taxonID	type	description	language	source
03DFB84ABB0B2B1E00F12EE2FE1AF397.taxon	type_taxon	Type species: Tico emmettcarri Bahder & Bartlett, 2021	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0B2B1E00F12EE2FE1AF397.taxon	diagnosis	Amended Diagnosis (updated from Bahder et al. 2021, 2022, Echavarria et al. 2022). Small (~ 2.5 – 4.2 mm), cenchreine derbids. Vertex roughly triangular or trapezoidal, widest posteriorly, disc concave, lateral margins keeled (bearing row of sensory pits), with transverse carinae near fastigium. Frons moderately compressed (length at midline about 4 x dorsal width), median carina absent, lateral carinae keeled, frontoclypeal margin nearly linear. Antennae short. Forewing with ScP branch reaching wing margin preceding apex of clavus (resulting in long marginal cell). Pygofer opening with expansions on lateral margin of opening, ventral margin without medioventral lobe. Gonostyli long and spatulate, apex rounded, with a medially directed dorsal lobe near midlength (or beyond; medial lobes absent on ventral margin) and sclerotized process on inner margin. Aedeagus nearly bilaterally symmetrical. Anal tube elongate, in lateral view ventral margin weakly concave, apex inflected downward, apex truncate; in dorsal view broadly spatulate.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0A2B1B00F12808FF3AF17D.taxon	description	(Figures 2 – 7)	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0A2B1B00F12808FF3AF17D.taxon	materials_examined	Type Locality. Costa Rica, Puntarenas Province, Osa Peninsula, La Tarde EcoLodge.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0A2B1B00F12808FF3AF17D.taxon	diagnosis	Diagnosis. Small, pale planthopper, ornately marked with red bands in apical cells of forewings, a large red patch in clavus, seven fuscous patches on forewing and a pair of red spots on mesonotum. Lateral margins of pygofer with relatively short, triangular projection. Endosoma with very wide and flattened projections with distinct serrations along margins.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0A2B1B00F12808FF3AF17D.taxon	description	Description. Color. General body color ivory white to pale yellow (Fig. 2); eyes red; two red spots on mesonotum, near tegulae; forewings with large red patch in clavus and red patches in apical cells; seven fuscous spots present primarily in posterior half of wing, one at base of claval cell. Structure. Body length (with wings), male 4.2 mm (n = 1, Table 2). Head. In dorsal view, vertex trapezoidal in shape (posterior margin about 1.3 x wider than vertex midlength), anterior margin linear at apical transverse carina, posterior margin broadly excavated, medially irregularly sinuate, single row of sensorial pits along lateral margins (Fig. 3 A). In lateral view, generally rounded, slightly angulate at fastigium (indicating position of transverse carina at apex of vertex), slightly projecting above and beyond eyes, lateral ocelli below ventral margin of eyes (Fig. 3 B). In frontal view, frons about 2.2 x longer at midlength than at frontoclypeal margin, dorsal margin linear (at transverse carina of vertex), lateral margins of frons constricting slightly from dorsal margin, expanding just below ventral margin of eyes to frontoclypeal suture; single row of weakly developed sensorial pits present along lateral keels of frons (Fig. 3 C). Thorax. Pronotum, in dorsal view, shorter at midline than vertex (0.67 x), with anterior margin convex, posterior margin concave; tricarinate, lateral carinae prominent and keeled, just reaching posterior margin, two small depressions on either side of median carina (Fig. 3). In lateral view, paradiscal region foliate, forming C-shape behind antennae. Mesonotum in dorsal view. about 1.7 x as long at midlength than vertex + pronotum cumulatively and approximately 1.25 x wider than long at midpoint, weakly tricarinate (Fig. 3). Metatibiotarsal spinulation 8 - 6 - 6. Forewings elongated, weakly spatulate; RA unbranched, RP 2 - branched, MP 5 - branched, CuA 2 - branched, branches anastomosed to form a closed C 5 cell with composite vein reaching margin, Claval apex near wing midlength (Fig. 4). Terminalia. Pygofer in lateral view with anterior margin sinuate, posterior margin irregularly sinuate, subtriangular projection arising in dorsal half, slightly curved dorsad (Fig. 5); in ventral view, posterior margin slightly concave (Fig. 5). Gonostyli in lateral view scoop-like, base expanded, projecting ventrally, ventral margin irregularly rounded, dorsal margin irregularly sinuate, large, subtriangular dorsal projection present, apex curved ventrally on outer lateral side, spinose (Fig. 5); in ventral view, widest at base, appearing rectangular, inner margin corner invaginated, constricting to narrowest point, immediately expanding, constricting subapically, apex subtriangular, paired sclerotized spines on inner margins visible, curving cephalad, posterior spine more robust and longer than anterior spine (Fig. 5). Aedeagal shaft simple; endosoma highly complex, comprises 10 processes in symmetrical pairs, with first pair (E 1 & E 2) arising at apex, broad and rounded at base, narrowing at midpoint, apices bifurcated; second pair (E 3 & E 4) arising subapically, broadly flattened, reaching midpoint, apices serrated and strongly bifurcated with outer bifurcation (E 3 a & E 4 a) being longer and more slender than inner bifurcations (E 3 b & E 4 b), curved distad, angled cephalad; third pair (E 5 & E 6) very broadly flattened, serrated on outer margins in distal half, apices spinose, curved mesad, angled cephalad, nearly reaching aedeagal base, lateral margins cupped dorsally; fourth pair (E 7 & E 8) similar to E 5 and E 6, extending slightly closer to aedeagal base, arising ventral to E 5 and E 6; fifth pair (E 9 & E 10) arising at midpoint, angled cephalad, reaching as far as E 7 and E 8, broad, not flattened or serrated (Figs 6 & 7). Anal segment in lateral view irregularly sinuate on ventral margin, apex concaves, extending just beyond lateral process of pygofer; in dorsal view, club-like, apex irregularly rounded, blunt (Fig. 5). Plant Associations. Unknown palm (Arecaceae).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0A2B1B00F12808FF3AF17D.taxon	distribution	Distribution. Costa Rica (Puntarenas Province), Osa Peninsula.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0A2B1B00F12808FF3AF17D.taxon	etymology	Etymology. The specific name is a patronym for Mr. Leonardo Rosseau, the nephew of the senior author.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0A2B1B00F12808FF3AF17D.taxon	discussion	Remarks. Superficially, T. leo sp. nov. most closely resembles the type species, T. emmettcarri based on body color and wing pattern. However, the former has many more distinct spots (both red and black on the wings and body) than T. emmettcarri. In addition, T. leo sp. nov. possesses a lateral process on the pygofer (lacking in T. emmettcarri) and a very broadly flattened and serrated endosoma whereas T. emmettcarri possesses a much narrower and compact endosoma. Finally, the medial process of the gonostyli differs significantly among the two species where T. leo sp. nov. has a strongly curved pair of spine-like processes (Fig. 5 E), whereas T. emmettcarri has an elongate ridge with minute serrations on the surface (Bahder et al. 2021, fig. 8 B).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0A2B1B00F12808FF3AF17D.taxon	materials_examined	Material Examined. Holotype male, “ Costa Rica, Puntarenas Pr., Drake / Bahia Drake Church lot / 12 - VI- 2021 / Coll.: B. W. Bahder / Sweeping coconut / Tico leo ♂ ” (FLREC). Holotype preserved in 95 % ethanol at - 80 ° C.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0E2B1700F12EC7FE2AF3ED.taxon	description	(Figures 8 – 11)	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0E2B1700F12EC7FE2AF3ED.taxon	materials_examined	Type Locality. Costa Rica, Puntarenas Province, Osa Peninsula, Palma.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0E2B1700F12EC7FE2AF3ED.taxon	diagnosis	Diagnosis. Small, pale planthopper, ornately marked with red band running near posterior margin of forewing with large fuscous patch within red band near forewing apex. Medioventral process of pygofer present, small, triangular. Sclerotized structure on inner margin of gonostyli elongate and serrated.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0E2B1700F12EC7FE2AF3ED.taxon	description	Description. Color. General body color yellowish ivory white (Fig. 8); eyes pale green; lateral carinae of vertex with fuscous anteriorly. Head in lateral view with fuscous marking on anterodorsal margin of heads. Females with reddish patch in lateral compartments of mesonotum (absent in males). Forewings with large red band near commissural margin from near wing base to wing apex (broken near claval margin in males; with darkened spot in distal portion of C 3 cell), two small fuscous band arising on anterior margin; females with two additional fuscous spots in proximal portion of precubital cell and a third near midlength of CuP (Fig. 8). Structure. Body length (with wings), male 3.2 – 3.9 mm (n = 3, Table 2), female 4.0 mm (n = 3, Table 2). Head. In dorsal view, vertex trapezoidal in shape (posterior margin about 1.9 x wider than vertex midlength), anterior margin linear at transverse carina, posterior margin pentagonally excavate, medially irregularly sinuate, single row of sensorial pits along lateral margins (Fig. 9). In lateral view, generally rounded, slightly angulate at fastigium (marking location of transverse carinae at vertex apex), slightly projecting above and beyond eyes, lateral ocelli below eyes (Fig. 9). In frontal view, frons about 2.3 x longer at midlength than at frontoclypeal margin, dorsal margin of frons linear, frons constricting slightly from dorsal margin, expanding just below ventral margin of eyes to frontoclypeal suture; single row of weakly developed sensorial pits present along lateral margins of frons (Fig. 9). Thorax. Pronotum, in dorsal view, about as long as vertex at midline, with anterior margin convex, posterior margin concave; tricarinate, lateral carinae prominent, keeled, reaching posterior margin, median carina nearly obsolete (Fig. 9). In lateral view, paradiscal region foliate, forming C-shape behind antennae. Mesonotum, in dorsal view, about 1.4 x as long at midlength than vertex + pronotum cumulatively and approximately 1.25 x wider than long at the midpoint, weakly tricarinate (Fig. 9). Metatibiotarsal spinulation 8 - 6 - 6. Forewings weakly spatulate; RA unbranched, RP 2 - branched, MP 5 - branched, CuA 2 - branched (branches anastomosed to form a closed C 5 cell with composite vein reaching margin), claval apex near wing midlength (Fig. 10). Terminalia. Pygofer in lateral view with anterior and posterior margin irregularly sinuate, narrowest dorsally, expanding immediately, constricting slightly after midpoint, expanding slightly at base; in ventral view, medioventral process present, small, subtriangular (Fig. 11). Gonostyli in lateral view spatulate, basal expansion extending ventrally, fin-like, gently curved on ventral margin, rounded at apex; dorsal process present, arising subapically, bifurcated, anterior bifurcation more elongate and curved than posterior bifurcation (Fig. 11); in ventral view, widest at base, expansions subtriangular, constricting, expanding at midpoint to widest point, narrowing to gently hooked apices, inner sclerotized processes elongate and serrated (Fig. 11). Aedeagal shaft with paired serrated flanges on lateral margins near midpoint (A 1 & A 2) (Fig. 12 & 13); endosoma complex with six pairs of processes, first pair (E 1 & E 2) arising subapically curving mesad, angled dorsad; second pair (E 3 & E 4) strongly bifurcated angled cephalad, inner bifurcation (E 3 a & E 4 A) slender, spinose, angled mesad, outer bifurcation (E 3 b & E 4 b) more robust, rounded at apex; third pair (E 5 & E 6) elongate, angled cephalad, curved dorsad, spinose; fourth pair (E 7 & E 8) angled cephalad and mesad, spinose, curved gently ventrad; fifth pair (E 9 & E 10) similar to E 7 and E 8, arising where E 7 and E 8 terminate; sixth pair spinose, angled cephalad and mesad, curved dorsad, nearly reaching aedeagal base (Figs. 12 & 13). Anal segment in lateral view curved on ventral margin, irregularly sinuate on dorsal margin, apex spatulate, subquadrate (Fig. 11); in dorsal view, goblet-shaped, apex slightly invaginated (Fig. 11). Plant Associations. Unknown palm (Arecaceae).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0E2B1700F12EC7FE2AF3ED.taxon	distribution	Distribution. Costa Rica (Puntarenas Province), Osa Peninsula.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0E2B1700F12EC7FE2AF3ED.taxon	etymology	Etymology. The specific name is a patronym for Mr. Logan Vetter.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0E2B1700F12EC7FE2AF3ED.taxon	discussion	Remarks. Similar in general appearance to both T. leo sp. nov. and T. emmettcarri, it has significantly fewer spots on the forewings (possessing only one near apex). Features of the genitalia in T. logani sp. nov. are much more similar to T. emmettcarri than to T. leo sp. nov.; lack of lateral process on pygofer, a more compact / narrow endosoma and T. logani sp. nov. has an elongate ridge on the inner margin of the gonostyli with minute serrations on the surface, very similar to the form observed in T. emmettcarri. Despite the similarities in general patterns of the genitalia to T. emmettcarri that would suggest a closer relationship than to T. leo sp. nov., the structures of the genitalia are all distinct among T. logani sp. nov. and T. emmettcarri.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0E2B1700F12EC7FE2AF3ED.taxon	materials_examined	Material Examined. Holotype male, “ Costa Rica, Puntarenas Pr. / Drake / 18 - VI- 2021 / Coll.: B. W. Bahder / Sweeping palm / Tico logani ♂ ” (FLREC). Paratypes (1 male, 2 females) same as holotype (FLREC).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB0E2B1700F12EC7FE2AF3ED.taxon	discussion	Sequence Data. For the COI gene, a 560 bp sequence was obtained, for the 18 S gene, a 1,334 bp sequence was obtained, for the D 8 expansion region of 28 S, a 714 bp sequence was obtained and for the D 9 – D 10 expansion region of 28 S, a 799 bp sequence was obtained. All sequences were deposited in GenBank. Accession numbers are presented in Supplementary Table 1. The Maximum Likelihood analyses for individual loci resulted in strong bootstrap support for the monophyly of Tico based on 18 S and all regions of 28 S analyzed; 98, 98 and 100 respectively for 18 S, D 8 and D 9 – D 10 of 28 S (Fig. 14). Additionally, both new taxa resolved within Tico (Fig. 14). The phylogeny derived from concatenated data for COI, 18 S and 28 S demonstrated strong support (100 / 100) for the monophyly of Tico and strong support (90 / 97) for T. logani sp. nov. resolving adjacent to T. emmettcarri and strong support (78 / 99) for T. leo sp. nov. resolving adjacent T. pseudosororius (Fig. 15). Remarks. The novel taxa possess morphological characters that are consistent with currently described species of Tico and molecular data also supports the placement of the novel taxa within Tico. While the molecular data suggests there are two distinct lineages within Tico, there are no apparent morphological features that are consistent with this dichotomy. Both the wing pattern and general structure of the terminalia of T. logani sp. nov. seemed more similar to T. emmettcarri than to other members in the genus, a relationship reflected in the molecular phylogeny. The morphological characters of the terminalia observed in T. leo sp. nov. in general, differed significantly from other members of the genus. Tico leo sp. nov. is unusual in the genus by possessing a process on the lateral portion of the pygofer (Fig. 5 A).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB042B1100F12BEDFD6BF78C.taxon	description	(Fig. 16)	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB042B1100F12BEDFD6BF78C.taxon	distribution	Distribution. St. Vincent and the Grenadines.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB042B1100F12BEDFD6BF78C.taxon	discussion	Remarks. This species was considered in Bahder et al. (2021) and compared it with Tico. At the time, it was not included in Tico because face was relatively broad and the lateral margins of the pygofer bares a large falciform processes, which was not found on the species of Tico known at that time and molecular data was lacking. Fennah (1952: 125) noted that Cenchrea dorsalis “ stands strongly apart from the other species ” and that they “ may be regarded as comprising two subgenera ”. We concur that C. dorsalis is unlike the other species in Cenchrea and the remaining taxa are an uncomfortable fit for the genus. Tico leo sp. nov., like Cenchrea exquisita, bears a lateral process on the opening of the pygofer. While the lateral process of the opening of the pygofer is much more ‘ exuberant’ (Fennah 1952: 131) than other members of Tico, the other features of the male terminalia of Cenchrea exquisita are similar enough to be congeneric. Also, the general coloration of C. exquisita (see Bahder et al. 2021, fig. 3 A) is comparable to Tico. While C. exquisita remains unusual in possessing a relatively broad face, and is found in the Caribbean, rather than the mainland, the morphological similarities listed above, plus strong molecular support for placement of T. leo sp. nov. (a species also possessing a lateral process on pygofer) in Tico, it is clear T. exquisita belongs in the genus Tico, and so herein Tico exquisita (Uhler, 1895), comb. nov. is transferred to Tico.	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
03DFB84ABB042B1100F12BEDFD6BF78C.taxon	materials_examined	Material Examined. “ ♂ // ST. VINCENT, St. / PatrickP., Belleisie / Hill, N. slope, 9 - 3 - 1991 / C. W. & L. B. O’Brien // UDCC _ TCN 00104866 ” (University of Delaware).	en	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2026): New planthoppers in the genus Tico from Costa Rica (Hemiptera: Auchenorrhyncha: Derbidae) and an updated molecular phylogeny of the New World Cenchreini. Zootaxa 5849 (1): 42-58, DOI: 10.11646/zootaxa.5849.1.2, URL: https://doi.org/10.11646/zootaxa.5849.1.2
