taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03DF87ECFFDF3670FF38860179B08EFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891248/files/figure.png	https://doi.org/10.5281/zenodo.17891248	FIGURE 4. Body and sole colouration of L. macedonicus macedonicus (A, B), Jakupica Mountains, North Macedonia and L. graecus (C), Shar Mountains, North Macedonia. Sole colouration of L. m. leucopus (D). The deep dark colouration of the body and distinctive dark lateral sole fields (A) of L. macedonicus macedonicus, as well as fingerprint pattern on the mantle (B) are well visible in contrast to L. graecus (C).	FIGURE 4. Body and sole colouration of L. macedonicus macedonicus (A, B), Jakupica Mountains, North Macedonia and L. graecus (C), Shar Mountains, North Macedonia. Sole colouration of L. m. leucopus (D). The deep dark colouration of the body and distinctive dark lateral sole fields (A) of L. macedonicus macedonicus, as well as fingerprint pattern on the mantle (B) are well visible in contrast to L. graecus (C).	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFDF3670FF38860179B08EFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891250/files/figure.png	https://doi.org/10.5281/zenodo.17891250	FIGURE 5. Vestigial shell of L. macedonicus macedonicus (A), and L. macedonicus leucopus (B).	FIGURE 5. Vestigial shell of L. macedonicus macedonicus (A), and L. macedonicus leucopus (B).	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFDF3670FF38860179B08EFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891254/files/figure.png	https://doi.org/10.5281/zenodo.17891254	FIGURE 7. Comparative pictures of the reproductive system of: A. L. macedonicus leucopus, North Macedonia Pelister Mts., B. L. macedonicus leucopus, North Macedonia Pelister Mts., female parts are still not developed; C. L. macedonicus macedonicus, North Macedonia, Jakupica Mts., D. L. graecus, after Wiktor (2001), Zoomed male parts of the reproductive systems: E. L. macedonicus leucopus, North Macedonia Pelister Mts., F. L. macedonicus macedonicus, North Macedonia, Jakupica Mts., G. L. graecus, SW Bulgaria, Kresna gorge.	FIGURE 7. Comparative pictures of the reproductive system of: A. L. macedonicus leucopus, North Macedonia Pelister Mts., B. L. macedonicus leucopus, North Macedonia Pelister Mts., female parts are still not developed; C. L. macedonicus macedonicus, North Macedonia, Jakupica Mts., D. L. graecus, after Wiktor (2001), Zoomed male parts of the reproductive systems: E. L. macedonicus leucopus, North Macedonia Pelister Mts., F. L. macedonicus macedonicus, North Macedonia, Jakupica Mts., G. L. graecus, SW Bulgaria, Kresna gorge.	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFDF3670FF38860179B08EFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891244/files/figure.png	https://doi.org/10.5281/zenodo.17891244	FIGURE 3. A maximum likelihood tree based on a COI fragment (573 bp).	FIGURE 3. A maximum likelihood tree based on a COI fragment (573 bp).	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFDF3670FF38860179B08EFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891242/files/figure.png	https://doi.org/10.5281/zenodo.17891242	FIGURE 2. Habitats: A–C L. macedonicus macedonicus, A—Jakupica Mts., the first site (red spot) detected in 2011 and general habitat of L. macedonicus, with Solunska Glava peak in the background, where the first topotype series was collected; Rep. Macedonia, Jakupica Mountains, site above Begovo Pole (to the left) just below Solunska Glava peak. Well visible the stony area of a sinkhole below the road, just behind the Pinus mugo bush; B—Jakupica Mts., the second area, detected in 2017, Marina Rupa, area of sinkholes, between the peaks of Cheples and Kantarica. Well visible the big rocky area of two sink holes. In the background the southern extending ridge of Solunska Glava with Kantarica peak; C—Jakupica Mts., the third site detected in 2017; Marina Rupa, area of sinkholes, between the peaks of Ceples and Kantarica; D—Pelister Mts., view of the area were L. macedonicus leucopus was found.	FIGURE 2. Habitats: A–C L. macedonicus macedonicus, A—Jakupica Mts., the first site (red spot) detected in 2011 and general habitat of L. macedonicus, with Solunska Glava peak in the background, where the first topotype series was collected; Rep. Macedonia, Jakupica Mountains, site above Begovo Pole (to the left) just below Solunska Glava peak. Well visible the stony area of a sinkhole below the road, just behind the Pinus mugo bush; B—Jakupica Mts., the second area, detected in 2017, Marina Rupa, area of sinkholes, between the peaks of Cheples and Kantarica. Well visible the big rocky area of two sink holes. In the background the southern extending ridge of Solunska Glava with Kantarica peak; C—Jakupica Mts., the third site detected in 2017; Marina Rupa, area of sinkholes, between the peaks of Ceples and Kantarica; D—Pelister Mts., view of the area were L. macedonicus leucopus was found.	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFDF3670FF38860179B08EFC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891252/files/figure.png	https://doi.org/10.5281/zenodo.17891252	FIGURE 6. Reproductive system of L. graecus, SW Bulgaria, Kresna gorge. AT—atrium, HG—hermaphrodite gland, HD— hermaphrodite duct, OD—oviduct; SD—sperm-oviduct; PA—penial appendage, P—penis, PR—penis retractor muscle, VD— vas deferens; AG—albumen gland, FO—free oviduct, BC—bursa copulatrix, BCD—bursa copulatrix duct.	FIGURE 6. Reproductive system of L. graecus, SW Bulgaria, Kresna gorge. AT—atrium, HG—hermaphrodite gland, HD— hermaphrodite duct, OD—oviduct; SD—sperm-oviduct; PA—penial appendage, P—penis, PR—penis retractor muscle, VD— vas deferens; AG—albumen gland, FO—free oviduct, BC—bursa copulatrix, BCD—bursa copulatrix duct.	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFD03676FF38841A786C8D59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891248/files/figure.png	https://doi.org/10.5281/zenodo.17891248	FIGURE 4. Body and sole colouration of L. macedonicus macedonicus (A, B), Jakupica Mountains, North Macedonia and L. graecus (C), Shar Mountains, North Macedonia. Sole colouration of L. m. leucopus (D). The deep dark colouration of the body and distinctive dark lateral sole fields (A) of L. macedonicus macedonicus, as well as fingerprint pattern on the mantle (B) are well visible in contrast to L. graecus (C).	FIGURE 4. Body and sole colouration of L. macedonicus macedonicus (A, B), Jakupica Mountains, North Macedonia and L. graecus (C), Shar Mountains, North Macedonia. Sole colouration of L. m. leucopus (D). The deep dark colouration of the body and distinctive dark lateral sole fields (A) of L. macedonicus macedonicus, as well as fingerprint pattern on the mantle (B) are well visible in contrast to L. graecus (C).	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFD03676FF38841A786C8D59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891250/files/figure.png	https://doi.org/10.5281/zenodo.17891250	FIGURE 5. Vestigial shell of L. macedonicus macedonicus (A), and L. macedonicus leucopus (B).	FIGURE 5. Vestigial shell of L. macedonicus macedonicus (A), and L. macedonicus leucopus (B).	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFD03676FF38841A786C8D59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891254/files/figure.png	https://doi.org/10.5281/zenodo.17891254	FIGURE 7. Comparative pictures of the reproductive system of: A. L. macedonicus leucopus, North Macedonia Pelister Mts., B. L. macedonicus leucopus, North Macedonia Pelister Mts., female parts are still not developed; C. L. macedonicus macedonicus, North Macedonia, Jakupica Mts., D. L. graecus, after Wiktor (2001), Zoomed male parts of the reproductive systems: E. L. macedonicus leucopus, North Macedonia Pelister Mts., F. L. macedonicus macedonicus, North Macedonia, Jakupica Mts., G. L. graecus, SW Bulgaria, Kresna gorge.	FIGURE 7. Comparative pictures of the reproductive system of: A. L. macedonicus leucopus, North Macedonia Pelister Mts., B. L. macedonicus leucopus, North Macedonia Pelister Mts., female parts are still not developed; C. L. macedonicus macedonicus, North Macedonia, Jakupica Mts., D. L. graecus, after Wiktor (2001), Zoomed male parts of the reproductive systems: E. L. macedonicus leucopus, North Macedonia Pelister Mts., F. L. macedonicus macedonicus, North Macedonia, Jakupica Mts., G. L. graecus, SW Bulgaria, Kresna gorge.	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFD03676FF38841A786C8D59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891256/files/figure.png	https://doi.org/10.5281/zenodo.17891256	FIGURE 8. Copulation of A., B. L. macedonicus leucopus, North Macedonia Pelister Mts., C. L. conemenosi.	FIGURE 8. Copulation of A., B. L. macedonicus leucopus, North Macedonia Pelister Mts., C. L. conemenosi.	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFD03676FF38841A786C8D59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891242/files/figure.png	https://doi.org/10.5281/zenodo.17891242	FIGURE 2. Habitats: A–C L. macedonicus macedonicus, A—Jakupica Mts., the first site (red spot) detected in 2011 and general habitat of L. macedonicus, with Solunska Glava peak in the background, where the first topotype series was collected; Rep. Macedonia, Jakupica Mountains, site above Begovo Pole (to the left) just below Solunska Glava peak. Well visible the stony area of a sinkhole below the road, just behind the Pinus mugo bush; B—Jakupica Mts., the second area, detected in 2017, Marina Rupa, area of sinkholes, between the peaks of Cheples and Kantarica. Well visible the big rocky area of two sink holes. In the background the southern extending ridge of Solunska Glava with Kantarica peak; C—Jakupica Mts., the third site detected in 2017; Marina Rupa, area of sinkholes, between the peaks of Ceples and Kantarica; D—Pelister Mts., view of the area were L. macedonicus leucopus was found.	FIGURE 2. Habitats: A–C L. macedonicus macedonicus, A—Jakupica Mts., the first site (red spot) detected in 2011 and general habitat of L. macedonicus, with Solunska Glava peak in the background, where the first topotype series was collected; Rep. Macedonia, Jakupica Mountains, site above Begovo Pole (to the left) just below Solunska Glava peak. Well visible the stony area of a sinkhole below the road, just behind the Pinus mugo bush; B—Jakupica Mts., the second area, detected in 2017, Marina Rupa, area of sinkholes, between the peaks of Cheples and Kantarica. Well visible the big rocky area of two sink holes. In the background the southern extending ridge of Solunska Glava with Kantarica peak; C—Jakupica Mts., the third site detected in 2017; Marina Rupa, area of sinkholes, between the peaks of Ceples and Kantarica; D—Pelister Mts., view of the area were L. macedonicus leucopus was found.	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
03DF87ECFFD03676FF38841A786C8D59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17891244/files/figure.png	https://doi.org/10.5281/zenodo.17891244	FIGURE 3. A maximum likelihood tree based on a COI fragment (573 bp).	FIGURE 3. A maximum likelihood tree based on a COI fragment (573 bp).	2025-11-11	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon		Zenodo	biologists	Dedov, Ivaylo K.;Schneppat, Ulrich E.;Glogger, Fabia Knechtle;Cornu, Regula;Hristovski, Slavčo;Borissov, Simeon			
