: figs 1 – 9) for a detailed description and illustrations of G. pavonis]: head proportionately narrower in G. claytonbushorum (Fig. 4) than in G. pavonis; male scape with distal seta on posterior margin thorn-like in G. pavonis, but slender in G. claytonbushorum (Fig. 4); male os macrosetae in G. pavonis, but mesosetae in G. claytonbushorum (Fig. 4); male occipital process more elongated, with sinuous lateral margin in G. claytonbushorum (Fig. 4); female coni elongated and bluntly rectangular in G. claytonbushorum (Fig. 5), but proportionately shorter and rounder in G. pavonis; posterior margin of pteronotum more concave in G. claytonbushorum (Figs 1 – 2) than in G. pavonis; male tergopleurite IX + X with lateral extension shorter and broader in G. claytonbushorum (Fig. 1) than in G. pavonis; lateral tufts of setae on male tergopleurite IX + X with 1 – 3 mesosetae in G. claytonbushorum (Fig. 1), but with 6 + in G. pavonis; male subgenital plate of more uniform width throughout in G. claytonbushorum (Fig. 1), but narrowed submedianly in G. pavonis; paratergal setae on male tergopleurite III much shorter in G. claytonbushorum (Fig. 1) than in G. pavonis; mesosome more rectangular in G. claytonbushorum (Fig. 6) than in G. pavonis; gonopore symmetrical and U-shaped in G. pavonis, but slightly asymmetrical proximally and narrowed at about half-length in G. claytonbushorum (Fig. 7); pst 1 – 2 situated close together in G. claytonbushorum (Figs 6 – 7), but far apart in G. pavonis; subvulval plates visible but faint in G. claytonbushorum (Fig. 8), but not visible in examined specimens of G. pavonis.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753341FFA8769D6A0ED893FCB9.taxon	description	Descriptions Both sexes. Head rounded rectangular, wider than long (Fig. 4). Frons gently rounded. Marginal carina relatively broad, with sinuous inner margin. Head chaetotaxy as in Fig. 4; os mesosetae; 8 – 12 dorsal postantennal sensilla on each side in arched row between pns; mts 4 – 5 situated on occipital process. Thoracic and abdominal segments and chaetotaxy as in Figs 1, 3. Posterior margin of pteronotum concave in section between ipts and smns. Male. Scape swollen and elongated compared to female, with posterior margin bearing distally squamous horn in proximal half; marginal seta distal to this horn not thorn-like. Pedicel slightly curved. Flagellomere I with long distal extension. Ocular setae similar to pos in length. Temples rounded, with slight restriction at mts 3, and occipital process with sinuous lateral margin. In both holotype and one paratype the occipital process is stunted on one side, but the opposite side and in other paratypes is symmetrical as illustrated. Tergopleurites II – IV may have tps microsetae intercalated with tcs medianly. Tergopleurite IX + X extended antero-laterally to reach tergopleurite VIII, with sinuous posterior and anterior margins. Inner margin of genital opening hidden under posterior margin of tergopleurite IX + X, simple and rounded with row of microsetae (Fig. 2) and not clearly visible sclerite, but with vague thickening as indicated by dotted line in Fig. 2. Distal end of abdomen soft, and compressed in examined specimens, and here illustrated approximately, based on the holotype. Basal apodeme trapezoidal, widening distally (Fig. 6), with bifurcated median thickening on ventral side not recurving anteriorly (Fig. 7). Mesosome roughly rectangular dorsally, with antero-lateral bulges slender, with distal ends slightly squamous, and close to main part of mesosome. Gonopore asymmetrical, with lateral margins concave in anterior half. Distal lobes of ventral mesosome slender, with distinct rugose areas laterally. Parameres with elongated, slender heads (Fig. 6), slender for most of length, but widening suddenly in distal fourth, with pst 1 – 2 situated closely together. Measurements as in Table 1. Female. Antennae slender (Fig. 5). Coni elongated and roughly rectangular, with slight kink on lateral margin near distal end. Temples somewhat flattened between mts 1 – 3, and occipital process less sinuous than in male. Only tergopleurites VI – VIII with tps microsetae, not intercalated. Abdominal segment IX – XI with lateral setae situated in narrow hyaline groove. Vulval margin straight to slightly convex (Fig. 8), with 10 – 12 long, slender setae laterally on each side, 8 – 12 shorter setae on each side anterior to these, and 7 – 8 short, slender setae on each side forming vaguely convergent lines. Bifid inner structure as in Fig. 8. Measurements as in Table 1.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753341FFA8769D6A0ED893FCB9.taxon	etymology	Etymology: This species is dedicated to the Clayton / Bush Lab, University of Utah, Salt Lake City, U. S. A., including all its past and present members. For over 20 years, this lab has researched the ecology and behaviour of chewing lice, their interactions with their bird hosts and the consequences of these interactions, producing a steady stream of unprecedented insights into how lice live and evolve. In short, the Clayton / Bush lab is truly the “ peacock in the global aviary ” of louse research, spreading its radiant tail of amazing colours. Also, the first author wishes to express his profound gratitude for having the privilege of working in this lab for 4.5 years, looking at and talking about lice on a daily basis.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753341FFA8769D6A0ED893FCB9.taxon	materials_examined	Material examined Types: Ex Pavo muticus imperator: Holotype ♂, Jingdong Yi Autonomous County, Yunnan Province, China, Aug. 1964, [coll. Zhixiang Li, but not written on slide], box E 0026200, slide 76 (NHMC) [marked with black dot on slide]. Paratypes: 4 ♂, 9 ♀, Jingdong Yi Autonomous County, Yunnan Province, China, Aug. 1964, coll. Zhixiang Li, box E 0026011, slide 37; box E 0026135, slide 22; box E 0026200, slides 75 – 76 (NHMC).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753341FFA8769D6A0ED893FCB9.taxon	discussion	Morphological separation between Goniodes and Goniocotes The genera Goniodes and Goniocotes as listed by Price et al. (2003) are two of the most speciose genera within Goniodidae. However, Clay (1951) suggested that there are no morphological characters separating these genera, and Gustafsson et al. (2021) noted that all of the characters proposed to separate them are inadequate. Whereas Goniocotes sensu Price et al. (2003) is relatively homogeneous morphologically, Goniodes sensu Price et al. (2003) is not, as any character regarded as diagnostic for Goniocotes is found in at least some species of Goniodes. The separation of Dictyocotes Kéler, 1940 and Pavoniocotes Gustafsson et al., 2023 from Goniocotes contributed to reduce the morphological variation within Goniocotes (Gustafsson et al. 2023, 2024 b) but did little to provide a clear separation of Goniocotes from Goniodes. By limiting Goniodes to the type species and G. claytonbushorum, we are able to provide clear morphological limits between these two genera for the first time. However, Goniocotes chapini Clay, 1938 does not fit into our definition of Goniocotes here, and likely deserves recognition as a separate genus or subgenus; a redescription of this species is necessary before such a taxon can be proposed. Hence, for the purposes of this study, G. chapini is not included in Goniocotes. Moreover, some recently described species, such as Goniocotes devius Gustafsson et al., 2023, differ somewhat from Goniocotes sensu stricto and may deserve recognition at the subgenus or genus level, but these differences are not so pronounced to contradict the comparison given below. Goniocotes can be separated from Goniodes as defined here by the following characters [see Gustafsson et al. (2023, 2024 a, b) for recent descriptions and illustrations of various species of Goniocotes]: male antennae sexually dimorphic in Goniodes (Figs 4 – 5), but monomorphic in Goniocotes; coni bent posteriorly in Goniodes (Figs 4 – 5), but not in Goniocotes; occipital process narrow, with both mts 4 – 5 situated subapically on process in Goniodes (Fig. 4), but process broader and blunter, with only mts 5 situated apically in Goniocotes; male postantennal head with U-shaped band of multiple sensilla in Goniodes (Fig. 4), but with at most s 1 – 7 present on the dorsal head, and no sensilla forming U-shaped band in Goniocotes; postero-lateral corner of pteronotum flattened so that both lpts and ipts are lateral in Goniodes (Figs 1, 3), but rounded and only lpts lateral in Goniocotes; smns submarginal with clearly associated sensillum in Goniodes (Figs 1, 3), but more central, without sensillum, in Goniocotes (in some species absent, or with patches of multiple smns on each side); mesosternum present in Goniodes (Figs 1, 3), but absent in Goniocotes (but metepisterna may be fused medianly in some Goniocotes, giving the appearance of a mesosternum); male tergopleurite IX fused to central plate in Goniodes (Fig. 1), but separate from central plate (but may overlap) in Goniocotes; each of abdominal segments III – V with multiple sts on each side in Goniodes (Figs 1, 3), but with only one sts on each side in Goniocotes; male genitalia with distinct parameres and mesosome in Goniodes (Figs 6 – 7), but male genitalia solenoid in Goniocotes; bifid structure present internally in female genitalia (Fig. 8), but not present in Goniocotes; female vulval chaetotaxy with lateral tufts of long setae, no marginal setae medially and no thorn-like setae in Goniodes (Fig. 8) but with vms distributed more or less evenly along vulval margin and vss thorn-like in Goniocotes. More distinguishing characters may become relevant after Goniocotes has been thoroughly revised. For instance, lateral accessory sternal plates are present in Goniodes (Figs 1, 3) and some species of Goniocotes, but not in the type species Goniocotes gallinae (De Geer, 1778). Similarly, neither sex of Goniocotes gallinae has as many dorsal abdominal setae as Goniodes, but other species currently placed in Goniocotes do; however, in these species tps microsetae are not evident, whereas such setae are found on at least some tergopleurites in Goniodes (Figs 1, 3). In some species of Goniocotes, the length of the os is sexually dimorphic, being macrosetae in males and micro- or mesosetae in females, but in Goniodes and Goniocotes gallinae this seta is sexually monomorphic, a mesoseta in both sexes. In Goniodes the rhombic sclerite is always separate from the pronotum (Figs 1, 3), whereas in Goniocotes gallinae, these are fused, and this character varies among other Goniocotes.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335AFFA9769D6BF8DEE6FA25.taxon	type_taxon	Type species: Goniodes meinertzhageni Clay, 1940: 9.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335AFFA9769D6BF8DEE6FA25.taxon	diagnosis	Diagnosis. Sulciferodes (Figs 9 – 15) is close to Goniodes sensu stricto, based on the following shared characters: female genitalia with bifid structure (Figs 8, 15); mesosternum present (Figs 1, 3, 9, 10); structure of the male antenna (Figs 4, 11). However, the relationships between Sulciferodes, Tragoniodes gen. nov. and Rhopaloncus gen. nov. require further study, because some of these characters are also found in these genera. Sulciferodes can be separated from Goniodes sensu stricto by the following characters: occipital process clearly present and somewhat pointed, bearing mts 4 – 5 in Goniodes (Fig. 4), but absent or at most a slight bulge in Sulciferodes (Figs 9 – 11); coni strongly curved in Goniodes (Figs 4 – 5), but not in Sulciferodes (Figs 11 – 12); metepisterna present and latitudinal plates in Sulciferodes (Figs 9 – 10), but absent in Goniodes (Figs 1, 3); male pteronotum fused to tergopleurite II medianly in Sulciferodes (Fig. 9), but not in Goniodes (Fig. 1); both sexes with multiple tcs on each side in Goniodes (Figs 1, 3), but with only one tcs on each side in Sulciferodes (Figs 9 – 10); male tergopleurite IX fused to central plate in Goniodes (Fig. 1), but plates separate in Sulciferodes (Fig. 9); lateral accessory sternal plates present in both sexes in Goniodes (Figs 1, 3), but absent in males of Sulciferodes (Fig. 9); mesosome and parameres fused to basal apodeme in Sulciferodes (Figs 13 – 14), but not fused in Goniodes (Figs 6 – 7); genitalia overall pseudo-solenoid in Sulciferodes (Figs 13 – 14), with mesosome simple and elongated and parameres seemingly flexible and attenuated distally, whereas in Goniodes, genitalia are larger and bulkier, with more complicated mesosome and distally expanded parameres (Figs 6 – 7); female subgenital plate absent in Goniodes (Fig. 8), but present in Sulciferodes (Fig. 15); vulval margin at most slightly concave in Goniodes (Fig. 8), but deeply concave in Sulciferodes (Fig. 15).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335AFFA9769D6BF8DEE6FA25.taxon	description	Description Both sexes. Large chewing lice of the Goniodes body louse ecomorph (Figs 9 – 10). Head broader than long (Fig. 11), frons gently rounded to somewhat flattened. Marginal carina uninterrupted, terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni prominent, not bent posteriorly. Antennae sexually dimorphic (Figs 11 – 12): in male, scape much swollen and elongated compared to that of female, with one distally squamous process on posterior margin; male pedicel similar to that of female in size, but noticeably curved; male flagellomere I much enlarged compared to that of female and to male flagellomeres II – III, extended distally into curved horn with squamous inner margin. Eyes large. Temples somewhat angular; posterior margin of head without pronounced occipital process, but bulging slightly at site of mts 4 – 5, particularly in female. Head chaetotaxy: dsms, ads, pas, pns, pts, pos, avs 2 – 3 all mesosetae; os sexually dimorphic, longer in male than in female; male postantennal head with row of sensilla from pns to pts on each side; head sensilla s 1 – 2, s 6 present. Rhombic sclerite moderate, clearly separated from pronotum (Figs 9 – 10). Pronotum flaring posteriorly, with ppss on postero-lateral corner. Pteronotum roughly pentagonal, in male fused submedianly to median end of tergopleurite II. Postero-lateral corner of pteronotum rounded, with lpts only setae on lateral margin; ipts intermediate; smns submedial, sexually dimorphic with that of female being microseta and that of male being macroseta, in both sexes without associated sensillum; mpts situated in hyaline tegument posterior to pteronotum. Mesosternum present, metasternum absent in male, but small, poorly sclerotised plate seemingly present in female; mets and mss present. Proepimera large, curling around coxae II, but not fused medianly. Metepisterna broadly sclerotised, but not connected to more lateral plates. Abdomen rounded in male, more elongate in female (Figs 9 – 10). Tergopleurites II – VIII medianly separated, with section medial to spiracular openings conspicuously narrowed in tergopleurite VIII. Male tergopleurite IX not fused to median plate, but these overlap, and median plate has distinct shape with postero-lateral hook. Female tergopleurites IX – XI fused together, and medianly continuous. Central sternal plates absent, but abdominal segments II – VI in female with lateral accessory sternal plates; no such plates visible in examined males. Abdominal chaetotaxy of both sexes: tergopleurites II – VIII with one tcs on each side, flanked laterally by row of tps microsetae; male tergopleurite II and VII with two psc on each side, tergopleurites III – VI with one psc on each side, and tergopleurite VIII with one psc macroseta and multiple shorter psc on each side in male, and multiple psc in female; ventral abdominal segments II – VI with median row of sts, on segments VII – VIII with only one sts on each side; female also with rows of microsetae following posterior margin of lateral accessory sternal plate on segments II – VI, with rows of segments V – VI extending medianly to be intercalated with longer sts. Male subgenital plate large, medianly continuous, expanded laterally. Female subgenital and subvulval plates present. Male genitalia large, reaching anteriorly to around tergopleurite III in non-everted state. Male genitalia pseudo-solenoid (Figs 13 – 14). Basal apodeme partially fused to mesosome and parameres, seemingly consisting of approximately half the length of genitalia, and with distally bifurcated structure overlapping mesosome. Mesosome elongated, narrowed distally, on ventral surface with what may be lateral sensilla. Parameres elongated, slender, somewhat flexible distally, with pst 1 – 2 on lateral margins widely separated. Female genitalia with bifid structure (sensu Clay 1940) internally, but without triangular processes laterally (Fig. 15). Subgenital plates elongated, fused only at vulval margin, and with distinct longitudinal furrows in anterior half. Vulval margin deeply concave, with marginal mesosetae (presumably vms) forming rows laterally. A vague band of microsetae across the genital area may comprise the vss, whereas a median longitudinal row of setae likely represents the vos. Subvulval plates elongated, curved.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335BFFAC769D6E0CD938FF1D.taxon	description	(Figs 9 – 15)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335BFFAC769D6E0CD938FF1D.taxon	biology_ecology	Type host: Pavo cristatus Linnaeus, 1758 — peacock. Host distribution: Known from one species of Pavo (Galliformes: Phasianidae).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335BFFAC769D6E0CD938FF1D.taxon	distribution	Geographical range: India.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335BFFAC769D6E0CD938FF1D.taxon	etymology	Etymology: This name is derived from “ sulcus ”, Latin for “ furrow ”, “ ferō ”, Latin for “ carry ”, and the arbitrary ending “ ides ”. This refers to the striated pattern of the female subgenital plate. Gender masculine.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335BFFAC769D6E0CD938FF1D.taxon	discussion	Remarks: Sulciferodes is currently monotypic. For more details on the morphology of this genus, see the redescription of the type species by Gustafsson et al. (2023) Note that Gustafsson et al. (2023) inadvertently omitted the metepisternum in the male and the smns in the female, characters which we confirm here as present.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335EFFAE769D691CD82CFA79.taxon	type_taxon	Type species: Goniocotes spinicornis Nitzsch [in Giebel], 1866: 389.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335EFFAE769D691CD82CFA79.taxon	diagnosis	Diagnosis. The presence of a bifid structure in the female genitalia, the squamous horn of the male scape, and the presence of numerous dorsal head sensilla situated in a roughly U-shaped arch in at least the male indicate that Tragoniodes is closely related to Goniodes sensu stricto, Sulciferodes and Rhopaloncus gen. nov. Two further characters in the male suggest that Goniodes sensu stricto may be the closest relative of Tragoniodes: the fusion of male tergopleurite IX with the central plate of abdominal segment IX + X and the presence of a bifid structure on the distal basal apodeme. Tragoniodes can be separated from Goniodes sensu stricto by the following characters: coni not bent posteriorly in Tragoniodes (Figs 18 – 19), but bent in Goniodes (Figs 4 – 5); dorsal head surface of both sexes with more than 30 sensilla on each side in Tragoniodes (Fig. 18), but with less than 20 sensilla in male and 3 – 4 in female in Goniodes (Figs 1, 3); pteronotum fused with tergopleurite II in at least male in Tragoniodes (Fig. 16), but not in Goniodes (Fig. 1); basal apodeme comprising less than half the length of genitalia, broadening markedly in distal end, and with ventral subsidiary plates in distal corners in Goniodes (Figs 6 – 7), but comprising more than half the length of genitalia, not widening or widening only slightly in distal end, and without subsidiary plates in Tragoniodes (Figs 20 – 21); bifid structure of distal basal apodeme small, slender, and without lateral lobes in Goniodes (Fig. 7), but large, broad, and with prominent lateral lobes in Tragoniodes (Fig. 21); mesosome roughly rectangular, with ventral section bearing fleshy, laterally rugose, lobes laterally and more sclerotised, slender appendages distally, and with gonopore prominent ventrally in Goniodes (Figs 6 – 7), but Y-shaped, without any ventral lobes [except Tragoniodes tragopan (Clay, 1940); see below], and with gonopore not evident in Tragoniodes (Figs 20 – 21); parameres widening distally with pst 1 – 2 on lateral margins in Goniodes (Fig. 7), but tapering distally with more than eight sensilla on dorsal surface in Tragoniodes (Fig. 20); genital sac absent in Goniodes (Figs 6 – 7), but present in Tragoniodes (Figs 20 – 21); setal row of vulval margin medianly continuous in Tragoniodes (Fig. 22), but interrupted medianly in Goniodes (Fig. 8); distal subgenital area with microreticulation and post-vulval area with dense covering of microtrichia in Tragoniodes (Fig. 22), but without such characters in Goniodes (Fig. 8).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335EFFAE769D691CD82CFA79.taxon	description	Description Both sexes. Large chewing lice of the Goniodes body louse ecomorph. Head broader than long, frons gently rounded. Marginal carina uninterrupted, terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni prominent, in males may be extended posteriorly to overlap scape on dorsal side; in females with attenuated point. Antennae sexually dimorphic: in male, scape much swollen and elongated compared to that of female, with one squamous process on posterior margin and posterior margin squamous; male pedicel similar to that of female in size, but noticeably curved (female pedicel may also be distinctly swollen or hooked distally); male flagellomere I much enlarged compared to that of female and to male flagellomeres II – III, extended distally into curved horn with squamous inner margin. Eyes large. Temples rounded laterally in male, extended into acute lateral angles in female; posterior margin of head without occipital process in male, but process present in female, with only mts 5 situated on process. Head chaetotaxy: dsms, ads, pns, pts, avs 2 all mesosetae or longer; mts 1, mts 3 macrosetae; os macrosetae in male, but mesosetae in female; much of central head surface in both sexes with multiple (in some species over 100) setae arranged roughly in a U- or V-shaped pattern; head sensilla s 1 – 2 present. Rhombic sclerite large, clearly separated from pronotum. Pronotum flaring posteriorly, with ppss on postero-lateral corner. Pteronotum roughly pentagonal, fused medianly with median end of tergopleurite II (except apparently in females of Tragoniodes yinhe sp. nov.). Postero-lateral corners of pteronotum rounded, with only lpts situated on corner, and ipts more median; smns with associated sensillum; mpts variable in position. Meso- and metasterna absent (poorly sclerotised mesosternum may be present in some specimens); mets and mss present. Metepisternum not sclerotised. Abdomen rounded in male, more elongate in female. Tergopleurites II – VIII medianly separated, with section median to spiracular openings conspicuously narrowed in more posterior segments. Male tergopleurite IX fused to central plate, medianly continuous, and extended laterally to reach tergopleurite VIII. Female tergopleurites IX – XI fused together, and medianly continuous. Central sternal plates absent, but abdominal segments II – VI in both sexes with lateral accessory sternal plates. Male abdominal chaetotaxy: tergopleurites II – VII with 1 – 2 tcs on each side, fewer and often shorter in more posterior segments, flanked by rows of tps microsetae that may be intercalated with tcs medial to tergopleurites; tergopleurite VIII with dense row of tps mesosetae along posterior margin; tergopleurites II – VII with 3 – 4 psc on each side, tergopleurite VIII with pcs as in more anterior segments or indistinguishable from tps row; ventral abdominal segments II – VI with dense rows of sts; on at least segments IV – VI with rows of microsetae that may be intercalated with sts centrally; on segments VII – VIII only one ventral sts on each side. Female abdominal chaetotaxy: tergopleurite II with multiple (≥ 5) tcs on each side, tergopleurites III – VIII with gradually fewer tcs on each side, with tergopleurites VII – VIII (in some species also VI) with only one tcs in each side; rows of tps microsetae present on segments II – VIII, with median microsetae typically intercalated with tcs; tergopleurites II – VII each with 2 – 4 psc on each side; ventral side of abdominal segments II – VI with 2 – 6 central sts on each side, and with partially intercalated rows of microsetae on at least segments III – VI; on segments VII – VIII only one ventral sts on each side. Male subgenital plate large, expanded laterally; female subgenital plate absent. Male genitalia large, reaching at least to segment IV when non-everted. Basal apodeme long, comprising approximately two-thirds the length of genitalia, seemingly fused to mesosome and parameres, with prominent bifid structure in distal end, possibly the gonopore or bifurcated structure seen in previous two genera; in all species, this structure has additional lateral lobes that extend to lateral margins of basal apodeme. Mesosome simple, roughly Y-shaped with distal end pointed; in Tragoniodes tragopan (Clay, 1940), apparently with broad rounded lateral flaps not present in other species. Parameres shorter than mesosome, broad proximally and tapering to distal tips; dorsal surface of parameres with multiple (more than eight) sensilla each. Genital sac present, either largely distal to mesosome or overlapping this; sac with bar-shaped lateral sclerites in distal end, and intensely rugose areas throughout (Figs 20 – 21, 27 – 28). Female genitalia with bifid structure (sensu Clay 1940) internally, but without triangular processes laterally. Vulval margin slightly concave to slightly convex, without lateral lobes; central section of subgenital area near vulval margin with distinct micro-reticulation. Vulval chaetotaxy: vulval margin with dense row of marginal setae (vms + vss?) and subgenital area with multiple scattered microsetae (vss + vos?). Subgenital plates strong curved; postvulval plates present. Much of post-vulval area covered in d. ense microtrichia.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335CFFAE769D6E74DFACF95F.taxon	biology_ecology	Type host: Tragopan temminckii (Gray, 1831) — Temminck’s tragopan.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335CFFAE769D6EA8DF44F81D.taxon	biology_ecology	Type host: Tragopan satyra (Linnaeus, 1758) — satyr tragopan.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFAF769D68C1DFA4FED9.taxon	biology_ecology	Type host: Tragopan melanocephalus (Gray, 1829) — western tragopan.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFAF769D69D6DE36FB5A.taxon	biology_ecology	Type host: Tragopan caboti (Gould, 1857) — Cabot’s tragopan. Host distribution: Species of Tragopan Cuvier, 1829 (Galliformes: Phasianidae).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFAF769D69D6DE36FB5A.taxon	distribution	Geographical range: China, Himalayas, parts of Southeast Asia.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFAF769D69D6DE36FB5A.taxon	etymology	Etymology: The name Tragoniodes is a portmanteau of the host genus Tragopan — itself derived from “ trágos ” (τρᾰ ́ ΓΟΣ), ancient Greek for “ male goat ” and “ Pán ” (Πᾱ ́ ν), the Greek god of shepherds and nature — and the genus Goniodes derived from “ gōnía ” (Γωνία), ancient Greek for “ angle ”, referring to the angular temples of many species in the genus. Gender masculine.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFAF769D69D6DE36FB5A.taxon	discussion	Remarks: A recently illustrated redescription of the type species of Tragoniodes, T. spinicornis, is available in Gustafsson et al. (2024 a: 534 – 535, figs 674 – 680). We tentatively accept the synonymy of Goniodes spinicornis blythi under the nominate subspecies, but note that the original description of this subspecies by Eichler (1950) is inadequate and needs redescription. No specimens of Tragoniodes tragopan have been examined. Clay (1940: fig. 9 d) described the bifid structure in the female genitalia of T. tragopan as more broadly rounded than in all other species, and the mesosome with small, rounded flaps on each side in (Clay 1940: fig. 11 c). Photos of the male genitalia of the holotype of T. tragopan available from the NHML’s homepage (https: // data. nhm. ac. uk) confirm the presence of these flaps, and no similar structure has been found in any of the other species examined. Furthermore, the distal section of the male scape in T. tragopan bulges markedly in photos of the holotype, similar to those of Rhopaloncus gen. nov. (see below), but this feature cannot be seen in any of the other species of Tragoniodes. Although T. tragopan needs redescription, it is clear that it is morphologically distinct and cannot be confused with any of the species of the new genus.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFA3769D6D59D978FA79.taxon	description	(Figures 16 – 22)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFA3769D6D59D978FA79.taxon	materials_examined	Type host: Tragopan temminckii (Gray, 1831) — Temminck’s tragopan. Type locality: Sichuan Province, China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFA3769D6D59D978FA79.taxon	diagnosis	Diagnosis. Tragoniodes gallowayi can be separated from T. spinicornis by the following characters: head proportionately wider with shorter preantennal area in T. gallowayi (Fig. 18) than in T. spinicornis; female pedicel slender with distal bulge in T. spinicornis, but more evenly broad in T. gallowayi (Fig. 19); male abdominal segment VI with two tcs on each side in T. gallowayi (Fig. 16), but with one tcs on each side in T. spinicornis; female tergopleurites II – IV each with more tcs on each side (II: 7; III 5 – 6; IV: 6 – 7) in T. spinicornis, than in T. gallowayi (Fig. 17; II: 5; III – IV: 3 each); bifid structure of ventral basal apodeme with concave proximal margin and longer distal appendages in T. gallowayi (Fig. 21), but with bilobed anterior margin and shorter distal appendages in T. spinicornis; vulval margin concave in T. spinicornis, but convex in T. gallowayi (Fig. 22).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFA3769D6D59D978FA79.taxon	description	Descriptions Both sexes. Head relatively wide, preantennal area rather short (Fig. 18); frons gently rounded. Marginal carina narrow, with sinuous inner margin. Head chaetotaxy as in Figs 17 – 18; os sexually dimorphic; both sexes with extensive dorsal head sensilla (more than 30 on each side) roughly forming a U-shape across much of head. Temples sexually dimorphic. Thoracic and abdominal segments and chaetotaxy as in Figs 16 – 17. Male. Scape swollen and elongated compared to female, with posterior margin bearing squamous horn in proximal half (Fig. 18); marginal seta distal to this horn thorn-like, and much of posterior margin of scape distal to horn also squamous; pedicel narrowing slightly distally, somewhat curved; flagellomere I much extended distally into curved horn with squamous inner surface. Ocular seta long, similar to mts 1 and mts 3 in length. Temples roughly rounded, but with irregular margin; occipital process absent. Abdominal chaetotaxy (Fig. 16): tergopleurites II – Vi with two tcs on each side; tergopleurite VII with one tcs on each side; tergopleurites II – VII with numerous tps microsetae, the medial of which may be intercalated with tcs; tergopleurite VIII with dense row of short setae, presumably comprising the tps and tcs; tergopleurites II – VII with 3 – 4 psc on each side, and tergopleurite VIII with lateralmost two setae in row longer than others; ventral surface of abdomen with 3 – 4 sts on each side of segments III – VI, one central sts on each side on segments VII – VIII, and rows of microsetae on segments II – VI, some of which may be intercalated with sts. Basal apodeme long, clearly more than half the length of genitalia (Fig. 20). Bifid structure with concave anterior margin, roughly triangular lateral lobes, and long distal appendages roughly parallel and reaching to distal half of parameres (Fig. 21). Mesosome comparatively slender. Measurements as in Table 1. Female. Antennae slender, with pedicel almost as broad as scape and somewhat bulbous distally (Fig. 19). Coni short (Fig. 19). Ocular seta short, similar to pos in length (Fig. 17). Temples expanded into acute angles; occipital process present. Abdominal chaetotaxy (Fig. 17): tcs: II: 4 – 5 on each side, III – IV: 3 on each side, V: two on each side, VI – VIII: one on each side; tergopleurites II – VIII with tps microsetae in rows, the medial of which may be intercalated with tcs; tergopleurites II – VIII with 3 – 4 psc on each side, the median-most 1 – 2 are microsetae; ventral surface of abdomen with 2 – 3 sts on each side of segments III – IV, four sts on each side of segments V – VI, and one sts on each side on segments VII – VIII; rows of microsetae, some of which are intercalated with sts, on segments II – VII, separated into setae following posterior margin of lateral accessory sternal plates and more central ones. Vulval margin slightly convex centrally (Fig. 22), with 15 – 23 marginal setae, the medial of which are shorter and sometimes more spike-liken than lateral ones, and 16 – 24 scattered microsetae in subgenital area. Bifid inner structure as in Fig. 22. Measurements as in Table 1.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFA3769D6D59D978FA79.taxon	etymology	Etymology: The specific epithet is in honour of our colleague Terry D. Galloway (University of Manitoba, Canada), in recognition of his long-standing research on the lice of Canada and Manitoba in particular, producing novel insights on the biology, ecology and population dynamics of lice from several hosts.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75335DFFA3769D6D59D978FA79.taxon	materials_examined	Material examined Types: Ex Tragopan temminckii: Holotype ♂, Szechwan [= Sichuan, China], Apr. 1908, coll. R. Meinertzhagen, 3756, NHMUK 010677091 (NHML) [marked with black dot on slide]. Paratypes: 3 ♂, Szechwan [= Sichuan, China], Apr. 1908, coll. R. Meinertzhagen, 3756, NHMUK 010677091 – 2 (NHML). 8 ♂, 7 ♀, 4 nymphs, Assam, India (Zoo), no date, coll. R. Meinertzhagen, 10868, NHMUK 010677090, NHMUK 010677093 – 94 (NHML).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753351FFA7769D6DB8DFA3F991.taxon	description	(Figures 23 – 29)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753351FFA7769D6DB8DFA3F991.taxon	materials_examined	Type host: Tragopan caboti (Gould, 1857) — Cabot’s tragopan. Type locality: Zhejiang Province, China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753351FFA7769D6DB8DFA3F991.taxon	diagnosis	Diagnosis. Tragoniodes yinhe can be separated from T. spinicornis by the following characters: female pedicel widened slightly distally in T. spinicornis, but developed distally into clear hook in T. yinhe (Fig. 26); male scape proportionately broader at base in T. yinhe (Fig. 25) than in T. spinicornis; male temples less expanded laterally in T. yinhe (Fig. 25) than in T. spinicornis; female tergopleurites III – IV with only two tcs on each side in T. yinhe (Fig. 24), but with 5 – 7 tcs on each side in T. spinicornis; bifid structure of distal basal apodeme with larger lateral lobes and entirely divergent distal appendages in T. yinhe (Fig. 28), but with smaller lateral lobes and distal ends of distal appendages parallel in T. spinicornis; mesosome proportionately larger and broader in T. yinhe (Fig. 27) than in T. spinicornis.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753351FFA7769D6DB8DFA3F991.taxon	description	Descriptions Both sexes. Head relatively narrow (Fig. 25); frons gently rounded. Marginal carina narrow, with sinuous inner margin. Head chaetotaxy as in Figs 23 – 24; os sexually dimorphic; both sexes with extensive dorsal head sensilla (more than 30 on each side) roughly forming a U-shape across much of head. Temples sexually dimorphic. Thoracic and abdominal segments and chaetotaxy as in Figs 23 – 24. Male. Scape swollen and elongated compared to female, with posterior margin bearing squamous horn in proximal half (Fig. 25); marginal seta distal to this horn somewhat stout, and much of posterior margin of scape distal to horn also squamous; pedicel curved; flagellomere I much extended distally into curved horn with squamous inner surface. Ocular seta long, but shorter than mts 1 and mts 3. Temples roughly rounded, not expanded much lateral to eye; occipital process absent. Abdominal chaetotaxy (Fig. 23): tcs: II and IV – VII with one on each side, III – IV with two on each side; tergopleurites II – VII with numerous tps microsetae, the medial of which may be intercalated with tcs; tergopleurite VIII with dense row of short mesosetae, the medial of which are markedly shorter than the lateral ones, possibly representing a division between short tcs and longer tps; tergopleurite II with 4 – 5 psc on each side, tergopleurites III – VII with 2 – 3 psc on each side, and tergopleurite VIII with lateralmost seta in row longer than others; ventral surface of abdomen with 3 – 5 sts on each side of segments III – VI, 0 – 2 sts on each side on segments VII – VIII, and rows of microsetae on segments III – VI, some of which may be intercalated with sts. Basal apodeme long, clearly more than half the length of genitalia (Fig. 27). Bifid structure with trilobed anterior margin, roughly triangular lateral lobes, and short distal appendages clearly divergent and reaching less than half length of parameres (Fig. 28). Mesosome proportionately large and wide. Measurements as in Table 1. Female. Antennae slender, with distal pedicel formed into hook on inner margin (Fig. 26). Coni elongated (Fig. 24). Ocular seta short, but longer than pos (Fig. 24). Temples expanded into acute angle; occipital process present. Abdominal chaetotaxy (Fig. 24): tcs: III with 5 – 7 on each side; III – V with two on each side; VI – VIII with one on each side; tergopleurites II – VIII with tps microsetae in rows, the medial of which may be intercalated with tcs; tergopleurites II – VIII with 2 – 3 psc macrosetae and 1 – 2 psc microseta on each side; ventral surface of abdomen with 4 sts on each side of segments II – VI, and one sts on each side of segments VII – VIII; rows of microsetae, some of which are intercalated with sts, on segments III – VII. Vulval margin more or less flat (Fig. 29), with 15 – 20 marginal setae, the medial of which are shorter and may be somewhat stouter than lateral ones, and 40 – 45 scattered microsetae in subgenital area. Bifid inner structure as in Fig. 29.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753351FFA7769D6DB8DFA3F991.taxon	etymology	Etymology: The specific epithet is derived from “ yínhé ” (DZñ), the Mandarin name for the Milky Way, referring to the dense band of dorsal head sensilla particularly evident in the male of this species, similar to a galaxy.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753351FFA7769D6DB8DFA3F991.taxon	materials_examined	Material examined Types: Ex Tragopan caboti: Holotype ♂, Zhejiang Province, China, 8 Dec. 1990, collector unknown, box E 0026010, slide 79 (NHMC) [marked with black dot on slide]. Paratypes: 14 ♂, 22 ♀, same data as holotype, box E 0026010, slides 77 – 80, 83 – 84 (NHMC). 1 ♂, 4 ♀, Fujian Province, China, 16 Dec. 1988, collector unknown, box E 0026010, slides 87, 100 (NHMC). 3 ♂, 6 ♀, Zhejiang Province, China, 17 Jun. 1984, collector unknown, box E 0026010, slides 91 – 92, 95 – 98 (NHMC). 1 ♂, South China, Jan. 1898, coll. R. Meinertzhagen, 1830, NHMUK 010677069 (NHML). 2 ♂, Fokien [= Fujian, China], Aug. 1908, coll. R. Meinertzhagen, 3761, NHMUK 010677068 (NHML). 3 ♂, 8 ♀, 4 nymphs, China, no date or collector, NHMUK 010677066 – 67 (NHML).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753355FF99769D6E91DDE1FE81.taxon	type_taxon	Type species: Goniodes megaceros Kellogg & Paine, 1914: 227.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753355FF99769D6E91DDE1FE81.taxon	diagnosis	Diagnosis. Rhopaloncus can be separated from Goniodes sensu stricto by the following characters (see below for comments on the identity of female Rhopaloncus): coni bent posteriorly in Goniodes (Figs 4 – 5), but not in Rhopaloncus (Figs 32 – 33); male scape with one distally squamous process in Goniodes (Fig. 4), but with two such processes in Rhopaloncus (Fig. 32); female with rounded temples in Goniodes (Fig. 3), but with temples extended laterally in Rhopaloncus (Fig. 31); postero-lateral corners of pteronotum flattened and both lpts and ipts lateral in Goniodes (Figs 1, 3), but rounded and only lpts lateral in Rhopaloncus (Figs 30 – 31); male pteronotum fused to tergopleurite II in Rhopaloncus (Fig. 30), but not in Goniodes (Fig. 1); male tergopleurite IX fused to central plate in Goniodes (Fig. 1), but not in Rhopaloncus (Fig. 30); ventral side of abdomen without rows of microsetae in Goniodes (Figs 1, 3), but with such rows in Rhopaloncus (Figs 30 – 31); female subgenital plate absent but bifid structure present in Goniodes (Fig. 8), but subgenital plate present and bifid structure absent in Rhopaloncus (Fig. 36); basal apodeme comprising less than half the genital length in Goniodes (Figs 6 – 7), but comprising more than three-quarters the length in Rhopaloncus (Figs 34 – 35); mesosome more complicated in structure in Goniodes (Figs 6 – 7) than in Rhopaloncus (Figs 34 – 35); parameres distally divergent and widening in Goniodes (Figs 6 – 7), but parallel or convergent and distal ends narrowly bifid or sinuous in Rhopaloncus (Figs 34 – 35).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753355FF99769D6E91DDE1FE81.taxon	description	Descriptions Both sexes. Large chewing lice of the Goniodes body louse ecomorph. Head broader than long, frons gently rounded to somewhat flattened. Marginal carina uninterrupted, terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni prominent, not bent posteriorly but narrowed to points in females. Antennae sexually dimorphic: in male, scape much swollen and elongated compared to that of female, with two distally squamous processes on posterior margin, the inner process being much longer than the outer process; male pedicel similar to that of female in size, but noticeably curved; male flagellomere I much enlarged compared to that of female and to male flagellomeres II – III, extended distally into curved horn with squamous inner margin. Eyes large. Temples sexually dimorphic, with that of male being largely rounded and that of female being extended laterally into distally flattened point. Occipital process absent in male, present as flattened bulge in female. Head chaetotaxy: dsms, ads, pns, pts, avs 2 all mesosetae or longer; mts 1, mts 3 macrosetae; mts 5 situated on occipital process in female; postantennal head with few (less than 10) central sensilla on each side; s 1 – 2, s 5 – 7 present. Rhombic sclerite small to moderate, clearly separated from pronotum. Pronotum flaring posteriorly, with ppss on postero-lateral corner. Pteronotum roughly pentagonal, fused submedianly to tergopleurite II in male. Postero-lateral corners of pteronotum rounded, with only lpts lateral, but ipts close to postero-lateral corners; smns without visible associated sensillum; mpts situated in hyaline tegument anterior to fusion between pteronotum and tergopleurite II in male. Mesosternum present, metasternum present or absent; mets and mss present. Proepimera large, curling around coxae II, but not fused medianly. Metepisterna not sclerotised medially. Abdomen rounded in male, more elongate in female. Tergopleurites III – VIII medianly separated, with section medial to spiracular openings conspicuously narrowed in more posterior segments in male. Male tergopleurite IX not fused to central plate, but plates overlapping. Female tergopleurites IX – XI fused together, and medianly continuous. Central sternal plates absent, but abdominal segments II – VI in both sexes with lateral accessory sternal plates. Male abdominal chaetotaxy: tergopleurites II – VIII with tergocentral mesosetae, gradually diminishing in number in more posterior segments, dense rows of tps microsetae, the medial of which may be intercalated with mesosetae, and 2 – 3 post-spiracular meso- or macrosetae on each side; ventral side of abdominal segment II – VII with multiple sts mesosetae, on segment VIII one macroseta on each side. Female abdominal chaetotaxy: tergopleurites II – V with dense rows of setae likely representing both tcs and tps, but no tps microsetae (more lateral setae in rows may be shorter than more central rows), tergopleurites VI – VII with dense rows of tcs and at least some tps microsetae on each side, but these not forming distinct rows; tergopleurites III – VII with 1 – 2 psc, tergopleurite VIII with numerous psc, the more medial of which are indistinguishable in length from a row of what is presumably tps; ventral side of abdominal segment II with dense median row of sts, segments III – VI with dense rows of sts and dense row of microsetae anterior to the sts (some intercalated with sts, particularly laterally); segment VII with 0 – 1 central sts. Male subgenital plate large, expanded laterally; female subgenital plate present, medianly continuous, but poorly sclerotised. Male genitalia large, reaching to tergopleurite III when non-everted. Basal apodeme long, comprising more than two-thirds the length of genitalia. Distal end may have bifurcated median structure, but it is unclear whether this structure is actually connected to basal apodeme, or comprises the proximal thickening of the mesosome. Mesosome variable among species, ventrally as rounded or semi-oval plate with rugose nodi at or near postero-lateral corners. Parameres long, not fused to basal apodeme or mesosome, distally either trifid or sinuous. Genital sac present, without conspicuously rugose areas. Female genitalia without bifid structure internally and without triangular processes laterally. Vulval margin concave, extended to very near distal end of abdomen, with medially interrupted row of typically very short setae (vms + vss?) and scattered microsetae (vss + vos?) in subgenital area. Subgenital plate with distal end reticulated. Subvulval plates present, small.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF99769D69A8D812FD47.taxon	biology_ecology	Type host: Lophophorus sclateri orientalis Davidson, 1974 — Sclater’s monal.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF99769D6AA3DD00F96A.taxon	biology_ecology	Type host: Lophophorus impeyanus (Latham, 1790) — Himalayan monal. Host distribution: Species of Lophophorus Temminck, 1813.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF99769D6AA3DD00F96A.taxon	distribution	Geographical range: High elevations in the Himalayas. A third species may be present in Yunnan, China (Gustafsson et al. 2024 a).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF99769D6AA3DD00F96A.taxon	etymology	Etymology: The name Rhopaloncus is constructed from “ rhópalon ” (ῥόπᾰλον), ancient Greek for “ club ”, and “ ónkos ” (ὄγκος), Ancient Greek for “ lump ”. This is in reference to the scape of the male, which has both a club-like and a more lump-like appendage.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF99769D6AA3DD00F96A.taxon	discussion	Remarks: Both Clay (1940) and Lakshminarayana (1982) discussed the species of Goniodes from Lophophorus spp. and noted that whereas two male morphs were present in collections (i. e., Margaritenes Kéler, 1940 and Rhopaloncus), only one female morph had ever been found. Males of Margaritenes eurygaster (Piaget, 1885) have the same head shape as the female of Rhopaloncus illustrated here (Fig. 31), but many post-cephalic characters of female M. eurygaster are the same as those in Rhopaloncus spp. and different from males of Margaritenes (see Gustafsson et al. 2024 a). Clay (1940) speculated that the females of these two species may not differ much morphologically. An alternative scenario is that they do differ, but the female of one species has not been found yet. We here include illustrations and a description of the females collected alongside male Rhopaloncus in the NHML collection, noting that no males of Margaritenes from the same host individuals are present in the NHML collection. However, Clay (1940) remarked that not all lice from these individuals may have been collected. Ultimately, more specimens are needed to establish whether the females of these two genera are morphologically different and, if so, to which genus the female illustrated and described here belongs. For a recent redescription of the type species with illustrations to compare with those below, see Gustafsson et al. (2024 a).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF9A769D6E89DDB2F81F.taxon	description	(Figs 30 – 36)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF9A769D6E89DDB2F81F.taxon	materials_examined	Type host: Lophophorus sclateri orientalis Davidson, 1974 — Sclater’s monal. Type locality: Yunnan Province, China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF9A769D6E89DDB2F81F.taxon	diagnosis	Diagnosis. Rhopaloncus lepidoceros can be separated from Rhopaloncus megaceros by the following characters [see Gustafsson et al. (2024 a) for a recent redescription and illustrations of R. megaceros]: inner squamous process of male scape reaches to site of pos in R. lepidoceros (Fig. 32), but shorter and stouter in R. megaceros; ventral mesosome rounded with deep median indentation of posterior margin in R. megaceros, but semi-oval with distal margin concave in R. lepidoceros (Fig. 35); parameres roughly parallel with broad, sinuous distal ends in R. megaceros, but convergent with distinctly trifid distal ends in R. lepidoceros (Figs 34 – 35). Male tergopleurites with generally fewer tergocentral mesosetae in R. lepidoceros (Fig. 30) than in R. megaceros, but as very few specimens of each species were examined this may be artefactual. More specimens are needed before adequate comparisons of abdominal chaetotaxy can be made.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF9A769D6E89DDB2F81F.taxon	description	Descriptions Both sexes. Head relatively wide, preantennal area rather short (Fig. 32); frons gently rounded. Marginal carina narrow, with sinuous inner margin. Head chaetotaxy as in Figs 30 – 32; os sexually monomorphic; dorsal area of postantennal head with few (less than 10) sensilla on each side, typically clustered centrally. Temples sexually dimorphic. Thoracic and abdominal segments and chaetotaxy as in Figs 30 – 31. Male. Scape swollen and elongated compared to female, with posterior margin bearing two squamous horns, the proximal being more than five times the length of the distal (Fig. 32); marginal seta situated on distal horn thorn-like; pedicel somewhat curved; flagellomere I much extended distally into curved horn. Ocular seta short, similar to pos in length. Temples roughly rounded, but outline irregular; occipital process absent. Abdominal chaetotaxy (Fig. 30): tergopleurites II – V with 3 – 5 tcs on each side; tergopleurites VI – VII with 1 – 2 tcs on each side; tergopleurites II – VIII with dense rows of tps microsetae, the most median of which may be intercalated with tcs; tergopleurites II – VII with three (rarely two, in which case the most median may be separated by a clear gap) psc on each side; tergopleurite VIII with 4 – 5 psc on each side; ventral surface of abdominal segments II – VII with dense central rows of sts; segment VIII with one sts on each side; segments IV – VI with microsetae either anterior to sts or intercalated with these. Basal apodeme long, dorsally apparently extended distally as a poorly sclerotised flap (Fig. 34). Mesosome semi-oval, flaring distally with postero-lateral corners rugose. Parameres slender, distally convergent, with distal ends trifid, with pst 1 – 2 situated subapically. Measurements as in Table 1. Female. Antennae slender, with flagellomere I somewhat bulbous (Fig. 33). Coni short, pointed distally. Ocular seta short. Temples expanded into acute angle; occipital process present as slight bulge of posterior head margin. Abdominal chaetotaxy (Fig. 31): tergopleurites II – V with dense rows of tcs all of roughly the same length; tergopleurites VI – VII with 1 – 2 tcs on each side, flanked laterally by rows of shorter tps, some of which may be microsetae; tergopleurites III – V with one psc on each side; tergopleurites VI – VII with 2 – 4 psc on each side; ventral surface of abdominal segments II – VI with dense rows of sts; segment VII with two sts on each side; segment VIII with one sts on each side, close to those of segment VII; segments III – VI with rows of microsetae, some of which are intercalated with sts. Vulval margin gently bilobed (Fig. 36), with 27 – 30 marginal setae on each side (one female with 23 on one side) of roughly equal length, and 13 – 19 microsetae on each side in subgenital area. Subvulval plates small. Measurements as in Table 1.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF9A769D6E89DDB2F81F.taxon	etymology	Etymology: The species epithet is constructed from “ lepís ” (λεπίς), ancient Greek for “ scale ”, and “ kéras ” (κέρας), ancient Greek for “ horn ”, referring to the intensely scaled horns of the male scape.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF9A769D6E89DDB2F81F.taxon	materials_examined	Material examined Types: Ex Lophophorus sclateri orientalis: Holotype: ♂, Yunnan Province, China, May 1906, coll. R. Meinertzhagen, 3626, NMHUK 010677303 [marked with black dot on slide; upper male near the “ M ” in Meinertzhagen] (NHML). Paratypes: 1 ♂, 3 ♀, same data as holotype (NMHL).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336BFF9A769D6E89DDB2F81F.taxon	discussion	Remarks: One male [NHMUK 010677304] from the Lophophorus sclateri orientalis collected in Tibet is not conspecific with R. lepidoceros, having distal parameres similar to those of R. megaceros, but with shorter and disproportionately broad genitalia. Also, there are differences in head shape and size between this male and all males of R. megaceros and R. lepidoceros. The host subspecies living in the collection locality is L. sclateri sclateri Jerdon, 1929, suggesting that different host subspecies are parasitised by different species of Rhopaloncus. Unfortunately, this male lacks most of its longer setae, the head is detached from the body, one of the parameres is broken, and its genitalia have been dissected. Therefore, the true identity of this male must be confirmed with further collections from L. sclateri sclateri.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336CFF9E769D68C1D860FA9D.taxon	description	Clay (1940) included a great variety of species in her Species Groups I – L, most of which belong to the genera Oulocrepis Kéler, 1940, Gonotyles Kéler, 1940, Solenodes Kéler, 1940 and “ Gonocephalus Kéler, 1940 ” [preoccupied; see below]. These groups share a mosaic distribution of morphological characters (Table 3), few of which are common to all species, but all are different from those of Goniodes sensu stricto and should be recognised as separate genera, as argued by Mey (2009). Based on the species we have examined, we recognise at least five genera in this group, including two new genera, as follows: Oulocrepis, Gonotyles, Solenodes, Complectodes new genus and Majiodes new genus. Apart from the genera listed in Table 3, a few other smaller groups may belong to the Oulocrepis - group, but as we have not examined specimens of those smaller groups, we do not treat them further here. The only named genus not included in Table 3 that definitely belongs to this group is Astrocotes Kéler, 1940. Most species included in the Oulocrepis - group share the following characters (although some characters are absent or different in some genera): male flagellomere I modified in distal end; temples sexually dimorphic; only mts 5 situated on occipital process; meso- and metasterna absent, and no setae on ventral thorax; male tergopleurite IX not fused with central plate (but may overlap); abdominal chaetotaxy generally with few meso- or macrosetae, particularly on ventral side; female abdominal segment IX + X expanded distally to form distinct lobes on which at least some macrosetae are situated; bifid process of female genitalia absent; vulval margin convex in almost all species, with lateral sections on each side often bulging or forming distinct lobes on which all or most of the marginal setae are situated; vulval submarginal setae typically clearly identifiable as either slender or thorn-like setae in submarginal row. All species of the Oulocrepis - group, except those in Gonotyles, have triangular processes lateral to the female vulval margin, suggesting that Gonotyles may be more distantly related. As many of the characters listed above are not present in Goniodes sensu stricto, it is possible that Gonotyles does not belong to the Oulocrepis - group, but it is retained in here with respect to the placement of species in Group J by Clay (1940), implying that she believed these were closely related. If Gonotyles does belong in the Oulocrepis - group, it seems plausible that Astrodes Kéler, 1940, Kelerigoniodes Conci, 1946 and possibly Stenocrotaphus Kéler, 1940 may also belong in this group, but more detailed examination of specimens is required. We regard Oulocrepis and Gonotyles as polytypic. Furthermore, considering the morphological differences between two groups within Gonotyles, we recognise them as subgenera, including the new subgenus Gonotyles (Syrmatichares) new subgenus. In addition, several species are placed as incerta sedis within Gonotyles because they need redescription and may represent a separate subgenus or genus. The differences among the male genitalia of all species of Oulocrepis also suggest that several subgenera are involved, but a more comprehensive revision of the genus is needed to determine generic boundaries. All other genera in this group are smaller and morphologically homogeneous. In Table 3, we have summarised the most important and most easily distinguishable features among these genera, but more differences are detailed under the diagnosis of each genus, below.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336CFF91769D6D9DD965FAE9.taxon	type_taxon	Type species: Goniodes dissimilis Denny, 1842: 57. By original designation.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336CFF91769D6D9DD965FAE9.taxon	diagnosis	Diagnosis. Oulocrepis is morphologically variable, particularly with regards to the male genitalia and the male abdominal chaetotaxy, but can be separated from the other genera in the Oulocrepis - group by the following combination of characters (see Table 2): male scape without process or with only slight bulge of posterior margin of scape; distal process of male flagellomere I broadly rounded distally; male genitalia not solenoid; female genitalia with triangular process laterally. Oulocrepis can be separated from Goniodes sensu stricto by the following characters: mesosternum and sternal thoracic setae (mss and mets) present in Goniodes (Figs 1, 3), but absent in Oulocrepis (Figs 45 – 46); male scape with distally squamous process on posterior margin in Goniodes (Fig. 4), but without posterior process or at most with slight bulge on posterior margin of scape (in which case one seta is positioned apically on bulge) in Oulocrepis (Fig. 47); mts 4 – 5 both situated on occipital process in Goniodes (Fig. 4), but only mts 5 on occipital process in Oulocrepis (Fig. 47); male and female coni may be bent posteriorly in Oulocrepis (Fig. 48), in which case they are always short and broad, never slender and extended across much of scape as in Goniodes (Fig. 5); male tergopleurite IX fused with central sclerite in Goniodes (Fig. 1), but separate in Oulocrepis (Fig. 45); both sexes with multiple sts on each side on abdominal segments II – VI in Goniodes (Figs 1, 3), but with only one sts on each side in Oulocrepis (Figs 45 – 46); female abdominal segment IX + X expanded distally to form setose lobes on each side in Oulocrepis (Fig. 46), but not expanded in Goniodes (Fig. 3); male genitalia variable within Oulocrepis, with Figs 49 – 50 showing one rather common type, but no genital type known within Oulocrepis is similar to those of Goniodes (Figs 6 – 7); bifid inner structure of female genitalia present, but triangular lateral processes absent in Goniodes (Fig. 8), whereas in Oulocrepis, the bifid structure is absent and the triangular processes are present (Fig. 51); vulval margin more or less straight in Goniodes (Fig. 8), being at most slightly concave or slightly convex, but strongly convex in Oulocrepis (Fig. 51), with lateral ends forming at least prominent bulges, but often with distinct, densely setose, lobes; vulval submarginal setae not distinguishable from vulval marginal setae in Goniodes (Fig. 8), but forming distinct rows of thorn-like setae in Oulocrepis (Fig. 51).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75336CFF91769D6D9DD965FAE9.taxon	description	Descriptions Both sexes. Large chewing lice of the Goniodes body louse ecomorph. Head broader than long, frons gently rounded to somewhat flattened. Marginal carina uninterrupted, broad with deep attendant canals of setal apertures, terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni prominent, in males generally protruding straight laterally, in females more variable among species and may be bent posteriorly to overlap with scape. Antennae sexually dimorphic: in male, scape much swollen and elongated compared to that of female, with no posterior process or at most with posterior margin bulging slightly, in which case one seta may be situated on bulge; male pedicel similar to that of female in size, but in many species noticeably curved; male flagellomere I much enlarged compared to that of female and to male flagellomeres II – III, extended distally into typically broad, blunt horn. Eyes large. Temples sexually dimorphic: in males variable among species, from gently rounded to protruding laterally; in females typically protruding laterally into acute angle, with ventral tubercle bearing the pos; occipital process present, but variable in extent among species. Head chaetotaxy: varying slightly among species, but dsms, ads, pas, os, pns, avs 1 – 2 all typically meso- or macrosetae; mts 1, mts 3 only temporal macrosetae; mts 5 situated on occipital process; head sensilla s 1 – 6 typically all visible, but variable among species. Rhombic sclerite generally small, clearly separated from pronotum in most species, but may be fused, especially in females of some species. Pronotum rounded distally, with ppss on postero-lateral corner. Pteronotum roughly pentagonal, but with posterior margin variable among species, in some species with postero-median point more extended. Postero-lateral corners of pteronotum not modified, only lpts lateral and ipts more median; smns not associated with sensillum; smns and mpts may be on pteronotum on in hyaline tegument posterior to pteronotum. Meso- and metasterna and associated setae absent. Proepimera large, curling around coxae II, but not fused medianly. Metepisterna not sclerotised medially. Abdomen rounded in male, more elongate in female; tergopleurite II not fused to pteronotum in either sex, and tergopleurites II – VIII medianly separated, with section median to spiracular openings conspicuously narrowed in more posterior segments in many species. Male tergopleurite IX not fused to central plate, but may overlap slightly. Female tergopleurites IX – XI fused, medianly continuous, in many species extended latero-distally to form distinct lobes on each side, which may be densely setose ventrally. Central sternal plates absent, but abdominal segments II – VI in both sexes may have lateral accessory sternal plates, though these are also absent in some species, particularly in males. Abdominal chaetotaxy of both sexes variable among species. Male subgenital plate variable among species, typically not extensive; female subgenital plate generally absent, but small plates may be present near vulval margin. Male genitalia variable among species, and likely form good basis for species groups. Basal apodeme typically fused at least to parameres in most species, but may be fused to mesosome as well; in some species parameres are fused to mesosome, which in turn is fused to basal apodeme. Parameres often very small, and pst 1 – 2 may be difficult to identify. Genital sac present in many species, and if present then typically with minute spicules. Female genitalia without bifid structure internally, but with triangular processes on each side of vulval margin. Structure of vulval margin variable among species, and both type species and Oulocrepis stylophorus sp. nov. are atypical. Most species have medianly bulging or slightly pointed vulval margin, with lateral ends forming smaller bulges or, in some species, clear lobes on which all or most marginal mesosetae (vms) are situated. In O. dissimilis and O. stylophorus, lateral lobes and more pronounced, and other parts of vulval margin strongly modified. In all examined species, vss clearly identifiable as thorn-like submarginal setae, which in some cases are dagger-like; vos scattered in subgenital area in most species, but present only as sensilla in type species. Subvulval plates absent.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753363FF91769D6DF0DFC8F9AF.taxon	biology_ecology	Type host: Ammoperdix griseogularis (Brandt, 1843) — see-see partridge.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753363FF91769D6ECBDF91F885.taxon	biology_ecology	Type host: Tetrao urogallus Linnaeus, 1758 — western capercaillie.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753366FF94769D68C1D802FED9.taxon	materials_examined	Type host: Bonasa umbellus umbelloides (Douglas, 1829) — gray ruffed grouse.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753366FF94769D69D8D805FDD7.taxon	materials_examined	Type host: Centrocercus urophasianus (Bonaparte, 1827) — greater sage-grouse.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753366FF94769D6AD2D84EFCCE.taxon	materials_examined	Type host: Chrysolophus amherstiae (Leadbeater, 1829) — Lady Amherst’s pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753366FF94769D6B2DDFA7FB98.taxon	materials_examined	Type host: Phasianus colchicus Linnaeus, 1758 — red-necked pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753366FF94769D6C98DFBBFA97.taxon	materials_examined	Type host: Canachites canadensis (Linnaeus, 1758) — spruce grouse.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753366FF94769D6D92DF9FF96C.taxon	materials_examined	Type host: Tetraogallus altaicus (Gebler, 1836) — Altai snowcock.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753366FF94769D6E8CDFEAF89B.taxon	materials_examined	Type host: Tympanuchus cupido (Linnaeus, 1758) — greater prairie-chicken.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753365FF97769D68C1DF25FE7C.taxon	materials_examined	Type host: Gallus gallus (Linnaeus, 1758) — red junglefowl.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753365FF97769D69BCDFEEFD6B.taxon	materials_examined	Type host: Ithaginis cruentus cruentus (Hardwicke, 1821) — blood pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753365FF97769D6AB6DF91FBFB.taxon	materials_examined	Type host: Lagopus lagopus (Linnaeus, 1758) — willow ptarmigan.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753365FF97769D6C26D85EFAF2.taxon	materials_examined	Type host: Lagopus leucura peninsularis Chapman, 1902 — white-tailed ptarmigan.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753365FF97769D6D31D8A4F9CF.taxon	materials_examined	Type host: Dendragapus obscurus richardsonii (Douglas, 1829) — Richardson’s dusky grouse.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753365FF97769D6E2AD81EF8C6.taxon	materials_examined	Type host: Tetrao urogalloides Middendorff, 1853 — black-billed capercaillie.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753365FF88769D6F25D8CCFF55.taxon	materials_examined	Type host: Tympanuchus phasianellus campestris (Ridgway, 1884) — prairie sharp-tailed grouse.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337AFF88769D6954DFFEFE43.taxon	materials_examined	Type host: Scleroptila shelleyi (Ogilvie-Grant, 1890) — Shelley’s francolin.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337AFF88769D69AFDF65FD7A.taxon	materials_examined	Type host: Catreus wallichii (Hardwicke, 1827) — cheer pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337AFF88769D6AB9DF99FC57.taxon	materials_examined	Type host: Callipepla californica (Shaw, 1798) — California quail.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337AFF88769D6C53D857FB4E.taxon	materials_examined	Type host: Bambusicola thoracicus (Temminck, 1815) — Chinese bamboo partridge.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337AFF88769D6CADDF37FA19.taxon	materials_examined	Type host: Lyrurus tetrix (Linnaeus, 1758) — black grouse.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337AFF89769D6E18DFB7FF1D.taxon	materials_examined	Type host: Tetraogallus tibetanus tibetanus Gould, 1854 — Tibetan snowcock.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337AFF89769D6E18DFB7FF1D.taxon	biology_ecology	Host distribution: Species of Oulocrepis parasitise a variety of host genera in the Phasianidae.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337AFF89769D6E18DFB7FF1D.taxon	distribution	Geographical range: Northern Hemisphere, but the presence of O. oreophilus on African francolins indicates that the geographical range of Oulocrepis may be wider than currently known. Some hosts of species of Oulocrepis have been introduced by humans into other parts of the world, expanding the geographical distribution of these lice (e. g., Palma 2017).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337AFF89769D6E18DFB7FF1D.taxon	discussion	Remarks: The species placed in Oulocrepis can be divided into a number of species groups, based on e. g., the structure of the male genitalia, the shape of the male temples, the abdominal chaetotaxy of primarily the male, and the structure of the female genitalia (Gustafsson & Johnson, in prep.). Gustafsson et al. (2024 a) redescribed and illustrated numerous species of Oulocrepis, including the type species.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337BFF8E769D691CDFC8FEF9.taxon	description	(Figures 45 – 51)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337BFF8E769D691CDFC8FEF9.taxon	materials_examined	Type host: “ Pyrrhocorax pyrrhocorax Tunstall, 1771, (Passeriformes: Corvidae) ” (error for Bambusicola thoracicus (Temminck, 1815) — Chinese bamboo partridge. See Remarks below. Type locality: Jianou, Fujian Province, China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337BFF8E769D691CDFC8FEF9.taxon	diagnosis	Diagnosis. Within Oulocrepis, O. stylophorus is morphologically unique, and not particularly close to any known species. Based on the presence of distinct lateral lobes of the vulval margin, dagger-like vss, and more prominent and ventrally more setose postero-lateral extensions of female abdominal segment IX + X (Figs 46, 51), O. stylophorus is likely most closely related to O. dissimilis (Denny, 1842). Males of these two species are not particularly similar, but the mesosome has a recurved dorsal thickening in both species (Fig. 49) and the parameres appear not to be fused to either the mesosome of the basal apodeme (Figs 49 – 50). Oulocrepis stylophorus can be separated from O. dissimilis by the following characters [see Gustafsson et al. (2024 a: 475 – 476, figs. 590 – 596) for a recent redescription and illustrations of this species]: preantennal area of female head broadly rounded to somewhat flattened in O. dissimilis, but narrowed distally in O. stylophorus (Fig. 46); female with large, slightly less sclerotised area on preantennal head in O. stylophorus (Fig. 46), but no such area in O. dissimilis; male tergopleurites II – VII with one tcs on each side in O. stylophorus (Fig. 45), but with 2 – 4 tcs on each side in O. dissimilis; male pteronotum with 3 – 4 ipts on each side in O. stylophorus (Fig. 45), but with two ipts on each side in O. dissimilis; male tergopleurite IX elongated, much wider than long in O. stylophorus (Fig. 45), but roughly quadratic in O. dissimilis; male without lateral accessory sternal plates in O. dissimilis, but with such plates on segments II – VIII in O. stylophorus (Fig. 45); ventral surface of female abdominal segments II – VI with dense rows of microsetae in addition to sts mesosetae in O. dissimilis, but without microsetae in O. stylophorus (Fig. 46); male genitalia with triangular, distally tapering parameres in O. stylophorus (Figs 49 – 50), but with slender parameres in O. dissmilis; mesosome with roughly crescent-shaped anterior plate ventrally in O. stylophorus (Fig. 50), but without such plate in O. dissimilis; median section of vulval margin narrowly rounded in O. dissimilis, but forming unique trapezoidal “ stylus ” in O. stylophorus (Fig. 51); area lateral to vulval margin in O. dissimilis with multiple small setae-bearing tubercles in O. dissimilis, but without such tubercles and no dense patches of setae in area in O. stylophorus (Fig. 51).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337BFF8E769D691CDFC8FEF9.taxon	description	Descriptions Both sexes. Head broad and long, shape sexually dimorphic (Figs 45 – 47). Marginal carina broad, with deep attendant canals of preantennal setae. Head chaetotaxy as in Figs 46 – 47; dorsal head without dense rows of sensilla, only s 1 – 4, s 6 – 8 present (not all visible in examined males). Antennae sexually dimorphic. Thoracic and abdominal segments as in Figs 45 – 46. Male. Preantennal head gently rounded, coni large and prominent, extending laterally, temples gently rounded, with pos on lateral head margin. Dorsal head surface without less sclerotised preantennal area. Scape swollen and elongated compared to female, with distal margin bulging slightly in distal half; pedicel swollen and elongated compared to female; flagellomere I extended distally into broad, blunt horn. Pterothorax with 3 – 4 ipts on each side (Fig. 45); smns short. Abdominal chaetotaxy (Fig. 45): tergopleurites II – VII with one tcs on each side; tergopleurites II – VI with rows of tps microsetae, more numerous in more anterior segments; tergopleurites II and VIII with 4 – 5 psc on each side, tergopleurites III – VI with three psc on each side, tergopleurite VII with four psc on each side; ventral side of abdominal segments II – VIII with one sts on each side, on some segments with one microsetae on each side flanking sts, but microsetae are irregular; lateral sternal seta of segment VI not present in examined males. Basal apodeme long and slender, seemingly fused with mesosome dorsally (Fig. 49). Mesosome dorsally with slender, distally curved, thickenings. Mesosome ventrally elongated, narrowing slightly distally, with latero-distal corners with recurved hooks (Fig. 50); separate set of slender, distally curved thickenings present on ventral side. Roughly crescent-shaped structure present proximal to mesosome. Parameres short, tapering gradually distally, somewhat convergent distally, with only pst 1 sensillum visible near distal ends. Genital sac present, small, weakly spiculate. Measurements as in Table 1. Female. Frons more arched than in male (Fig. 46), coni small, strongly recurved posteriorly to overlap slightly with scape (Fig. 48), temples flaring into acute angles, with pos positioned on ventral tubercle. Preantennal area with large, less sclerotised area. Antennae as in Fig. 48. Pterothorax with two ipts on each side, and smns longer. Tergopleurites IX + X extended distally into hooked lobe. Abdominal chaetotaxy (Fig. 46): tergopleurites II, VI – VII with two tcs on each side, tergopleurites III – V with three tcs on each side, tergopleurite VIII with one tcs on each side; tps microsetae absent; tergopleurite II without psc, tergopleurite III with two psc on each side, tergopleurites IV – VII with three psc on each side, tergopleurite VIII with two psc on each side, flanked medianly by row of short setae (tps); ventral surface of abdominal segments II – VIII with one sts on each side. Genital area unique within genus (Fig. 51), with vulval margin swollen into distinct lateral lobes and medianly modified to roughly trapezoidal stylus, the lateral corners of which are slightly fringed. Vulval chaetotaxy: 20 – 24 long, stout setae (vms?) on each side on lateral lobes of vulval margin, with 30 – 36 shorter, more slender setae (vss?) on each side at base of lateral lobes; 5 – 6 large, recurved setae (vss?) on each side along lateral margins of stylus, and 11 – 14 microsetae (vos) on each side on stylus and in subgenital area. Triangular processes slender, slightly recurved. Ventral side of lobes formed by abdominal segments IX + X with dense patches of mesosetae. Measurements as in Table 1.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337BFF8E769D691CDFC8FEF9.taxon	etymology	Etymology: The species epithet is formed from “ stylus ”, Latin for writing implements — referring to the hardened distal appendage of the abdomen or subgenital plate in various Phthiraptera — and “ - phóros ” (- φόρος), ancient Greek for “ bearing ”.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337BFF8E769D691CDFC8FEF9.taxon	materials_examined	Material examined Types: Ex “ Pyrrhocorax pyrrhocorax ” (error for Bambusicola thoracicus): Holotype ♂, Jianou, Fujian Province, China, 7 Jan. 1997, collector unknown, box E 0026139, slide 8 (NHMC). Ex “ Psilopogon virens ” (error for Bambusicola thoracicus): Paratype: 1 ♂, same data as holotype, box E 0026195, slide 37 (NMHC). Ex Bambusicola thoracicus: Paratypes: 1 ♀, Guilin, Guangxi Province, China, 29 Jul 1975, collector unknown, box E 0026136, slide 88 (NHMC). 4 ♂, 2 ♀, no collection data, box E 0026195, slide 37 (NMHC). Non-types: Ex Bambusicola thoracicus: One nymph, Guilin, Guangxi Province, China, 29 Jul 1975, collector unknown, box E 0026136, slide 88 (NHMC). One nymph, Jianou, Fujian Province, China, 7 Jan. 1997, collector unknown, box E 0026195, slide 37 (NHMC). Two nymphs, no collection data, box E 0026195, slide 37 (NMHC) [Nymphs identified by host association].	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337BFF8E769D691CDFC8FEF9.taxon	discussion	Remarks: We reluctantly designated as the holotype and one paratype of Oulocrepis stylophorus, two specimens labelled with incorrect, not natural hosts. The reason behind this unusual designation is that the available specimens with host labels written as Bambusicola thoracica held at the NHMC are all in extremely poor condition. Pyrrhocorax pyrrhocorax Tunstall, 1771, (Passeriformes: Corvidae) does not occur at the type locality (Liu & Chen 2024) and there is no goniodid known from corvid hosts. Equally, there is no goniodid known from Psilopogon virens (Boddaert, 1783) (Piciformes: Megalaimidae), the host of the paratype. Notably, the labels of the slides bearing the holotype and the paratype (ex Psilopogon virens) are both marked in pencil with the word “ thoracicus ”, suggesting that whoever made the slides suspected an error, but did not correct it. Six other slides with specimens from Bambusicola fytchii Anderson, 1871 mostly include a species similar to O. stylophorus, further indicating that the natural host of this species is a member of Bambusicola Gould, 1863. However, these specimens are also poorly preserved, with the glass coverslip of one slide broken, making adequate comparisons impossible. Some specimens on these slides are more similar to Gonotyles lophurus, but so poorly preserved that they cannot be identified. The series of slides containing the holotype of O. stylophorus includes specimens of Philopterus Nitzsch, 1818, Menacanthus Neumann, 1912, Myrsidea Waterston, 1915, Sturnidoecus Eichler, 1944, Priceiella Gustafsson & Bush, 2017, Picophilopterus Ansari, 1947, and a poorly preserved unidentified non-parasitic psocopteran. All of these specimens are unidentified to species level, and many are poorly preserved. Another slide from the same series contains a male and a female of Kelerigoniodes processus (Kellogg & Paine, 1914), but on its label two hosts are given: P. pyrrhocorax and Arborophila gingica (Gmelin, 1789), the latter a known host of K. processus (Price et al. 2003). In summary, it seems clear to us that the given hosts on the type slides are the result of a mix-up during the slidemounting process and not the bird species from which they were collected. Considering the available evidence, we conclude that Bambusicola thoracica is the natural, regular host of O. stylophorus. However, this association could only be confirmed when new collections from B. thoracica are examined.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337CFF8F769D69E0DCD3F9B5.taxon	type_taxon	Type species: Goniodes cervinicornis Giebel, 1874: 199, by original designation.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337CFF8F769D69E0DCD3F9B5.taxon	diagnosis	Diagnosis. Gonotyles can be separated from Goniodes sensu stricto by the following characters [see Gustafsson et al. (2024 a) for recent redescriptions and illustrations of G. cervinicornis]: male tergopleurite IX not fused to central plate in Gonotyles (Fig. 37), but fused in Goniodes (Fig. 1); male abdominal segments II – V with only one tergocentral mesoseta on each side in Gonotyles (Fig. 37), but with multiple tergocentral mesosetae in Goniodes (Fig. 1); female temples extended into angles and with pos situated on ventral tubercle in Gonotyles (Fig. 38), but temples rounded and without tubercle in Goniodes (Fig. 3); occipital process with only mts 5 apically in Gonotyles (Figs 37 – 39), but with both mts 4 – 5 subapically in Goniodes (Fig. 4); coni bent posteriorly in female Goniodes (Fig. 5), but not bent in Gonotyles (Fig. 40); postero-lateral corners of pteronotum rounded, with only lpts lateral, in Gonotyles (Figs 37 – 38), but flattened, with both lpts and ipts lateral in Goniodes (Figs 1, 3); mesosternum absent in Gonotyles (Figs 37 – 38), but present in Goniodes (Figs 1, 3); male genitalia clearly divided into basal apodeme, mesosome and paired parameres in Goniodes (Figs 6 – 7), but with all elements partially fused in Gonotyles (Figs 41 – 42); male genital opening may have serrated structures laterally or anteriorly in Gonotyles (Fig. 43; not known for all species), but such structures absent in Goniodes; vulval margin strongly convex with distinct sublateral lobes bearing all or most of the lateral setae in Gonotyles (Fig. 44), but at most slightly convex and without sublateral lobes in Goniodes (Fig. 8); bifid structure of female genitalia absent in Gonotyles (Fig. 44), but present in Goniodes (Fig. 8); female abdominal segment IX + X extended postero-laterally into distinct lobes bearing multiple macrosetae in Gonotyles (Figs 38, 44), but not extended into lobes in Goniodes (Figs 3, 8). We consider Gonotyles to comprise two subgenera: Gonotyles (Gonotyles) Kéler, 1940, and Gonotyles (Syrmatichares) new subgenus. In addition to the characters listed above, some characters that are unique to each subgenus also differ from Goniodes. Both subgenera of Gonotyles can be separated from the other members of the Oulocrepis - group by the following characters: either male scape has a bifid process or the male genital opening has a serrated sclerite (Fig. 43) in Gonotyles (serrated sclerite may be separate from other sclerites), but neither of these characters are found in other Oulocrepis - group genera; female genitalia without triangular process laterally in Gonotyles (Fig. 44), but this is present in other genera of this group. Gonotyles (Gonotyles) can be separated from Goniodes sensu stricto by the following additional characters: male scape with bifid process and two stout mesosetae near posterior margin distal to process in G. (Gonotyles), but with process simple and at most one stout seta distal to process in Goniodes; male abdominal segment VII ventrally with dense, irregular row of macrosetae in G. (Gonotyles), but with one sts macroseta on each side in Goniodes (Fig. 1); parameres slender and distally convergent, with pst 1 – 2 ventral and close to median margin of parameres in G. (Gonotyles), but broad, widening and diverging distally with pst 1 – 2 near lateral margin in Goniodes (Figs 6 – 7); dorsal mesosome long, distally bifurcated and with anterior, apically rugose, lobes in Goniodes (Fig. 6), but roughly Y-shaped, with distal end shallowly concave and proximal lobes not rugose in G. (Gonotyles); ventral genitalia with large papillate areas in anterior half and near distal end in G. (Gonotyles), but without such structures in Goniodes (Fig. 7); female subgenital plates present as slender submarginal sclerites in G. (Gonotyles), but absent in Goniodes (Fig. 8).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337CFF8F769D69E0DCD3F9B5.taxon	description	Descriptions Both sexes. Medium-sized chewing lice of the Goniodes body louse ecomorph. Head shape variable, typically broader than long in females but may be approximately as long as wide in males. Frons gently rounded to somewhat flattened. Marginal carina broad and uninterrupted, but typically with deep attendant canals to setae making the inner margin strongly sinuous; carina terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni prominent, not bent posteriorly in females, variable between subgenera in males. Antennae sexually dimorphic: in male, scape much swollen and elongated compared to that of female, with one process on posterior margin; male pedicel similar to that of female in size, but noticeably curved; male flagellomere I much enlarged compared to that of female and to male flagellomeres II – III, extended distally into curved horn with squamous inner margin. Eyes large. Temples sexually dimorphic, in males slightly rounded to almost flattened, in females extended laterally into angles, bearing a ventral tubercle on which the pos is situated. Occipital process present. Head chaetotaxy: dsms, ads, pns, os, avs 2 all mesosetae or longer; mts 1, mts 3 macrosetae; postantennal head with few (less than 10) central sensilla on each side. Rhombic sclerite small to large, clearly separated from pronotum. Pronotum flaring slightly posteriorly, with ppss on postero-lateral corner. Pteronotum roughly pentagonal, with postero-lateral corners rounded so that only lpts are lateral and ipts are more medial; smns without associated sensillum; mpts in hyaline tegument. Pteronotum not fused to tergopleurite II. Meso- and metasterna, mets and mss absent. Metepisterna not sclerotised. Abdomen rounded in male, more elongate in female; tergopleurite II not fused to pteronotum in either sex, and tergopleurites II – VIII medianly separated. Male tergopleurite IX not fused to central plate. Female tergopleurites IX – XI fused together, and medianly continuous. Central sternal plates absent, but abdominal segments II – VI in at least female with lateral accessory sternal plates. Male abdominal chaetotaxy: tergopleurites II – VII with one tcs on each side; tergopleurites II – VIII with rows (tergopleurite VIII generally with only 2 – 4) tps microsetae, some of which may be medial to tcs; tergopleurites II – VIII with 3 – 5 psc on each side; ventral side of abdominal segments III – VI with one sts on each side. Female abdominal chaetotaxy: tergopleurites II – V or II – VI with 3 – 6 tcs on each side, VI – VII or VII – VII with 1 – 2 tcs on each side; some tergopleurites (typically II – VII; variable among species) with a few (typically 1 – 4 on each side) tps microsetae, which may be intercalated with tcs; ventral side of abdominal segment II without sts (but microsetae may be present; however, not always visible); segments III – VI with one sts on each side; segments II – VI with rows of microsetae (on II often only one microseta on each side), some of which may be median to st; segments VII – VII with one sts on each side. Male subgenital plate variable among subgenera; female subgenital plate present, medianly divided. Subvulval plates present or absent. Female abdominal segment IX + X extended posteriorly to form distinct lateral lobes, typically bearing at least some macrosetae dorsally and ventrally; in some species, ventral surface of these lobes bears a smaller, densely setose lobe, and multiple setae typically present in this area, even if no distinct ventral lobe is discernable. Male genitalia variable between subgenera (see below). In both subgenera, basal apodeme, mesosome and parameres at least partially fused. Ventral side of distal genitalia with densely papillate area, variable in extent between subgenera. Female genitalia without bifid structure internally and without triangular processes laterally. Vulval margins strongly convex, laterally with prominent slender lobes on which all or most marginal mesosetae (vms) are situated; area medial to lobes either without setae or with setae submarginal (vss), in some species with vss clearly thorn-like.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75337DFF8F769D6F65D83AF812.taxon	materials_examined	Type host: Lophura nycthemera nycthemera (Linnaeus, 1758) — silver pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753372FF80769D68C1DF66FED8.taxon	materials_examined	Type host: Lophura diardi (Bonaparte, 1856) — Siamese fireback.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753372FF80769D6980D840FD9F.taxon	materials_examined	Type host: Bambusicola fytchii fytchii Anderson, 1871 — mountain bamboo partridge.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753372FF80769D6A90DF81FC02.taxon	materials_examined	Type host: Lophura ignita (Shaw, 1798) — Bornean crested fireback.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753372FF80769D6C7CD82BFB26.taxon	materials_examined	Type host: Lophura leucomelanos leucomelanos (Latham, 1790) — kalij pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753372FF81769D6D50DF70FC65.taxon	materials_examined	Type host: Lophura leucomelanos lineata (Vigors, 1831) — lineated pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753372FF81769D6D50DF70FC65.taxon	biology_ecology	Host distribution: Most species parasitise hosts in the genera Lophura Fleming, 1822, Pucrasia Gray, 1841 and Syrmaticus Wagler, 1832. These host genera all belong to the “ Phasianidae II ” clade in the phylogeny of Kimball et al. (2021) but are not closely related within that clade. Most of the host species in this clade belonging to other genera are known to be parasitised by goniodid species of other genera. The exception is Gonotyles bambusicolus, known from a member of the clade “ Phasianidae I ” in the phylogeny of Kimball et al. (2021). However, the host association of this species is dubious (see below).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753372FF81769D6D50DF70FC65.taxon	distribution	Geographical range: Central and Southeast Asia.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753372FF81769D6D50DF70FC65.taxon	discussion	Remarks: Most of the species of Gonotyles can be placed in a subgenus, but four species do not fit in either subgenus. Hence, they are listed here as incerta sedis within Gonotyles. We have not seen any specimens of G. longus, but the male genitalia as illustrated by Clay (1940: fig. 48 a) and Kéler (1940: fig. 25) suggest this species may not belong to either of the two subgenera of Gonotyles. Kéler (1940: fig. 24) illustrated the male scape without process, and the vulval margin as being concave rather than convex, further highlighting morphological differences between this species and all other Gonotyles; these characters have not been confirmed by us and require verification. Gonotyles longus requires redescription before it can be adequately placed in a genus. We have examined the alleged type specimens of both Gonotyles lophurus and Gonotyles bambusicolus, but they are all poorly preserved. We have reason to believe that other, now lost, specimens were originally included in the type series, and that non-type specimens may have been marked as types for these species and many other species described by Sikong Liu (Gustafsson et al. 2024 a). The specimens of G. lophurus and G. bambusicolus we have examined are nearly identical, and it seems probable that those of G. bambusicolus represent stragglers from Lophura leucomelanos, but this assumption needs verification. In all males of both species, the male genitalia are distorted, but appear to represent the same general type as in Gonotyles longus. Moreover, no females of G. lophurus and G. bambusicolus could be examined in detail, due to the poor state of the specimens at the NHMC. Fresh, good quality specimens are needed from Lophura leucomelanos and Bambusicola fytchii to redescribe and elucidate the true identity of these species. However, partial illustrations and descriptions were published by Gustafsson et al. (2024 a). We have not seen specimens of Gonotyles ocellatus, but we place it in Gonotyles incerta sedis based on the illustration of the male genitalia by Clay (1940: fig. 36, as Goniodes dentatus), regarded as of the same type as the other three species here considered incerta sedis. Gonotyles ocellatus needs a detailed redescription before it can be reliably placed in any genus. It seems probable that the four species regarded here as incerta sedis are closely related but, as explained above, new collections are needed to prove it or otherwise. Their inclusion in Oulocrepis is equally plausible; however, Kéler’s (1940: figs 23, 25) detailed illustrations of the female of G. longus lack the triangular process of the genitalia found in all species of Oulocrepis. Hence, Kéler (1940) placed Goniodes longus in Gonotyles and we accept that generic association here. Gustafsson et al. (2024 c) listed these four species under Oulocrepis in their table 2, having overlooked the lack of the triangular process in Kéler’s drawing.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753373FF82769D6B84DF01FD15.taxon	type_taxon	Type species: Gonotyles (Syrmatichares) selenautes new species.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753373FF82769D6B84DF01FD15.taxon	diagnosis	Diagnosis. Gonotyles (Syrmatichares) can be separated from Goniodes sensu stricto by the following additional characters: male scape with small, non-squamous process with apical stout seta in G. (Syrmatichares) (Fig. 39), but with process longer, squamous and without apical seta in Goniodes (Fig. 4); male occipital process wide and bulky, giving the postantennal head a squarish look, in G. (Syrmatichares) (Fig. 39), but more slender in Goniodes (Fig. 4); coni not bent posteriorly in either sex in G. (Syrmatichares) (Figs 39 – 40), but bent posteriorly in Goniodes (Figs 4 – 5); male subgenital plate small, roughly T-shaped, not reaching either lateral accessory sternal plates not paratergal plate VIII in G. (Syrmatichares) (Fig. 37), but much expanded, reaching to near lateral accessory sternal plates and paratergal plate VIII in Goniodes (Fig. 1); dorsal mesosome roughly rectangular, with distal half deeply bifurcated in Goniodes (Fig. 6), but small, with species-specific thickenings and lobes in G. (Syrmatichares) (Fig. 41). Further comparisons of male genital elements are difficult, as it is not clear which structures in the genitalia of G. (Syrmatichares) represent the parameres. Most likely, the parameres are the ventral structures that bear two subapical setae, in which case the following characters can separate Goniodes and G. (Syrmatichares): parameres largely fused to mesosome, slender and of irregular shape, with pst 1 – 2 as visible setae subapically in G. (Syrmatichares) (Fig. 42), but large and bulky, widening distally, not fused to mesosome, and with pst 1 – 2 as lateral sensilla (Figs 6 – 7). Gonotyles (Syrmatichares) can be separated from Gonotyles (Gonotyles) by the following characters: male scape with bifid process bearing two stout setae at its base in G. (Gonotyles), but with a monofid process bearing one stout seta apically in G. (Syrmatichares) (Fig. 39); male temples expanded much lateral to eye in G. (Gonotyles), but not or barely expanded lateral to eye in G. (Syrmatichares) (Fig. 39); male abdomen with dense central row of sts on ventral side of segment VII in G. (Gonotyles), but with at most 1 – 2 sts on each side on this segment in G. (Syrmatichares) (Fig. 37); male subgenital plate expanded, with lateral section recurving towards paratergal plate IX + X in G. (Gonotyles), but small, roughly T-shaped in G. (Syrmatichares) (Fig. 37); mesosome dorsally with prominent, roughly Y-shaped plate that overlaps other dorsal mesosomal features, and ventrally with prominent central funnel-shaped structure (gonopore?), but these structures are absent in G. (Syrmatichares) (Figs 41 – 42) and no characters in G. (Syrmatichares) appear to be homologous to them; ventral surface of genitalia with distinct papillate structures in both proximal and distal halves in G. (Gonotyles), but only distally in G. (Syrmatichares) (Fig. 42); female subgenital plate elongated following almost entire vulval margin in G. (Gonotyles), but much reduced in G. (Syrmatichares) (Fig. 44); distal lobes of female abdominal segment IX + X more pronounced in G. (Syrmatichares) (Fig. 44) than in G. (Gonotyles); subvulval plates present in G. (Gonotyles), but absent in G. (Syrmatichares) (Fig. 44). Serrated structure of male genital opening absent in specimens of G. (G.) cervinicornis, but present in all species of G. (Syrmatichares). However, G. (G.) diardi was not examined, and many of the specimens of G. (G.) cervinicornis examined were either dissected or poorly preserved. The absence of this structure in the nominate subgenus needs confirmation. Furthermore, the parameres are long in specie of G. (Gonotyles), clearly visible throughout, with hook-like processes laterally and triangular processes near median edge and pst 1 – 2 as sensilla on ventral surface, but in G. (Syrmatichares) there are three different distal processes on each side of the mesosome, labelled A – C in Figs 41 – 42. The dorsal processes (A in Fig. 41) are likely homologous to a similar structure found in G. (Gonotyles). One ventral process (B in Fig. 42) appears to be completely fused to the mesosome, and may be just a distal elongation of some mesosomal structure. The third process (C in Fig. 42) bears a pair of setae subapically, suggesting that these may be the parameres. If correct, the parameres of G. (Syrmatichares) are irregular in shape, with pst 1 – 2 near apical ends, and smaller in G. (Syrmatichares) than in G. (Gonotyles).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753370FF82769D6CD7DFB7FAC2.taxon	materials_examined	Type host: Syrmaticus humiae (Hume, 1881) — Mrs Hume’s pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753370FF82769D6D21DFE1F9F9.taxon	materials_examined	Type host: Pucrasia macrolopha darwini Swinhoe, 1872 — koklass pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753370FF82769D6BDCDF65FBCB.taxon	materials_examined	Type host: Syrmaticus ellioti (Swinhoe, 1872) — Elliot’s pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753370FF82769D6E38DFAEF80F.taxon	materials_examined	Type host: Syrmaticus mikado (Ogilvie-Grant, 1906) — Mikado pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753370FF82769D6E38DFAEF80F.taxon	biology_ecology	Host distribution: Species of Syrmaticus and Pucrasia.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753370FF82769D6E38DFAEF80F.taxon	distribution	Geographical range: Central and Southeast Asia, Taiwan.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753370FF82769D6E38DFAEF80F.taxon	etymology	Etymology: Syrmatichares is formed by the host genus of most known species included in the subgenus, i. e., Syrmaticus, itself derived from “ syrma ” (σύρμα), ancient Greek for a robe with a long train, referring to the long tails of these birds, and “ khaírō ” (χαίρω), ancient Greek for “ to enjoy ”.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753371FF84769D68C1DE5EFE35.taxon	description	(Figs 37 – 44)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753371FF84769D68C1DE5EFE35.taxon	materials_examined	Type host: Syrmaticus ellioti (Swinhoe, 1872) — Elliot’s pheasant. Type locality: Jianou, Fujian Province, China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753371FF84769D68C1DE5EFE35.taxon	diagnosis	Diagnosis. Gonotyles (Syrmatichares) selenautes lacks the arched transverse structure of the distal basal apodeme found in G. (S.) sinensis [see Emerson & Stojanovich (1964: fig. 3)]. Based on the illustrations of the male genitalia by Clay (1940: fig. 45) and Gustafsson et al. (2024 a: figs 631 – 632), G. (S.) selenautes is likely closest to G. (S.) humiae. It should be noted that Clay (1940) and Emerson & Stojanovich (1964) did not illustrate the distal end of the genitalia, including the papillate distal structure. Gonotyles (S.) selenautes can be separated from G. (S.) humiae by the following characters [see Gustafsson et al. (2024 a) for a recent redescription and illustrations of G. (S.) humiae]: seta-bearing process of male scape more prominent in G. (S.) selenautes (Fig. 39) than in G. (S.) humiae; dorsal mesosome shorter with more prominent antero-lateral processes and stouter distal processes in G. (S.) selenautes (Fig. 41) than in G. (S.) humiae; presumed parameres largely overlapping with ventral mesosomal appendage B in G. (S.) selenautes (Fig. 42), but widely separated in G. (S.) humiae; dorsal mesosomal appendage A and ventral mesosomal appendage B in G. (S.) selenautes (Figs 41 – 42) of different shape than in G. (S.) humiae; distal extensions of female abdominal segment IX + X longer and more prominent in G. (S.) selenautes (Fig. 44) than in G. (S.) humiae; vulval submarginal setae clearly thorn-like in G. (S.) selenautes (Fig. 44), but slender in G. (S.) humiae.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753371FF84769D68C1DE5EFE35.taxon	description	Descriptions Both sexes. Head relatively wide, preantennal area rather short (Fig. 39); frons gently rounded. Marginal carina broad, with deep attendant canals of setae and sinuous inner margin. Head chaetotaxy as in Figs 38 – 39: os sexually monomorphic; head sensilla sexually dimorphic. Temples sexually dimorphic. Thoracic and abdominal segments and chaetotaxy as in Figs 37 – 38. Male. Scape swollen and elongated compared to female, with posterior margin with short process bearing spike-like seta apically; pedicel narrowing slightly distally, somewhat curved; flagellomere I much extended distally into curved horn. Coni large, rounded triangular, not curved. Temples slightly rounded, with section between pos and mts 5 almost straight line; occipital process prominent. Abdominal chaetotaxy: as in genus description. Genital opening with serrated structure laterally (Fig. 43), seemingly continuous with dorsal section of subgenital plate, and additional anterior sclerites without serration. Genitalia overall broad (Figs 41 – 42). Dorsal mesosome short, with near-circular anterior processes and long, distally somewhat recurved, distal appendages (A in Fig. 41); lateral margins of mesosome with two sensilla on each side. Ventral mesosome with appendages B straight to slightly curved, with rugose postero-lateral edges, and slender distal tips. Presumed parameres (C in Fig. 42) hidden below appendages B for most of length, with presumed pst 1 – 2 clearly visible as setae near apical ends. Papillate area of distal genitalia small. Measurements as in Table 1. Female. Antennae slender, coni short and rounded, slightly curved (Fig. 40). Temples expanded, almost into right angles, with pos situated apically on ventral tubercle (Fig. 38). Abdominal chaetotaxy (Fig. 38): tergopleurites II – VII with 3 – 4 tcs on each side, tergopleurites VIII with one tcs on each side; tergopleurites III – VIII with few (2 – 4) tps microsetae on each side; tergopleurites II – VII with 2 – 3 psc on each side; ventrally as in genus description. Subgenital plates small, bilobed, widely separated medianly (Fig. 44). Vulval margin strongly convex, with lateral lobes slender. Vulval chaetotaxy: lateral lobes with 24 – 31 long, slender vms on each side; 5 – 9 short, slender setae (vms?) situated anterior to lobes on each side; 4 – 5 thorn-like vss medial to lobes on each side; subgenital area with 17 – 22 short, slender vos on each side. Post-vulval chaetotaxy as in Fig. 44.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753371FF84769D68C1DE5EFE35.taxon	etymology	Etymology: The species epithet is formed by “ Sel ḗnē ” (Σελήνη), the goddess of the Moon in ancient Greek mythology, and “ naútēs ” (ναύτης), ancient Greek for “ sailor ”. This is in honour of Tsukino Usagi, a manga and anime character created in the 1990 s by Takeuchi Naoko (b. 1967), in recognition of the startling similarities between the dorsal thickenings of the male genitalia of this louse species, and the headdress and hair style of Usagi in her transformed superhero persona, Sailor Moon. The Sailor Moon series was very close to the first author’s heart during his formative years and remains an inspiration to the first author to this day. It is our great pleasure to be able to dedicate this species “ in the name of the Moon ”.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753371FF84769D68C1DE5EFE35.taxon	materials_examined	Material examined Types: Ex Syrmaticus ellioti: Holotype ♂, Jianou, Fujian Province, China, 7 Jan. 1997, collector unknown, box E 0026195, slide 13 (NHMC) [right-most male in row with only two males, marked with black dot on slide]. Paratypes: 7 ♂, 11 ♀, same data as holotype, box E 0026195, slides 10 – 13, 20 (NHMC). 4 ♂, 5 ♀, Wuyi Mountain, Fujian Province, China, 20 Feb. 199, collector unknown, box E 0026195, slides 15 – 17 (NHMC). Non-types: 8 nymphs, Jianou, Fujian Province, China, 7 Jan. 1997, collector unknown, box E 0026195, slides 10, 11, 14 (NHMC). 15 nymphs, Wuyi Mountain, Fujian Province, China, 20 Feb. 199, collector unknown, box E 0026195, slides 15, 17 – 18 (NHMC).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753376FF85769D6A75DD42FB21.taxon	type_taxon	Type species: Goniodes dispar Burmeister, 1838: 432. By original designation.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753376FF85769D6A75DD42FB21.taxon	diagnosis	Diagnosis. Solenodes can be separated from members of the Oulocrepis - group by the following combination of characters: male genitalia solenoid or pseudo-solenoid; male scape and pedicel not or only little modified compared to female; male flagellomere I with small distal extension that either wraps around base of flagellomere II, or is at least not curved towards head; triangular process present lateral to vulval margin; vulval margin with slight bulges laterally. Solenodes can be separated from Goniodes sensu stricto by the following characters: postero-lateral corners of pterothorax flattened and with both lpts and ipts on lateral margins in both sexes of Goniodes (Figs 1, 3), but corners rounded with only lpts on lateral margin in Solenodes (Figs 52 – 53); male tergopleurite IX fused to central plate in Goniodes (Fig. 1), but separate (but may overlap) in Solenodes (Fig. 52); male subgenital plate much smaller and of different structure in Solenodes (Fig. 52) than in Goniodes (Fig. 1); male tergopleurites with rows of tps microsetae in Goniodes (Fig. 1), but with rows of tps mesosetae indistinguishable from tcs in Solenodes (Fig. 52); mesosternal plate and both mss and mets present in Goniodes (Figs 1, 3), but all absent in Solenodes (Figs 52 – 53); ventral sides of abdominal segments II – VI with multiple sts on each side in Goniodes (Figs 1, 3), but with only one sts on each side in Solenodes (Figs 52 – 53); scape and pedicel strongly sexually dimorphic in Goniodes (Figs 4 – 5), but more or less sexually monomorphic, or at least male scape is not elongated and swollen and has no process on posterior margin in Solenodes (Figs 54 – 55); male flagellomere I extended into long distal horn in Goniodes (Fig. 4), but smaller and wrapped around the base of flagellomere II or at least not recurved in Solenodes (Fig. 54); male temples rounded, pos situated on head margin, and both mts 4 – 5 situated on occipital process in Goniodes (Fig. 4), but male temples extended laterally into angular process, pos on ventral tubercle, and only mts 5 on occipital process in Solenodes (Fig. 54); male pas, os and pns all mesosetae in Goniodes (Fig. 4, macrosetae in Solenodes (Fig. 54); male genitalia solenoid or pseudo-solenoid in Solenodes (Fig. 56), but more complicated, with distinct mesosome and parameres in Goniodes (Figs 6 – 7); bifid structure of female genitalia present but triangular processes lateral to vulval margin absent in Goniodes (Fig. 8), but bifid structure absent and triangular processes present in Solenodes (Fig. 57); vulval submarginal setae not identifiable in Goniodes (Fig. 8), but present as median rows of thorn-like setae in Solenodes (Fig. 57); subvulval and postvulval plates present in Goniodes (Fig. 8), but absent in Solenodes (Fig. 57).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753376FF85769D6A75DD42FB21.taxon	description	Description Both sexes. Large chewing lice of the Goniodes body louse ecomorph. Head broader than long, frons gently rounded. Marginal carina uninterrupted, terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni generally small, rounded, not bent posteriorly. Antennae sexually dimorphic: male flagellomere I in all except one species with distal expansion that may wrap partially around flagellomere II, or is at least not recurving; other antennal segments generally similar between sexes, but scape may be slightly swollen in male compared to female. Eyes large. Temples protruding laterally in both sexes, with pos generally on ventral tubercle at apical end of extension (not clearly illustrated for all species); occipital process present. Head chaetotaxy: pas, pns, os, mts 1, mts 3 all macrosetae; ads may also be at least mesoseta; mts 5 only seta situated on occipital process; postantennal head generally with only s 1 – 3 present, but these are not always clearly visible. Rhombic sclerite small, not fused to pronotum. Pronotum rounded posteriorly, with ppss on postero-lateral corner. Pteronotum roughly pentagonal, with postero-lateral corners rounded; only lpts situated laterally; ipts sublateral; smns not associated with sensillum; mpts on or just posterior to pteronotum. Meso- and metasterna and associated setae absent. Proepimera large, curling around coxae II, but not fused medianly. Metepisterna not sclerotised medially. Abdomen typically rounded but abruptly truncate posteriorly in males of most species, such that the posterior margin of abdominal segments VIII, IX + X and IX may be aligned, or VIII, IX + X, or both may even extend more posterior than XI; in female more elongate and not truncate posteriorly. Tergopleurites II not fused to pteronotum; tergopleurites II – VII medianly separate, in males often narrowing conspicuously medial to spiracular openings. Male tergopleurite IX separate from central plate, but may overlap. Female tergopleurites IX – XI fused, with lateral section extended distally. Central sternal plates absent, but abdominal segments II – VI in both sexes may have lateral accessory sternal plates (not clearly visible in all species). Male abdominal chaetotaxy: tergopleurites II – VII with rows of tcs + tps on each side, often with no clear gap between psc and tcs + tps, but with 2 – 3 lateralmost setae (psc) clearly longer than more central setae; ventral side of abdominal segments II – VIII with one sts on each side. Female abdominal chaetotaxy: tergopleurites II – VIII with 1 – 5 tcs on each side, often more numerous in more anterior segments, and 1 – 3 psc on each side; tps microsetae or short mesosetae may be present between these setal sets, but often not illustrated and their distribution poorly known; ventral side of abdominal segments II – VIII with one sts on each side. Male subgenital plate small, deeply concave anteriorly, not expanded laterally; female subgenital, subvulval and postvulval plates absent. Male genitalia either solenoid (with parameres fused to basal apodeme and mesosome absent) or pseudo-solenoid (with parameres fused to basal apodeme, and mesosome reduced to small, often poorly characterised, central structure completely fused to basal apodeme. Parameres generally asymmetrical or flexible, with pst 1 prominent near distal tip. In species with pseudo-solenoid genitalia, remnant of mesosome limited to central, poorly sclerotised rod-like structure surrounded by hyaline bulging structures, but variable among species. Female genitalia without bifid structure, but with triangular processes on each side of vulval margin. Vulval margin convex centrally, and typically with slight lateral setae-bearing bulges (but these are not seen in S. ortygis; see Clay 1940: fig. 62). Vulval chaetotaxy: slender mesosetae (vms?) on lateral bulges, with a few shorter setae (vss?) immediately proximal to these; centrally with rows of thorn-like vss and vos microsetae in scattered, distally convergent rows.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753377FF85769D6D1CDFC7FA06.taxon	materials_examined	Type host: Pternistis leucoscepus (Grey, 1867) — yellow-necked spurfowl.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753377FF85769D6E70DF73F907.taxon	materials_examined	Type host: Pternistis capensis (Gmelin, 1789) — Cape spurfowl.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753377FF85769D6F63DFDCF83B.taxon	materials_examined	Type host: Phasianus colchicus Linnaeus, 1758 — ring-necked pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753374FF86769D68C1DF20FE10.taxon	materials_examined	Type host: Perdix perdix (Linnaeus, 1758) — grey partridge.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753374FF86769D6A10DFAFFD0F.taxon	materials_examined	Type host: Pternistis nobilis (Reichenow, 1908) — handsome spurfowl.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753374FF86769D6B6ADFA5FC06.taxon	materials_examined	Type host: Alectoris magna (Przevalski, 1876) — Przevalski’s partridge.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753374FF86769D6C65DFAAFAF2.taxon	materials_examined	Type host: Colinus virginianus (Linnaeus, 1758) — northern bobwhite.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753374FF86769D6D31D865F9E8.taxon	materials_examined	Type host: Scleroptila gutturalis levalliantoides (Smith, 1836) — Orange River francolin.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753374FF87769D6E08D903FEF9.taxon	materials_examined	Type host: Alectoris barbara (Bonnaterre, 1790) — Barbary partridge.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753374FF87769D6E08D903FEF9.taxon	biology_ecology	Host distribution: Most species of Solenodes occur on either francolins or partridges. The presence of this otherwise Afro-Eurasian genus on C. virginianus requires further study, but Clay (1940) recorded S. ortygis from multiple subspecies of that host from across North America, indicating that this host association is genuine.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753374FF87769D6E08D903FEF9.taxon	distribution	Geographical range: Most species are known from Africa and Eurasia, except for S. ortygis which is from North America. Some hosts of species of Solenodes have been introduced by humans into other parts of the world, expanding the geographical distribution of these lice (e. g., Palma 2017).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753374FF87769D6E08D903FEF9.taxon	discussion	Remarks: Given the differences in abdominal chaetotaxy between species from francolins and species from other hosts, it is possible that this genus may represent two separate groups, united by the reduction of the male genitalia and similarities in the male antennae. Most species placed in Solenodes have not been properly described or illustrated. Hence, redescriptions of almost all species are needed to evaluate the limits of the genus.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753375FFFB769D6938D810FCB9.taxon	description	(Figures 52 – 57)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753375FFFB769D6938D810FCB9.taxon	materials_examined	Type host: Alectoris magna (Przevalski, 1876) — Przevalski’s partridge. Type locality: Haiyuan, Ningxia Province, China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753375FFFB769D6938D810FCB9.taxon	diagnosis	Diagnosis. Solenodes ningxiaensis is most similar to S. capitatus, S. dispar, S. scleroptilus and S. securiger, mainly in the shape of the male flagellomere I, which is extended distally to wrap around flagellomere II (Fig. 54). In all other species, the distal extension of flagellomere I is directed more posteriorly (e. g., Tendeiro 1960: fig. 6). Solenodes ningxiaensis can be separated from the four species listed above by having completely solenoid male genitalia, with no trace of a mesosome. In S. dispar and S. scleroptilus, the remnant of the mesosome is large, with a central rod-like structure (Kéler 1953: fig. 20 c; Gustafsson et al. 2024 a; fig. 588), in S. capitatus this structure appears to extend beyond the distal tips of the parameres (Kéler 1940: fig. 54) and in S. securiger there are apparently paired thickenings at about half-length of the genitalia (Clay 1940: fig. 61 a). Solenodes ningxiaensis is further separated from S. securiger by having male flagellomere I with larger distal modification in the former than in the latter, and by the many fewer tergal setae in S. securiger than in S. ningxiaensis (see table in Clay 1940: 90). Solenodes ningxiaensis is further separated from S. scleroptilus by having narrower marginal carina (Fig. 54) and by the male tergal chaetotaxy, which in S. scleroptilus consists of six tcs on each side of tergopleurite II, four tcs on each side of tergopleurite III, two tcs on each side on tergopleurites IV – V, and one tcs on each side in tergopleurites VI – VII (see Kéler 1953: fig. 20 a). Moreover, in S. scleroptilus s 1 – 2 are apparently mesosetae (Kéler 1953: fig. 20 a), whereas in S. ningxiaensis these are microsetae. Solenodes ningxiaensis is further separated from S. capitatus by the following characters: marginal carina narrower in S. ningxiaensis (Figs 52 – 53) than in S. capitatus; lateral lobes of female vulval margin narrower in S. capitatus than in S. ningxiaensis (Fig. 57). Setal numbers of the abdomen were not given for S. capitatus by Kéler (1940), but from his fig. 54, it is clear that S. capitatus is more setose; for instance, tergopleurites II – III of S. capitatus have at least 10 tcs + tps illustrated on each side, whereas in S. ningxiaensis there are only 6 – 7 on each side of these segments. In the female, Kéler’s illustration appears to have one more tcs seta in each side of at least segments II – III and VI – VII. Solenodes ningxiaensis is further separated from S. dispar by the following characters: distal expansion of male flagellomere I less pronounced in S. ningxiaensis (Fig. 54) than in S. dispar; lateral bulges of female vulval margin more pronounced in S. ningxiaensis (Fig. 57) than in S. dispar; female tergopleurites III – VII with rows of microsetae in S. dispar, but these are absent in S. ningxiaensis (Fig. 53); frons more gently rounded in S. ningxiaensis (Fig. 54) than in S. dispar.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753375FFFB769D6938D810FCB9.taxon	description	Descriptions Both sexes. Head relatively wide, frons gently rounded (Fig. 54). Marginal carina narrow, with sinuous inner margin. Head chaetotaxy as in Figs 53 – 54. Antennae sexually dimorphic. Temples similar between sexes. Thoracic and abdominal segments as in Figs 52 – 53. Male. Scape slightly longer than in female but not swollen (Fig. 54); flagellomere I with slight distal expansion wrapping around flagellomere II. Abdominal chaetotaxy: tergopleurites II – VI with rows of 4 – 7 tcs + tps on each side, numbers decreasing gradually in more posterior segments; tergopleurites II – III with four psc on each side, tergopleurites IV – VIII with three psc on each side; ventral side of abdominal segments II – VIII with one sts on each side. Male genitalia solenoid, with parameres fused to basal apodeme and mesosome absent. Measurements as in Table 1. Female. Scape shorter and flagellomere I not modified (Fig. 55). Much of central dorsal head lightly sclerotised, but the contrast between central area and lateral is only apparent laterally and posteriorly (Fig. 53). Abdominal chaetotaxy: tergopleurites II – V with 4 – 5 tcs on each side, tergopleurites VI – VIII with 1 – 2 tcs on each side; tergopleurite II with one psc on each side, tergopleurites III – VII with three psc on each side, with innermost seta gradually being more median in posterior segment, tergopleurite VIII with four psc, the two more median on each side being markedly shorter than the two lateral; tps microsetae present on segments VI – VII; ventral side of abdominal segments II – VIII with one sts on each side. Vulval margin with clear lateral bulges (Fig. 57); chaetotaxy: 8 – 12 long, slender vms on each lateral lobe, with 4 – 8 shorter setae submarginal to these; 4 – 5 thorn-like vss on each side centrally, with 5 – 8 short, slender vos scattered centrally on each side. Measurements as in Table 1.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753375FFFB769D6938D810FCB9.taxon	discussion	Remarks: One slide (box E 0026135, slide 25) with the same collection data as the holotype but labelled “ Lot 19 - IV ”, thus presumably from the same host individual, contains one female of an unidentified Complectodes, suggesting that Solenodes and Complectodes may co-occur on the same host individuals, but it needs confirmation.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753375FFFB769D6938D810FCB9.taxon	materials_examined	Material examined Types: Ex Alectoris magna: Holotype ♂: Haiyuan, Ningxia Province, China, Apr. 1984, coll. Xueli Bai, “ Lot 19 - VII ” box E 0026135, slide 32 (NHMC). Paratypes: 2 ♀, same data as holotype, “ Lot 19 - I ”, box E 0026135, slides 31, 33. 1 ♀, same data as holotype, “ Lot 19 - B 11 ”, box E 0026076, slide 87 (NMHC). Non-types: Two nymphs, same data as holotype, box E 0026193, slides 51 – 52.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753309FFFC769D6BF8DD42F895.taxon	type_taxon	Type species: Goniodes graecus Liu, 1994: 173.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753309FFFC769D6BF8DD42F895.taxon	diagnosis	Diagnosis. Complectodes belongs to the Oulocrepis - group, but it can be separated from the other genera by the following combination of characters (Table 2): male scape expanded compared to that of female, with a large, blunt process bearing one seta at its base; male flagellomere I extended into a slender, tapering distal process; mesosome and parameres present and separate from basal apodeme; parameres and other genitalic elements unique within Ischnocera; triangular process present lateral to female vulval margin; vulval margin with narrow lateral lobes bearing all the marginal setae. Complectodes can be separated from Goniodes sensu stricto by the following characters: pronotum flaring posteriorly and pteronotum with postero-lateral corners flattened so that both lpts and ipts are lateral in Goniodes (Figs 1, 3), but pronotum not flaring and pteronotum not modified, placing ipts sublaterally in Complectodes (Figs 58 – 59); both sexes of Goniodes with more than three sts on each side of abdominal segments II – VI (Figs 1, 3), but with only one sts on each side in Complectodes (Figs 58 – 59); male tergopleurite IX fused to central plate in Goniodes (Fig. 1), but plates separate in Complectodes (Fig. 58); male subgenital plate smaller, not expanded laterally, in Complectodes (Fig. 58) compared to Goniodes (Fig. 1); mesosternum, mss, and mets all present in Goniodes (Figs 1, 3), but absent in Complectodes (Figs 58 – 59); male scape with distally squamous process bearing no setae in Goniodes (Fig. 4), but with process not squamous and bearing seta at base in Complectodes (Fig. 60); pas and pns mesosetae in Goniodes (Figs 3 – 4), but macrosetae in Complectodes (Figs 59 – 60); coni bent posteriorly in both sexes in Goniodes (Figs 4 – 5), but not bent in males (Fig. 60) and barely arched posteriorly in females of Complectodes (Fig. 61); both mts 4 – 5 situated on occipital process in Goniodes (Fig. 4), but only mts 5 on occipital process sin Complectodes (Fig. 60); pos dorsal in Goniodes (Figs 3 – 4), but ventral in Complectodes, with that of female situated on tubercle (Figs 59 – 60); parameres roughly parallel, expanded but blunt distally, and with narrowly pointed heads in Goniodes (Figs 6 – 7), but strongly curved, with distal ends bifid and heads bearing complicated hatchet-like structures and partially enveloped in lateral sclerites in Complectodes (Figs 62 – 63); mesosome large, distally bifurcate, and associated proximally with bifurcate thickening of basal apodeme in Goniodes (Figs 6 – 7), but small, pointed distally and bifurcate thickening of basal apodeme absent in Complectodes (Figs 62 – 63); bifid inner structure of female genitalia present but triangular processes absent in Goniodes (Fig. 8), but bifid structure absent and triangular processes present in Complectodes (Fig. 64); vulval margin more or less flat, without lateral lobes in Goniodes (Fig. 8), but strongly convex with distinct lateral lobes in Complectodes (Fig. 64).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753309FFFC769D6BF8DD42F895.taxon	description	Description Both sexes. Large chewing lice of the Goniodes body louse ecomorph (Figs 58 – 59). Head about as broad as long, frons gently rounded (Fig. 60). Marginal carina narrow and uninterrupted, terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni prominent in male, less so in female, in both sexes not bent posteriorly. Antennae sexually dimorphic: male scape much swollen and elongated compared to that of female, with a non-squamous process on posterior margin, bearing one seta at its base; male pedicel longer than that of female and slightly curved; male flagellomere I extended distally into slender, tapering process. Eyes large. Temples rounded in male, more angular in female; occipital process present, more prominent in male than in female. Head chaetotaxy: ads, pas, os, pns all meso- or macrosetae; pos ventral, in female situated on tubercle; mts 1 and mts 3 only temporal macrosetae; only mts 5 situated on occipital process; dorsal head with few sensilla, and s 1 – 2 visible as micro- or mesosetae; s 6 present. Rhombic sclerite small, clearly separated from pronotum. Pronotum rounded posteriorly, with ppss situated on postero-lateral corner. Pteronotum roughly pentagonal, but with median section of posterior margin flattened; postero-lateral corners rounded, so that only lpts situated lateral, and ipts sublateral; smns without visible associates sensillum; mpts situated on or just posterior to pteronotum. Meso- and metasterna and associated setae absent. Proepimera large, curling around coxae II, but not fused medianly. Metepisterna not sclerotised medially. Abdomen rounded in male (Fig. 58), more elongate in female (Fig. 59); tergopleurite II not fused to pteronotum in either sex, and tergopleurites II – VIII medianly separated. Male tergopleurite IX not fused to central plate. Female tergopleurites IX – XI fused together, and medianly continuous, with postero-lateral end extended posteriorly to form small lobe with macroseta apically. Central sternal plates absent, but abdominal segments II – VI in both sexes with lateral accessory sternal plates. Male abdominal chaetotaxy: tergopleurites II – VII with one median tcs on each side; tergopleurites II – VIII with sparse rows of tps on each side, longer on II and VIII than on other segments; tergopleurites II – VIII with 2 – 3 psc on each side; ventral side of abdominal segments II – VIII with one sts on each side. Female abdominal chaetotaxy: tergopleurites II – VII with multiple tcs, gradually decreasing in number in more posterior segments; tergopleurites III – VI with at least one tps microseta lateral to tcs on each side; tergopleurites III – VIII with one tps mesosetae on each side, progressively shorter in more posterior segments; tergopleurites II – VI with two psc on each side, and tergopleurites VII – VIII with three psc on each side; ventral side of abdominal segments II – VIII with one sts on each side, on VI also with one microseta on each side. Male subgenital plate large, not expanded laterally or anteriorly; female subgenital plate absent, but slightly more sclerotised area present centrally anterior to vulval margin (Fig. 64). Subvulval and postvulval plates absent. Male genitalia unique within known Ischnocera (Figs 62 – 63). Basal apodeme relatively short, distally seemingly reduced to slender thickening on each side that reaches to proximal base of mesosome. Mesosome small, pointed distally, with accessory lateral plates. Dorsally, accessory plates arise proximally as slender, curved bands, that bend almost 90 ˚ laterally at midpoint of mesosome, and expand distally into densely rugose plates that overlap the parameres. Ventrally, these plates lie roughly parallel to the mesosome, being irregular in shape and tapering gently to reach about as far distally as mesosome. Gonopore and mesosome setae not visible. Parameres with complicated, hatchet-shaped heads bearing a median posterior process. Parameral blades strongly curved medianly to overlap distally, distal ends bifid and asymmetrical, with area between distal tips palmate by hyaline region. Overlapping parameral heads large, asymmetrical structure on each side, which curves around parameral heads laterally, and extends medially on ventral side to reach to mesosome; on this extension one sensillum is visible basally. Female genitalia without bifid structure, but with triangular processes laterally (Fig. 64). Vulval margin strongly convex, with lateral ends forming distinct lobes on which all marginal setae are situated. Vulval chaetotaxy: vms mesosetae clustered on lateral lobes, vss thorn-like setae in rows more centrally, and vos microsetae in roughly convergent rows.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFFD769D6889DFB2FE60.taxon	materials_examined	Type host: Scleroptila psilolaema (Gray, 1867) — moorland francolin.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFFD769D6983DD1FF957.taxon	description	(Figs 58 – 64)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFFD769D6983DD1FF957.taxon	materials_examined	Type host: Alectoris magna (Przevalski, 1876) — Przevalski’s partridge.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFFD769D6983DD1FF957.taxon	biology_ecology	Host distribution: Distantly related hosts within the Phasianidae.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFFD769D6983DD1FF957.taxon	distribution	Geographical range: Ethiopia, China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFFD769D6983DD1FF957.taxon	etymology	Etymology: The genus name Complectodes is derived from “ complector ”, Latin for “ to embrace ”, with the arbitrary ending “ - odes ” to denote a relationship with Goniodes. This refers to the unique shape of the male parameres.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFFD769D6983DD1FF957.taxon	discussion	Remarks: The two species of Complectodes have been collected from hosts belonging to different genera from different parts of the world, and both hosts are local endemics with very restricted ranges. This suggests that the diversity of Complectodes may be greatly underestimated, and that it may be found on many more hosts across Africa and Eurasia. The host genera of Complectodes species overlap with the host genera parasitised by species of Solenodes (see above). Furthermore, Alectoris magna is host to both Complectodes graecus and Solenodes ningxiaensis. Although it is not entirely clear from the slide labels, it seems likely that the specimens examined of S. ningxiaensis and those of C. graecus were derived from the same host individual. The co-occurrence of species of Complectodes and Solenodes on the same host species / individual requires verification by new collections. Gustafsson et al. (2024 a) redescribed Complectodes graecus. Hence, here we include illustrations of this species (Figs 58 – 64) and refer to the previous publication for a text description. Complectodes emersoni needs redescription and confirmation of its identity, as the two species are similar and the photos of C. emersoni published by Tendeiro (1965) are neither clear nor detailed. However, from the original description, C. emersoni appears to differ from C. graecus in the head shape of both sexes, the width of the marginal carina, and possibly the shape of the basal apodeme and mesosome, but these differences require confirmation.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFFD769D6983DD1FF957.taxon	materials_examined	Material examined Types of Complectodes graecus: Ex Alectoris magna: Holotype ♂, Haiyuan, Ningxia Province, China, Apr. 1984, coll. Bai Xueli, type collection slide 185035 (NHMC). Paratype: 1 ♂, Haiyuan, Ningxia Province, China, Apr. 1984, coll. Bai Xueli, type collection slide 185036 (NHMC). Non-types: 1 ♀, Haiyuan, Ningxia Province, China, Apr. 1984, coll. Bai Xueli, box E 0026135, slide 25 (NHMC).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFF2769D6F52DFB1FC65.taxon	type_taxon	Type species: Goniodes crossoptilon Clay, 1940: 58.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFF2769D6F52DFB1FC65.taxon	diagnosis	Diagnosis. Majiodes belongs to the Oulocrepis - group, but can be separated from all other genera in this group by the following combination of characters (Table 2): male scape swollen and elongated compared to that of female, with distally pointed process on posterior margin bearing setae at base; male flagellomere I extended distally into tapering process; male genitalia asymmetrical, with parameres and mesosome separate from basal plate; triangular processes present lateral to female vulval margin; lateral sections of vulval margin forming distinct lobes on which all marginal setae are situated. Majiodes can be separated from Goniodes sensu stricto by the following characters: male tergopleurite IX fused to central plate and inner margin of male genital opening with all setae of roughly equal length in Goniodes (Figs 1 – 2), but plates not fused (but overlap) and lateral-most setae of inner margin much longer than median setae in Majiodes (Figs 65 – 67); pteronotum with flattened postero-lateral corners and both lpts and pts lateral in Goniodes (Figs 1, 3), but corners rounded and only lpts lateral in Majiodes (Figs 65, 68); mesosternum, mss, and mets present in Goniodes (Figs 1, 3), but absent in Majiodes (Figs 65, 68); ventral side of abdominal segments II – VI with more than three sts on each side in Goniodes (Figs 1, 3), but with 0 – 2 sts on each side in Majiodes (Figs 65, 68); temples rounded in females of Goniodes (Fig. 3), but flaring into angles in Majiodes (Fig. 68); male scape with distally squamous and rounded posterior process bearing no setae in Goniodes (Fig. 4), but with distally non-squamous and pointed process bearing at least one seta at base in Majiodes (Fig. 69); coni bent posteriorly to overlap extensively with scape in Goniodes (Figs 4 – 5), but not bent posteriorly and only marginally overlapping with scape in Majiodes (Figs 69 – 70); pas and pns mesosetae in Goniodes (Fig. 4), but macrosetae in Majiodes (Fig. 69); occipital process narrow, with mts 4 – 5 both situated subapically on process in Goniodes (Fig. 4), but process broader, with only mts 5 situated on process in Majiodes (Fig. 69); basal apodeme comprising less than 50 % the length of male genitalia in Goniodes (Figs 6 – 7), but more than 50 % the length of genitalia in Majiodes (Figs 71 – 72); basal apodeme ventrally with bifurcated structure in Goniodes (Fig. 7), but without such structure in Majiodes (Fig. 72); parameres fused to basal apodeme, distally narrowing and convergent in Majiodes (Figs 71 – 72), but separate from basal apodeme, distally expanding and blunt in Goniodes (Figs 6 – 7); mesosome strongly asymmetrical in Majiodes (Figs 71 – 72), but symmetrical in Goniodes (Figs 6 – 7); bifid structure present but triangular processes absent in Goniodes (Fig. 8), but bifid structure absent and triangular processes present in Majiodes (Fig. 73); vulval margin with lateral setae-bearing lobes in Majiodes (Fig. 73), but without such lobes in Goniodes (Fig. 8); parts of subvulval area covered in dense microstructures in Majiodes (Fig. 73), but no such microstructures in Goniodes (Fig. 8).	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE75330FFFF2769D6F52DFB1FC65.taxon	description	Description Both sexes. Large chewing lice of the Goniodes body louse ecomorph (Fig. 65). Head broader than long (Fig. 69), frons gently rounded to somewhat flattened. Marginal carina uninterrupted, terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni prominent, not bent posteriorly. Antennae sexually dimorphic: in male, scape much swollen and elongated compared to that of female, with one distally pointed process bearing at least one seta basally; pedicel longer and slightly curved in male; flagellomere I extended distally I to curved, apically blunt process. Eyes large. Temples rounded laterally in male, extended laterally into angles in female (Fig. 68); occipital process present, broadly rounded. Head chaetotaxy: dsms, ads, pas, pns, os, avs 1 – 2, pcs all meso- or macrosetae; mts 1 and mts 3 only temporal macrosetae; pos ventral, not situated on tubercle in female; dorsal head sensilla s 1 – 2 and s 6 present, but central group of sensilla variable among species; mts 5 only seta situated on occipital process. Rhombic sclerite small, clearly separated from pronotum. Pronotum variable in shape, may flare slightly posteriorly, with ppss on postero-lateral corner. Pteronotum roughly trapezoidal to pentagonal, with posterior margin bulging only slightly medianly and postero-lateral corners rounded; only lpts lateral; ipts sublateral; smns not associated with visible sensillum; mpts on pteronotum. Meso- and metasterna and associated setae absent. Proepimera large, curling around coxae II, but not fused medianly. Metepisterna not sclerotised medially. Abdomen rounded in male, more elongate in female; tergopleurite II not fused to pteronotum in either sex, and tergopleurites II – VIII medianly separated. Male abdominal segment XI generally very broad. Male tergopleurite IX not fused to central plate, but may overlap; central plate of distinct shape (Fig. 66), with deeply bifid posterior margin and lateral, densely setose lobes; interior margin of genital opening convex, hidden underneath distal ends of central plate (Fig. 67), with lateralmost setae much longer than central setae. Female tergopleurites IX – XI fused together, and medianly continuous. Central sternal plates absent, but abdominal segments II – VI in both sexes with lateral accessory sternal plates. Male abdominal chaetotaxy: tergopleurites II – VII with one tcs on each side, flanked laterally by rows of tps that may be microsetae medially and gradually longer laterally, and lateral-most tps may be indistinguishable from median-most psc in length; tergopleurites II – VII or VIII with at least one psc on each side; ventral side of abdominal segments II – VIII with 1 – 2 sts on each side (variable among species), on segments II – VI with rows of microsetae on each side of sts. Female abdominal chaetotaxy: tergopleurites II – VIII with multiple (more than four) tcs on each side, gradually fewer in more posterior segments, flanked laterally by tps microsetae on at least tergopleurites III – VIII; tergopleurites II – VIII with 3 – 4 psc on each side; ventral side of abdominal segments II – VIII with one (rarely two) sts on each side, with rows of microsetae present on at least some segments, but distribution variable among species. Male subgenital plate large, expanded somewhat anteriorly to reach lateral accessory plate of segment VII, but not much laterally; female subgenital plate present but poorly sclerotised, flanked on each side by poorly sclerotised inner plates (subvulval plates?); postvulval plates present. Male genitalia large, reaching to around abdominal segment III when non-everted (Figs 71 – 72); basal apodeme constitutes more than 50 % the length of genitalia. Latero-distal corners of basal apodeme may be extended distally, and basal apodeme partially fused with parameres. Mesosome asymmetrical in all known species, and consists of several elements that are difficult to describe, and which vary among species. Parameres distally convergent, narrowing. Female genitalia without bifid structure internally, but with triangular processes laterally (Fig. 73). Vulval margin convex, with small lateral lobes. Vulval chaetotaxy: lateral lobes with multiple vms mesosetae, and small numbers of shorter setae (vss?) proximal and median of these; vss as rows of thorn-like setae median to lobes; vos microsetae scattered centrally, with anterior vos forming convergent but irregular rows. Parts of subvulval area with dense microstructures, the distribution of which varies among species.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF2769D6C50DFB5FB26.taxon	description	(Figs 65 – 73)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF2769D6C50DFB5FB26.taxon	materials_examined	Type host: Crossoptilon auritum (Pallas, 1811) — blue-eared pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF2769D6D45DF3EF919.taxon	materials_examined	Type host: Crossoptilon crossoptilon crossoptilon Hodgson, 1838 — Tibetan white-eared pheasant.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF2769D6D45DF3EF919.taxon	biology_ecology	Host distribution: Species in the genus Crossoptilon Hodgson, 1838.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF2769D6D45DF3EF919.taxon	distribution	Geographical range: China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF2769D6D45DF3EF919.taxon	etymology	Etymology: The genus name Majiodes is derived from “ maji ” (ḶḚ), the Chinese name for the eared pheasants of the genus Crossoptilon and the ending “ - odes ”, to symbolise the relationship between Majiodes and Goniodes.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF2769D6D45DF3EF919.taxon	discussion	Remarks: Both species of Majiodes were redescribed and illustrated by Gustafsson et al. (2024 a). Hence, we do not provide redescriptions here, but refer to the illustrations of the type species (Figs. 65 – 73). Note that the post-spiracular macrosetae of male tergopleurite VIII were inadvertently omitted from the illustration of Gustafsson et al. (2024 a: fig. 575), but have been added to Fig. 65, here.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF7769D6F17DE2DF992.taxon	type_taxon	Type species: Goniodes tetraophasis Chou & Liu, 1986: 317.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF7769D6F17DE2DF992.taxon	diagnosis	Diagnosis. The relationships between species of Sikongia and species placed in Goniodes by Price et al. (2003) are unknown, as the only known species of Sikongia is not similar morphologically to any other known species of goniodid. The closest relatives may be in Gonotyles, particularly in the subgenus Syrmatichares, with which Sikongia shares the following characters: meso- and metasterna and associated setae absent (Figs 74 – 75); male scape swollen and elongated, but with posterior process present only as a small bulge with an apical seta (Fig. 76); coni not bent posteriorly in either sex (Figs 76 – 77); male tergopleurite IX not fused to central plate (Fig. 78); female abdominal segment IX – XI extended posteriorly to form lateral seta-bearing lobes (Fig. 79); triangular processes and bifid structure both absent in female genitalia (Fig. 80). However, many of these characters are widely distributed within Goniodes sensu Price et al. (2003) and may be homoplasies. Sikongia can be separated from Gonotyles (Syrmatichares) by the following characters: pteronotum with three lpts on each side in Sikongia (Fig, 74), but with two lpts on each side in Gonotyles (Syrmatichares) (Fig. 37); ventral side of male abdominal segments IX – XI with multiple (more than five) macrosetae on each side in Gonotyles (Syrmatichares) (Fig. 37), but without ventral macrosetae except on distal margin in Sikongia (Fig. 74); tergopleurites with rows of tps microsetae separating tcs and psc in Gonotyles (Syrmatichares) (Figs 37 – 38), but with tps mesosetae in Sikongia (Figs 74 – 75) and only tergal microsetae intercalated with mesosetae near median end of tergopleurites; paratergal plate II in both sexes of Sikongia with dense rows of macrosetae one each side (Figs 74 – 75), but with few (fewer than three) ps on segment II in Gonotyles (Syrmatichares) (Figs 37 – 38); male genital opening with serrated sclerite on each side in Gonotyles (Syrmatichares) (Fig. 43), but without such sclerite in Sikongia (Fig. 78); female vulval margin not extending beyond distal margin of abdomen, strongly convex with lateral setae-bearing lobes in Gonotyles (Syrmatichares) (Fig. 44), but extending beyond distal margin of abdomen, deeply bilobed but without setae-bearing lobes in Sikongia (Figs 79 – 80); vulval setae divided into vms mesosetae, vss thorn-like setae, vss (?) slender setae, and vos microsetae in Gonotyles (Syrmatichares) (Fig. 44), but all vulval setae more or less similar as microsetae in Sikongia (Fig. 80). Male genitalia of Sikongia were not illustrated in detail by Chou & Liu (1986) and are absent in all known specimens of S. tetraophasis (see Gustafsson et al. 2024 a: 544 – 545). Based on the illustrations of Chou & Liu (1986: fig. 3 a), the male genitalia of S. tetraophasis differ from those of Gonotyles (Syrmatichares) in having parameres distinct and not fused to either basal apodeme or mesosome, and by having an elongated T-shaped structure on the distal basal apodeme. New material of S. tetraophasis is needed to evaluate the differences in the male genitalia adequately. Sikongia can be separated from Goniodes sensu stricto by the following characters: male scape with distally squamous process not bearing any setae in Goniodes (Fig. 4), but with only a slight seta-bearing bulge in Sikongia (Fig. 76); coni bent posteriorly in both sexes of Goniodes (Figs 4 – 5), but not in Sikongia (Figs 76 – 77); head setae ads, pas, pns, os all macrosetae in Sikongia (Fig. 76), but mesosetae in Goniodes (Fig. 4); mts 4 and mts 5 both situated on occipital process in Goniodes (Fig. 4), but only mts 5 on occipital process in Sikongia (Fig. 76); postero-lateral corners of pteronotum flattened so that both lpts (of which there are two on each side) and ipts are lateral in Goniodes (Figs 1, 3,), but rounded so that only lpts (of which there are three on each side) are lateral in Sikongia (Figs 74 – 75); mesosternum, mss, and mets all present in Goniodes (Figs 1, 3), but all absent in Sikongia (Figs 74 – 75); male tergopleurite IX fused to central plate in Goniodes (Fig. 1), but not fused in Sikongia (Fig. 74); female abdominal segment IX + X expanded distally to form setae-bearing lobes in Sikongia (Figs 79 – 80), but no such lobes in Goniodes (Fig. 3); male tergopleurites with rows of tps microsetae separating tcs and psc in Goniodes (Fig. 1), but with tps mesosetae in Sikongia (Fig. 74) and only tergal microsetae intercalated with tcs near median end of tergopleurites; paratergal plate II in both sexes of Sikongia with dense rows of macrosetae one each side (Figs 74 – 75), but with no ps on segment II in Goniodes (Figs 1, 3); both sexes of Sikongia with rows of microsetae on ventral surface of at least some segments (Figs 74 – 75), but no microsetal rows in Goniodes (Figs 1, 3); bifid structure of female genitalia present in Goniodes (Fig. 8), but absent in Sikongia (Fig. 80); female vulval margin not extending beyond distal margin of abdomen, straight with lateral setae-bearing patches in Goniodes (Fig. 8), but extending beyond distal margin of abdomen, deeply bilobed but without setae-bearing lobes in Sikongia (Figs 79 – 80); vulval setae divided into vms mesosetae, vss (?) slender setae, and vos microsetae in Goniodes (Fig. 8), but all vulval setae more or less similar as microsetae in Sikongia (Fig. 80). Based on the illustrations of the male genitalia of S. tetraophasis by Chou & Liu (1986), Sikongia can be separated from Goniodes sensu stricto on the following additional characters: mesosome elongated triangular, distally bifurcated, with rugose lateral corners and a distinct structure presumed to be the gonopore in Goniodes (Figs 6 – 7), but rounded rectangular with no distinct structures in Sikongia (Chou & Liu 1986: fig. 3 a); parameres widening distally in Goniodes (Figs 6 – 7), but tapering distally in Sikongia.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753300FFF7769D6F17DE2DF992.taxon	description	Description Both sexes. Large chewing lice of the Goniodes body louse ecomorph (Figs 74 – 75). Head broader than long, frons gently rounded to somewhat flattened (Fig. 76). Marginal carina uninterrupted, terminating proximally in elongated preantennal nodi. Ventral carina uninterrupted; clypeo-labral suture absent. Coni not bent posteriorly (Figs 76 – 77). Antennae sexually dimorphic: male scape swollen and elongated compared to that of female, with posterior margin bulging slightly at about mid-length, bearing one seta on the bulge; male pedicel swollen and slightly modified in shape compared to female; male flagellomere I extended distally into broad, distally tapering process. Eyes large. Temples somewhat angular laterally; occipital process present, in males directed somewhat lateral. Head chaetotaxy: dsms, ads, pas, os, pns, pcs and avs 2 all meso- or macrosetae; mts 1 and mts 3 only temporal macrosetae; only mts 5 situated on occipital process; head sensilla s 1 – 2 and s 6 present in both sexes; in male a patch of dorsal sensilla present centrally in post-antennal head. Rhombic sclerite present, may be fused to pronotum. Pronotum rounded posteriorly, with ppss on postero-lateral corner. Pteronotum roughly pentagonal, with postero-lateral corners rounded so that only lpts (of which there are three) are lateral; ipts sublateral; smns without associated sensillum; mpts on or just posterior to pteronotum. Meso-and metasterna and associated setae absent. Proepimera large, curling around coxae II, but not fused medianly. Metepisterna not sclerotised medially. Abdomen rounded in male, more elongate in female (Figs 74 – 75); tergopleurite II not fused to pteronotum in either sex, and tergopleurites II – VIII medianly separated. Male tergopleurite IX not fused to central plate, but may overlap (Fig. 78). Female tergopleurites IX – XI all fused together and medianly continuous, with lateral section extended distally to form setae-bearing lobes; central part of female tergopleurite IX – XI with distinct reticulation (Fig. 79). Central sternal plates absent, but abdominal segments II – VI in both sexes with lateral accessory sternal plates. Male abdominal chaetotaxy: tergopleurites II – VII with rows of tcs + tps, with small numbers of microsetae intercalated with tcs + tps in more anterior segments, particularly near the median ends of tergopleurites; tergopleurites II – VIII with at least three psc on each side, but median ones indistinguishable in length from tcs + tps rows; ventral side of abdominal segments II – VIII with one (two on VII) sts on each side, on II – VI with additional rows of microsetae. Female abdominal chaetotaxy: tergopleurites II – III with dense double rows of tcs + tps, tergopleurites IV – VIII each with one row of tcs, gradually more sparse in more posterior segments, with (tps?) microsetae around median ends of tergopleurites on some more posterior segments; on tergopleurites II – III psc e cannot be distinguished from tergal rows, on tergopleurites IV – VII three psc on each side; on tergopleurite IX – XI a row of short mesosetae submarginally (Fig. 79); ventral sides of abdominal segments II – VIII with one sts on each side, at least on segments III – V, with additional microsetae. Male subgenital plate small; female subgenital, subvulval, and postvulval plates absent. Male genitalia not present in any examined specimens; based on illustrations by Chou & Liu (1986: fig. 3 a) of (now lost) type specimens, the basal apodeme is long (reaching abdominal segment II with not everted), not fused to either mesosome or parameres, but bears an elongated T-shaped structure centrally. Mesosome rounded rectangular, seemingly without any distinct structures apart from perhaps centrally. Parameres elongated, slender, tapering distally. Female genitalia without bifid structure or triangular processes (Fig. 80). Vulval margin extended distally to reach beyond distal margin of abdomen, forming a narrowly pointed lobe on each side separated by a narrow, deep emargination. Vulval setae not easily separated into sets, with numerous (more than 60) microsetae or sensilla on each side scattered in subvulval area.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753305FFEC769D6F59DF4AFE49.taxon	description	(Figs 74 – 80, 83)	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753305FFEC769D6F59DF4AFE49.taxon	materials_examined	Type host: Tetraophasis obscurus (Verreaux, 1869) — chestnut-throated monal-partridge.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753305FFEC769D6F59DF4AFE49.taxon	biology_ecology	Host distribution: One species of Tetraophasis Elliot, 1871.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753305FFEC769D6F59DF4AFE49.taxon	distribution	Geographical range: Sichuan Province, China.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753305FFEC769D6F59DF4AFE49.taxon	etymology	Etymology: Sikongia is named in honour of Sikong Liu, formerly from the Beijing Museum of Natural History, in recognition of his work on Chinese goniodids, including the description of the type species of this genus.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
03DBDE753305FFEC769D6F59DF4AFE49.taxon	discussion	Remarks: Sikongia tetraophasis was recently redescribed and illustrated by Gustafsson et al. (2024 a). In addition to the genus-level description above, we refer to this publication for further details of the species. Illustrations provided here are for comparative purposes. Also, note the comments of Gustafsson et al. (2024 a) about the status of the type series and the specimens illustrated here.	en	Gustafsson, Daniel R., Li, Zhu, Tian, Chunpo, Ren, Mengjiao, Sun, Xiuling, Zou, Fasheng (2025): Revision of genera in Goniodidae (Phthiraptera: Ischnocera) parasitising gamefowl (Aves: Galliformes) with descriptions of six new genera, one new subgenus and seven new species. Zootaxa 5731 (1): 1-99, DOI: 10.11646/zootaxa.5731.1.1, URL: https://doi.org/10.11646/zootaxa.5731.1.1
