taxonID	type	description	language	source
03DF87FEFFF8F132FF3154A6FA6F872F.taxon	discussion	This species group (sensu Moravec 2007), which has been recently supplemented by two new species (Moravec et al 2020, Moravec & VybÍral 2022) and the new species described here, now comprises 15 species of a medium body size (generally smaller than those within other species-groups of the nominotypical subgenus, except for the new species described here). Head robust with mostly notably short temples. Labrum 6 – 10 - setose with 4 – 8 and 2 lateral setae; shape of labrum rather variable in both sexes, but in most species male labrum short with truncate anterior margin while female labrum more prolonged with slightly elongate anterior lobe, almost semicircular or with slightly truncate, anterior margin (except for the uniformly shape labrum in new species described here). Lacinia generally shorter than that in other species groups and more distinctly (though gradually), spatulate-dilated. Antennae black, antennomere 1 – 4 usually with metallic blue lustre. Elytral surface with faint or moderate discal impression, punctation covering whole elytral surface, exceptionally partly effaced on extreme base; punctures predominantly isolated and nearly regular with narrow, sometimes carinulate, or else wider and flat intervals, only occasionally anastomosing in chains with declined lateral intervals (except for P. (P.) meridionale, whose elytral punctation is partly denser and notably irregularly anastomosing), but never cavernous; elytral apices variously shaped. Setal vesture whitish, in two species partly or predominantly black, exceptionally ferrugineous (in P. (P.) meridionale). Legs black to black-brown. Aedeagus variously shaped; that in P. (P.) majunganum extraordinarily short.	en	Moravec, Jiří, Wiesner, Jürgen (2026): New or rare Madagascar tiger beetles- 27. Pogonostoma (Pogonostoma) andasibense sp. nov., and a revised key to the Pogonostoma (P.) elegans species-group (Coleoptera: Cicindelidae). Zootaxa 5748 (2): 215-229, DOI: 10.11646/zootaxa.5748.2.3, URL: https://doi.org/10.11646/zootaxa.5748.2.3
03DF87FEFFFFF13AFF3157BAFAD48529.taxon	description	(Figs 1 – 20)	en	Moravec, Jiří, Wiesner, Jürgen (2026): New or rare Madagascar tiger beetles- 27. Pogonostoma (Pogonostoma) andasibense sp. nov., and a revised key to the Pogonostoma (P.) elegans species-group (Coleoptera: Cicindelidae). Zootaxa 5748 (2): 215-229, DOI: 10.11646/zootaxa.5748.2.3, URL: https://doi.org/10.11646/zootaxa.5748.2.3
03DF87FEFFFFF13AFF3157BAFAD48529.taxon	materials_examined	Type locality. Analamazaotra National Park (previously Andasibe-Mantadia N. P. with Perinet Special Reserve), Mangoro Region, Eastern Madagascar. Type material. Holotype ♂, labelled: “ Madagascar 19 ° 00 S, / 48 ° 27 E. Toamasina prov., Andasibe Nat. Park, Perinet / 19. - 31.12.2001, 950 m, / V. Dolin et R. Andreeva leg. ” [printed] // “ Holotype / Pogonostoma (s. str.) / andasibense sp. nov. / det. JiřÍ Moravec & / Jürgen Wiesner 2025 “ [red label, printed]. Allotype ♀, labelled: “ Madagascar E / Alaotra Mangoro Reg. / Andasibe-Mantadia NP / 1000 m., 17. I. - 2. II. 2020 / leg. S. Dementiev ” [printed] // “ Allotype / Pogonostoma (s. str.) / andasibense sp. nov. / det. JiřÍ Moravec & / Jürgen Wiesner 2025 ” [red label, printed]. Type depository. Both type specimens deposited in ZSM (temporarily in JWCM).	en	Moravec, Jiří, Wiesner, Jürgen (2026): New or rare Madagascar tiger beetles- 27. Pogonostoma (Pogonostoma) andasibense sp. nov., and a revised key to the Pogonostoma (P.) elegans species-group (Coleoptera: Cicindelidae). Zootaxa 5748 (2): 215-229, DOI: 10.11646/zootaxa.5748.2.3, URL: https://doi.org/10.11646/zootaxa.5748.2.3
03DF87FEFFFFF13AFF3157BAFAD48529.taxon	diagnosis	Differential diagnosis. Pogonostoma (Pogonostoma) andasibense sp. nov. may resemble Pogonostoma (Pogonostoma) elegans (Brullé in Audouin & Brullé, 1834), yet apart from its somewhat larger size and generally wider pronotal disc, which is covered with finer sculpture (Figs 13 – 14), the new species clearly differs in having bulging head temples (Figs 1 – 2, 5 – 6) and 10 - setose labrum (with 8 anterior setae), which is uniformly shaped in both sexes (Figs 8 – 9). In contrast, the labrum in P. (P.) elegans (as in most other taxa of this species-group) has primarily only 4 – 6 anterior setae and is sexually dimorphic, notably anteriad-prolonged in females (see Moravec 2007, figs 867 – 872, where that of the female lectotype (MNHN) is in fig. 870). The pronotal disc in P. (P.) elegans is always covered with coarser, more fragmented rugae and has mostly subparallel lateral margins (see Moravec 2007, figs 857 and 1733 of the male lectotype and figs 1734 – 1735 of females where the fig. 1735 is of the holotype (SDEI) of synonymous Pogonostoma klugi W. Horn, 1893). Males of these two species are immediately distinguished by the boomerang-bent aedeagus with rounded apex in the new species, while the aedeagus in P. (P.) elegans is much straighter and with short, dorsally hooked apex (see Moravec 2007, figs 860, 862 – 863, 865, where the fig. 865 shows aedeagus apex of the lectotype (MFNB) of synonymous Pogonostoma coerulescens Klug, 1835). Last but not least, in contrast to the whitish setae densely covering dorsal surface of meso- and metafemora, the setae on the femoral surface in P. (P.) elegans are sparser, stiffer and blackened. Other species within the group resembling this new species are Pogonostoma (Pogonostoma) brullei (Laporte de Castelnau & Gory, 1835) and P. (Pogonostoma) differens Cassola & Andriamampianina, 2001. While the former is known only from its female holotype (in MNHN) – see Moravec (2007, figs 1002 – 1004, 1747), the former is known only from its male holotype (in FCCR-MCZR) – see Cassola & Andriamampianina (2001, figs 1 – 3) and Moravec (2007, figs 1009 – 1012 and 1748). As the differences may represent only common sexual dimorphism, these two taxa might be conspecific (see “ Remarks ” by Moravec 2007: 285 and 287). Nonetheless, both are immediately distinguished from P. (P.) andasibense sp. nov. by their much smaller size (length of the former 10.4 mm, 9.75 mm of the latter), much longer and oblique-arcuate temples (only 1.6 times shorter than eyes) and by their labrum possessing only five anterior setae. Furthermore, the aedeagus of the holotype of P. (P.) differens is clearly distinguished from that in the new species particularly by its gradually arcuate-bent basal portion (see the line drawing by Cassola & Andriamampianina (2001, fig. 3) and the photograph by Moravec (2007, fig. 1011). It should be noted here that P. (P.) brullei was probably confused with several other species by Horn (1934 a, 1934 b) – see “ Remarks ” by Moravec (2007: 285). Pogonostoma (P.) praetervisum Moravec, 2005 (of the Pogonostoma (s. str.) alluaudi species-complex within this species-group) also may resemble the new species by shape of its temples, yet the temples are extremely short, 3.2 – 4.2 times shorter than eyes. Furthermore, apart from its much smaller body, P. (P.) praetervisum clearly differs from the new species in having coarser pronotal surface, much larger elytral punctures which are commonly anastomosing in irregular chains, and sexually dimorphic labrum with only 4 – 5 anterior setae. Also, its arcuateboomerang-bent aedeagus is much shorter, and with much narrower, dorsally moderately bent apex (for relevant characters see figs 911 – 918 and 1738 by Moravec 2007). Pogonostoma (Pogonostoma) externespinosum W. Horn, 1927 also has its aedeagus distinctly boomerang-bent (see fig. 795 of the lectotype (MNHN) and of another male in fig. 803 by Moravec 2007). This species (of the P. (s. str.) srnkai species-group), described by Horn (1927) from Montagne d’Ambre, is known only from the type locality and from Montagne des Français (both situated near Antsiranana (= Diego Suarez), Northern Madagascar). However, apart from its smaller size (10.6 – 12.1 mm), P. (P.) externespinosum differs from P. (P.) andasibense sp. nov. in having its head temples arcuate (in contrast to the abruptly bulging temples in the new species) and pronotal disc narrower and ellipsoidal, covered with finer rugae (see Moravec 2007, figs 792 and 1725), its elytra have deeper discal impression and finer punctures which are more spaced (partly almost effaced) on postero-median area of elytral disc including anteapical convexity, and elytral apices (in both sexes) possess more spiny-like and generally distinctly externally directed teeth (see Moravec 2007, fig. 793 of the female lectotype and figs 800 – 802 of other specimens); furthermore, the labrum of P. (P.) externespinosum is notably sexually dimorphic and with only 5 – 6 (– 7) anterior setae (see Moravec 2007, figs 797 – 799). Pogonostoma (Pogonostoma) spinipenne (Laporte de Castelnau & Gory, 1835) also has its aedeagus similar to the new species (see Moravec 2007, figs 639 – 641), as well as somewhat similar elytral apices, yet the apices have more spiny-like and externally directed teeth (see Moravec 2007, figs 647 – 649). However, apart from its larger size (14.5 – 17.0 mm) and longer, arcuate temples (1.6 times shorter than eyes), this species (of the P. (s. str.) chalybaeum species-group) is immediately distinguished from P. (P.) andasibense sp. nov. by its 11 – 14 - setose labrum (with 9 – 12 anterior setae) possessing markedly sinuous anterior margin in both sexes, shallowly but notably emarginated in middle (see Moravec 2007, figs 643 – 646); extremely rarely the labrum may have anomalously only 8 anterior setae, but the characteristic shape of the labrum clearly distinguishes all specimens of P. (P.) spinipenne from the new species. Moreover, the pronotal disc of P. (P.) spinipenne has almost parallel lateral margins (see figs 637 and 1713 of the lectotype (MNHN) by Moravec 2007).	en	Moravec, Jiří, Wiesner, Jürgen (2026): New or rare Madagascar tiger beetles- 27. Pogonostoma (Pogonostoma) andasibense sp. nov., and a revised key to the Pogonostoma (P.) elegans species-group (Coleoptera: Cicindelidae). Zootaxa 5748 (2): 215-229, DOI: 10.11646/zootaxa.5748.2.3, URL: https://doi.org/10.11646/zootaxa.5748.2.3
03DF87FEFFFFF13AFF3157BAFAD48529.taxon	description	Description. Body (Figs 1 – 2) medium-sized (somewhat larger within the species-group), length 13.7 – 15.0 mm, width 3.30 – 3.50 mm (male HT, female AT), black with faint blue lustre; setal vesture primarily whitish to blonde. Head (Figs 1 – 2, 5 – 6) large, width 2.85 – 3.00 mm, wider than pronotum and slightly narrower than body; temples rather short and abruptly bulging (notably dorsad), 0.60 – 0.70 mm long (male, female), approximately two times shorter than eyes which are 1.20 – 1.30 mm long. Frons notably wide, indistinctly separated from clypeus and not differentiated from vertex, nearly flat, with two elongate anterolateral impressions forming elongate central-anterior hump, and with distinct, rather wide posterior impression; supraantennal keels consisting of elevated, sharp anterior crest and indistinct, blunter posterior one; frons-vertex surface irregularly scabrous-rugulose, posterolateral areas almost vermicular-rugulose, occipital area transverse-wavy to vermicular rugose; dorsal surface of head covered with whitish, decumbent or erect, short or longer hairlike setae, which are much better visible in lateral view as a dense fur, while in dorsal view appearing dense and forming occasional clusters only on anterior vertexal area. Genae irregularly subparallel yet scabriculous-rugulose, with scattered long, semierect hairlike setae. Clypeus slightly elevated in middle, coriaceous-wrinkled, covered with dense, irregular whitish setae. Labrum (Figs 8 – 9) with 8 anterior and 2 lateral brownish setae, metallic black, matt-shiny with faint blue-green lustre uniformly shaped in both sexes; surface finely coriaceous, and glabrous, with large yet moderate central convexity; male labrum (Fig. 8) short (yet not transverse), length 0.75 mm, width 1.45 mm, lateral margins moderately arcuate towards shallow lateral indentations, then notably towards anterolateral teeth which are in the same level with truncate, irregularly dentate anterior margin; female labrum (Fig. 9) practically of the same shape, yet larger and only slightly longer, length 1.00 mm, width 1.65 mm, Maxillae. Galea metallic black, of normal slender shape; lacinia (Fig. 10) metallic black with faint blue-green lustre and black-brown or brownish setae, apical portion moderately spatulate-dilated (width 0.35 mm). Palpi of usual length and shape; maxillary palpi (Figs 1 – 2, 7) with longest and penultimate palpomeres black with usual white microtrichia and long black-brown setae; labial palpi metallic-black with faint green-blue lustre, terminal palpomeres with brownish inner margin or also apical area, longest palpomere covered with long, black-brown or blackish-violet setae. Mandibles (Fig. 12) normally shaped, subsymmetrical, metallic black with indistinct blue-green lustre, except for dark reddish-brown apical areas of teeth; left mandible in male with the second tooth smaller than third one, while the inner teeth are almost of the same length in right mandible. Antennae (Figs 1 – 2) very long, only slightly shorter in female, but surpassing body in both sexes, entirely black, antennomeres 1 – 4 with metallic blue lustre; scape with one or two long apical setae, its surface covered with several microsetae. Thorax. Pronotum (Figs 13 – 14) elongate, length 3.50 – 3.70 mm, width 2.25 – 2.40 mm, anterior lobe very slightly narrower than posterior one, its surface distinctly vermicular to wavy rugose; disc robust, dorsally convex, almost subglobose, nearly as long as wide, lateral margins convex, in male indistinctly subparallel in middle; notopleural sutures invisible in dorsal view; median line thin yet developed on whole discal length; discal surface rather densely wavy rugulose; wavy rugae transverse or obliquely directed, occasionally anastomosing, becoming fragmented into elongate tubercles on anterior area; large lateral areas rather distinctly irregularly tuberculate; pronotal median surface appearing glabrous, yet with sparse, easily abraded microsetae, lateral areas with scattered, average-long, decumbent or erect setae; posterior lobe irregularly wrinkled, lateral areas almost smooth, but with sparse setigerous microtubercles bearing whitish setae; prosternum, mesosternum and metasternum black, nearly smooth, only shallowly wrinkled, covered with dense, rather long, erect whitish setae; proepisterna broad, rather distinctly irregularly sculptured on large juxta notopleural area, with few whitish setae, particularly on their ventral area; mesepisterna smooth metallic black, glabrous except for few setae on anterior and ventral area, lacking any remarkable sulcus or pit; metepisterna smooth, with occasional whitish microtrichia. Elytra (Figs 17 – 18) elongate, 8.00 – 8.50 mm long, surface convex, discal impression rather shallow but distinct, basodiscal convexity moderate, posterior discal area rather distinctly convex; humeri arcuate; lateral margins subparallel, but with small arcuate protrusion above the middle in male (Fig. 17); angles of anteapical convexity arcuate; elytral apices sexually dimorphic: apex in male (Figs 15, 17) with shallow but elongate sutural emargination towards indistinct, blunt sutural spine; inner tooth large, almost right-angled and acute; external tooth smaller but acute downward directed; apex in female (Figs 16, 18) with larger sutural emargination towards blunt sutural spine, inner tooth large but blunt, notably inward directed, external tooth rather large and blunt; surface punctate on whole elytral length, including elytral base; punctures comparatively large, predominantly isolated and with flat, shiny intervals, yet those on elytral discal median area occasionally anastomosing in chains with declined intervals, but never cavernous (in terms by Moravec 2007), smaller and with large smooth intervals on anteapical convexity; elytral setal vesture consisting of nearly erect, copious, moderately long, whitish or dirtily blonde or darkened ornamental setae; few long, erect and feeble setae are present on basal area. Abdomen. Ventrites black, with scattered whitish setae and longer sensory setae at either side of their lateral margins. Legs. All segments entirely metallic-black, setal vesture whitish, yet some setae may appear darkened depending on angles of illumination; femora with sparse whitish setae which are even sparser on metafemora; tibiae, covered with short white setae which are on their dorsal surface (particularly on meso- and metatibiae) notably dense; hooks dark reddish-brown. Aedeagus (Fig. 11) rather long, length 3.20 mm (measured in the same position when photographed), width 0.40 mm, distinctly boomerang-shaped due to its basal portion abruptly ventrad-bent from the middle of the median portion; apical half rather wide, conically attenuated and dorsally slightly arcuate-emarginated towards rather wide apex with rounded tip.	en	Moravec, Jiří, Wiesner, Jürgen (2026): New or rare Madagascar tiger beetles- 27. Pogonostoma (Pogonostoma) andasibense sp. nov., and a revised key to the Pogonostoma (P.) elegans species-group (Coleoptera: Cicindelidae). Zootaxa 5748 (2): 215-229, DOI: 10.11646/zootaxa.5748.2.3, URL: https://doi.org/10.11646/zootaxa.5748.2.3
03DF87FEFFFFF13AFF3157BAFAD48529.taxon	etymology	Etymology. Named after the type locality, previously known as Andasibe-Mantadia National Park.	en	Moravec, Jiří, Wiesner, Jürgen (2026): New or rare Madagascar tiger beetles- 27. Pogonostoma (Pogonostoma) andasibense sp. nov., and a revised key to the Pogonostoma (P.) elegans species-group (Coleoptera: Cicindelidae). Zootaxa 5748 (2): 215-229, DOI: 10.11646/zootaxa.5748.2.3, URL: https://doi.org/10.11646/zootaxa.5748.2.3
03DF87FEFFFFF13AFF3157BAFAD48529.taxon	distribution	Distribution. The two type specimens of Pogonostoma (P.) andasibense sp. nov. were caught over the course of many years in the area that is today known as the Analamazaotra National Park, following a change in the type of protected area status in 2015 (see Goodman et al. 2018 and Goodman 2022), which holds evergreen rain forest (Forêt d’Analamazaotra). In the past, it was known or referred to as Andasibe-Mantadia National Park or Périnet Special Reserve, 30 km east of Moramanga. Furthermore, several differences and mistaken spellings occur in the literature and on-line resources. It must also be noted here that this portion of eastern Madagascar was previously administrated as the Province de Toamasina and in 2009 the national administration hierarchy was changed from provinces to regions, with the area now falling within the Alaotra-Mangoro Region. Many places throughout Madagascar have been renamed recently, and the system of administrative division presented by Viette (1991) has changed in the past years as well (see also Moravec 2022). Unfortunately, natural habitats in many localities have been rapidly disappearing over the past few years. Some of the unprotected areas near Moramanga, once full of tiger beetle species including those described as new for science, were totally destroyed by fire between 2001 and 2006 (see Moravec 2010, 2022, Moravec & VybÍral 2018); M. Trýzna and J. VybÍral pers. com. Severe fires also destroyed other parts of strictly preserved areas throughout Madagascar quite recently (e. g. Vyawahare 2020 and Andrianambinina et al. 2025). Fortunately, the Analamazaotra National Park remains one of the best protected areas. For the protected areas (map Fig. 20), phytogeography, ecosystems and biotopes see Goodman et al. (2018) and Goodman (2022). It is noteworthy that according to the coordinates written by the collector on the locality label, the male holotype of the new species was caught outside of the national park, somewhat southward from the NR 2, as obvious from the map Fig. 19. Nevertheless, as the national park is written on the locality label, we may speculate that the coordinates might have been written erroneously. We were unable to receive exact data on the habitats of the new species from the two collectors, yet the adult specimens were almost certainly caught on trunks of trees in the evergreen Analamazaotra forest. Adults and larvae of all species of the genus are strictly silvicolous and arboreal, confined to trees of all types of forest (see Emden (1935), Vadon (1935), Jeannel (1946 a, b), Rivalier (1970) Putchkov & Dolin (2005) and Moravec 2007).	en	Moravec, Jiří, Wiesner, Jürgen (2026): New or rare Madagascar tiger beetles- 27. Pogonostoma (Pogonostoma) andasibense sp. nov., and a revised key to the Pogonostoma (P.) elegans species-group (Coleoptera: Cicindelidae). Zootaxa 5748 (2): 215-229, DOI: 10.11646/zootaxa.5748.2.3, URL: https://doi.org/10.11646/zootaxa.5748.2.3
