identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D0F70EFFBFFFAEFF55FC47FA1DF95D.text	03D0F70EFFBFFFAEFF55FC47FA1DF95D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stylogeocoris Montandon 1913	<div><p>Genus Stylogeocoris Montandon, 1913</p><p>Stylogeocoris Montandon, 1913: 55 . Type species: Stylogeocoris biroi Montandon, 1913, by monotypy.</p><p>Diagnosis. See Malipatil (1994: 316–317).</p><p>Key to species of Stylogeocoris</p><p>[Modified from Malipatil (1994: 317)]</p><p>1. Dorsum of entire head orange or pale yellow............................................................ 2</p><p>- Dorsum of at least basal 3/4 of head excluding postero-lateral areas of head behind eyes fuscous or black............ 3</p><p>2. Ocular sulcus on dorsum of head each with one fuscous red spot at about 1/2 length of head; (short, broad species)............................................................................... S. maculatus Malipatil, 1994</p><p>- Ocular sulcus on dorsum of head without spots but with fuscous areas..................... S. biroi Montandon, 1913</p><p>3. Abdominal venter bicolorous, sterna III to anterior half of V dirty yellow, posterior to this fuscous; corial inner-most row of punctures along claval furrow as long as apical 1/2 of clavus (Figs. 9, 11).......... S. bicoloratus Malipatil, sp. nov.</p><p>- Abdominal venter almost similarly coloured throughout, not bicolorous; corial inner-most row of punctures represented only by a few punctures, often irregularly placed, and occupying at most apical 1/3 of clavus (Fig. 1)............... 4</p><p>4. Lateral margin of pronotum either partly or entirely pale in contrast to fuscous rest of pronotum (Figs. 1, 2)..................................................................................... S. provisus (Bergroth, 1895)</p><p>- Lateral margin of pronotum fuscous as rest of pronotum................................................... 5</p><p>5. Hind margin of pronotum with a narrow uniform pale-yellow band...................... S. convivus (Distant, 1901)</p><p>- Hind margin of pronotum fuscous as rest of pronotum........................... S. capricornutus (Kirkaldy, 1908)</p></div>	https://treatment.plazi.org/id/03D0F70EFFBFFFAEFF55FC47FA1DF95D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Malipatil, M. B.;Liu, Yingqi	Malipatil, M. B., Liu, Yingqi (2026): Additions to the knowledge of Stylogeocoris and Umbrageocoris of Australia (Hemiptera: Heteroptera: Geocoridae). Zootaxa 5768 (3): 370-380, DOI: 10.11646/zootaxa.5768.3.3, URL: https://doi.org/10.11646/zootaxa.5768.3.3
03D0F70EFFBFFFABFF55F88AFCC1FABF.text	03D0F70EFFBFFFABFF55F88AFCC1FABF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stylogeocoris provisus (Bergroth 1895) Malipatil & Liu 2026	<div><p>Stylogeocoris provisus (Bergroth, 1895)</p><p>(Figs. 1–7)</p><p>Geocoris provisus Bergroth, 1895: 294 .</p><p>Geocoris elongatus Distant, 1901: 470 . Syn. nov.</p><p>Geocoris provisus: Slater (1964: 580), Malipatil (1994: 301).</p><p>Geocoris elongatus: Slater (1964: 548).</p><p>Stylogeocoris elongatus: Malipatil (1994: 317, 321).</p><p>This species was redescribed in detail by Malipatil (1994) as Stylogeocoris elongatus . The redescription is updated here based on examination of a syntype and new information about the morphology and distribution that has become available from the additional specimens listed below.</p><p>Type specimens examined: Syntype of Geocoris provisus: female (Figs. 1, 2), with labels (Fig. 3): Geocoris provisus Bergr. VICTORIA / Geocoris provisus Bergr. (in Bergroth’s handwriting) / Nat. Mus. Victoria / 122. / [circular green label]; antennae, most of legs, distal 3 segments of labium missing. Deposited in MV.</p><p>Lectotype of Geocoris elongatus (Malipatil 1994): female, Gayndah, 257 / Distant coll., 1911-383 / syntype / elongatus Dist.; glued on triangle; right antenna missing. Paralectotype female, same data as lectotype; glued on triangle, pinned below lectotype on the same pin. Both deposited in NHM.</p><p>Other specimens examined: AUSTRALIA. Queensland: 1 female, Cairns, vii.1940, ‘predatory on the lantana bug’ (QDPI) ; 1 male, Mt Spec, 22.ii.1994, L.J. Cookson (MV) ; 1 female (dissected), 40 km S of Roma, 10.i.1984, K.L. Walker, on Eucalyptus sp. (MV) ; 1 male, Mt Edwards, iii.1934, A.R.B. (QDPI) ; 1 female, Stanthorpe, 8.xi.1926 (QDPI) ; 1 female, Maryvale, vi.1926 (QDPI) ; 1 female, Emerald, 17.ix.1975, G.K. Waite, ex. linseed (QDPI) ; 1 female, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=152.57&amp;materialsCitation.latitude=-26.51" title="Search Plazi for locations around (long 152.57/lat -26.51)">Beerwah</a>, 26.51S 152.57E, 29.x.-23.xi. 1986, B.K. Cantrell, Malaise trap (QDPI) ; 1 male, MHRS [ Maroochy Horticulture Research Station], 26. x.1984, G. K. Waite, ex peaches (QDPI) ; 1 female, Ashgrove, vii.1932, A. Brimblecombe (QDPI) ; 1 female, Brisbane, 26.i.1977, J. Cantrell (QDPI) ; 1 male (dissected), Indooroopilly, 28.i- 7.ii.1983, Malaise trap (QDPI) , 1 female, same data except, 6–12.xii.1984 (QDPI) . Specimens from additional localities from Queensland and New South Wales were listed by Malipatil (1994: 321–322) as Stylogeocoris elongatus .</p><p>Additionally, as of 7 March 2025, the following observation records were available on Atlas of Living Australia via ‘ iNaturalist Australia ’ and ‘ NatureMapr ’: 21 from Queensland, 12 from New South Wales, 10 from Australian Capital Territory, and 1 from Victoria (Cranbourne). No specimens from these observations were available for examination.</p><p>Diagnosis. Head with most of posterolateral area behind eyes dorsally and ventrally yellow. Pronotal lateral margins for most length (at least posterior 1/2–3/4) yellow. Posterior 1/2–3/4 of pronotum covered with large dense punctures not arranged in distinct rows (Figs. 1, 2).</p><p>Redescription</p><p>Colour generally black; head ventrally and entire anterior margin, most of posterolateral area of head behind eyes dorsally and ventrally, pronotal lateral margins for most length (at least posterior 1/2–3/4), and middle of scutellum not reaching base yellow; antennae brown, base of second and third segments often dark brown, distal 5/6 of fourth segment dark brown; entire or almost entire length of corial outer margins and connexiva broadly yellow or yellowish brown, intersegmental sutures of connexiva often fuscous; membrane dark brown, inner basal area slightly dirty white. Legs and labium generally yellow, distal one or two labial segments fuscous.</p><p>Body elongate ovate in outline (Fig. 1). Total length 5.00 (4.06–4.39); maximum width 2.10 (1.54–1.70).</p><p>Head. Dorsally between eyes almost flat, rugose and indistinctly punctate, ocular sulcus reaching to apex of head, head ventrally slightly depressed in apical half between antennifers; bucculae short, extending caudad as low ridges forming a V and continuing in a line, ending well before base of head; head length 0.80 (0.74–0.76); width across eyes 2.00 (1.86–1.97); interocular space 1.11 (0.91–1.10); interocellar space 0.72 (0.63–0.64); eye-ocellar space 0.17 (0.11–0.16); eye length 0.68 (0.75–0.76). Antennae relatively short, segment I slightly thicker than II and III, IV rather heavy, narrowed at ends, covered with fine golden pubescence; length of segments I missing (0.22–0.26); II missing (0.50–0.53); III missing (0.38); IV missing (0.52–0.56). Labium reaching to middle coxae, segment I ending well before base of head, length of segments I 0.55 (0.49–0.54); II missing (0.49–0.50); III missing (0.41–0.47); IV missing (0.39–0.43).</p><p>Thorax. Pronotum trapezoidal, covered with coarse dense and irregular punctures all over except for callosities and posterior margin; anterior collar distinct; lateral margins narrowly carinate, posterior margin almost straight; length 1.25 (1.10–1.14); maximum width 1.90 (1.67–1.79). Scutellum equilateral, margins coarsely punctate, basally slightly impressed in middle, median trifurcate carina distinct; length 1.02 (0.79–0.83); width 1.11 (0.93–1.03). Clavus with a complete row of punctures along corial margin, in addition a few punctures near base and apex of inner margin. Hemelytra fully covering abdomen, partly exposing connexiva; corium with punctures as follows: a row of punctures close to claval margin, extending onto posterior margin, inner to this row a few often irregularly placed punctures representing the inner most row only in apical 1/3 of clavus, and a row along outer lateral margin but gradually turning inwards about midlength, in addition several irregular punctures on disc in apical 1/3–1/2; membrane with distinct veins; length hemelytron 3.77 (2.80–2.95); length corium 2.90 (2.03–2.10).</p><p>Abdomen. Female with terga III–V with numerous coarse punctures, V with lesser number; VI and VII almost lacking any punctures (Fig. 4). Male as in female except VI with only a few punctures, VII with dense minute punctures.</p><p>Female genitalia. Spermatheca as in Fig. 5, bulb moderately sclerotised, with a distinct basal external flange, duct short.</p><p>Male genitalia. Pygophore posteriorly rounded. Paramere with outer lobe more developed than inner lobe, blade long, narrow and gradually curved (Fig. 6). Aedeagus with phallotheca heavily and uniformly sclerotised; helicoid process about 1.5 coiled, gonoporal process robust, about 4 coiled, coils get larger towards distal end which is gradually thickened; body and wings of ejaculatory reservoir well developed (Fig. 7).</p><p>Distribution. Known from Queensland, New South Wales, Australian Capital Territory and Victoria. Malipatil (1994) recorded this species (as S. elongatus) also from two localities in Western Australia: Broomehill and Swan View east of Midland Junction. However, upon further review, the specimen from Broomehill was identified as S. bicoloratus Malipatil, sp. nov. (see details below). The specimen from Swan View has not been located for reexamination, thus the presence of this species in Western Australia remains unverified at present.</p><p>Notes. Geocoris provisus was described based on an unspecified number of specimens (Bergroth 1895). Malipatil (1994) could not locate the type specimen, and he assumed that they were lost. Recently, an incomplete female specimen was discovered in the collections of Museum Victoria, Melbourne. Based on a comparison of the specimen with the inadequate original description, furthermore an examination of its labels, considering that the species name is written in Bergroth’s hand (see Fig. 3) as judged from comparison with handwritten labels on type specimens of other species described in the same paper (Bergroth 1895), e.g., Germalus victoriae Bergroth, 1895, it was concluded that it indeed is the specimen or one of the specimens on which Bergroth based his description of Geocoris provisus . Accordingly, it is considered a syntype.</p><p>Although the species was originally placed in genus Geocoris, it belongs to Stylogeocoris according to the current generic classification of Australian Geocoridae (Malipatil 1994), therefore the new combination Stylogeocoris provisus (Bergroth, 1895), comb. nov., is proposed.</p><p>This species is related to S. capricornutus but differs from the latter in the coloration of the head and the pronotum (the posterolateral area of the head behind the eyes dorsally and ventrally, and the lateral margin of the pronotum are fuscous or black as the rest of the pronotum in S. c apricornutus).</p><p>There is some variation in colour markings among the specimens examined:a specimen from Cairns (Queensland) has the posterolateral area of the head behind the eyes dorsally almost entirely fuscous; another specimen from Emerald (Queensland) has the abdominal ventral surface pale castaneous. A male from Mt. Spec (Queensland) is considerably smaller in size (body length 3.75 mm) and its head has coarse sparse punctures dorsally rather than being rugose and indistinctly punctate as in other specimens.</p></div>	https://treatment.plazi.org/id/03D0F70EFFBFFFABFF55F88AFCC1FABF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Malipatil, M. B.;Liu, Yingqi	Malipatil, M. B., Liu, Yingqi (2026): Additions to the knowledge of Stylogeocoris and Umbrageocoris of Australia (Hemiptera: Heteroptera: Geocoridae). Zootaxa 5768 (3): 370-380, DOI: 10.11646/zootaxa.5768.3.3, URL: https://doi.org/10.11646/zootaxa.5768.3.3
03D0F70EFFBAFFA9FF55FAECFA9FFAE1.text	03D0F70EFFBAFFA9FF55FAECFA9FFAE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stylogeocoris bicoloratus Malipatil 2026	<div><p>Stylogeocoris bicoloratus Malipatil, sp. nov.</p><p>(Figs. 8–11, 14–16)</p><p>Type specimens: Holotype female, Lake Hattah Victoria J.E. Dixon / J. E. Dixon Collection DON Jan 1940 / HEM-4808 /; deposited in MV . Paratypes, 2 males and 3 females, same label data as holotype (MV); 2 males, same label data as holotype (SAM); 1 female, Western Australia, Norseman, 8.iii.2015, L.J. Cookson (MV) .</p><p>Diagnosis. (Figs. 8–11). Head with posterolateral narrow area behind eyes dorsally and ventrally almost uniformly yellow. Pronotal lateral margins with posterolateral areas dirty yellow. Posterior 1/2–3/4 of pronotum covered with large dense punctures not arranged in distinct rows. Abdominal sterna III to anterior half of sternum V dirty yellow, posterior to this fuscous.</p><p>Description</p><p>Colour of body mostly black with markings as follows: head below and entire anterior margin extending above to 1/3–1/2 head length, posterolateral area of head behind eyes, posterolateral areas of pronotum, middle of scutellum and arms of trifurcate carina not reaching base yellow. Antennae yellowish, distal and proximal ends of segments I, II, III and about distal 1/2 of IV dark brown. Hemelytra with entire length of corial outer margins broadly yellow; connexiva yellow; corial inner corner area fuscous; membrane dark brown or fuscous, inner basal area slightly dirty white. Legs and labium generally yellow, distal two labial segments fuscous.</p><p>Body generally elongate ovate (Fig. 8). Total length 5.05 (4.48); maximum width 1.97 (1.63).</p><p>Head. Dorsally between eyes almost flat, coarsely rugose, ocular sulcus reaching to apex of head, head ventrally slightly depressed in apical half between antennifers; bucculae short, extending caudad as low ridges forming a V and continuing in a line, ending well before base of head; head length 0.68 (0.71); width across eyes 1.90 (1.78); interocular space 1.14 (1.02); interocellar space 0.66 (0.57); eye-ocellar space 0.15 (0.15); eye length 0.68 (0.68). Antennae relatively short, segment one slightly thicker than two and three, four rather heavy, narrowed at ends, covered with fine golden pubescence; length of segments I 0.34 (0.30); II 0.57 (0.53); III 0.39 (0.34); IV 0.57 (0.57). Labium reaching to middle coxae, segment one ending well before base of head, length of segments I 0.39 (0.42); II 0.49 (0.49); III 0.45 (0.45); IV 0.40 (0.32).</p><p>Thorax. Pronotum trapeziform, covered with coarse, dense and irregular punctures all over except for callosities and posterior margin; anterior collar narrow and indistinct; lateral margins narrowly carinate, posterior margin almost straight; length 1.10 (0.95); maximum width 1.71 (1.59). Scutellum subequilateral, margins coarsely punctate, basally slightly depressed in middle, median trifurcate carina bulging, medially rather explanate; length 0.90 (0.83); width 0.95 (0.87). Clavus with a complete row of punctures along corial margin, also inner margin with an incomplete row with a few punctures near base and at apex of clavus. Hemelytra fully covering abdomen, partly exposing connexiva; corium with punctures arranged as follows: a complete row parallel to claval margin extending onto posterior margin, inner to this an additional but incomplete row along corial inner margin distinct only in about apical half-length of clavus, this row also with a few irregularly placed punctures near apical area of clavus, and also a complete row along outer lateral margin of corium but gradually turning inwards about midlength, in addition several irregular and faint punctures on disc in apical 1/3-1/2; membrane with distinct veins; length hemelytron 3.53 (3.04); length corium 2.31 (2.25).</p><p>Abdomen. Tergum uniformly sclerotised. Male and female with terga III to V, particularly in middle area with coarse sparse punctures. Trichobothria on sternum III in triangle; those on IV in linear series. Male with apodeme on anterior margin of sternum VII broad, gradually narrowed and pointed apically, approximately 3/4 as long as VI.</p><p>Female genitalia. Spermatheca (Fig. 14), bulb spherical and with distinct flange near base; duct short.</p><p>Male genitalia. Pygophore posterior end gradually rounded. Paramere with lobes distinct, outer lobe much larger; blade narrow, shallowly sickle-shaped (Fig. 15). Aedeagus with phallotheca uniformly sclerotised; helicoid process 1–2 coiled; gonoporal process robust, 3–4 coiled; secondary gonopore markedly thickened (Fig. 16).</p><p>Etymology. The epithet is the Latin adjective bicoloratu s, - a, - um, in allusion to the species’ bicoloured abdominal venter, i.e., sterna III to anterior half of V dirty yellow, posterior to this fuscous.</p><p>Distribution. Known from Victoria and Western Australia.</p><p>Notes. This species can be readily distinguished from all other known species of the genus in the coloration of the abdomen with the sterna III to the anterior half of V are dirty yellow, and the segments posterior to this are uniformly fuscous (all abdominal sterna excluding connexiva uniformly fuscous in all other species).</p><p>This species is related to S. provisus but differs from the latter, besides the above noted abdominal coloration, in the presence of an additional but incomplete row of punctures along the corial inner margin, the punctures distinct only in about apical half length of the clavus, this row also has a few irregular additional punctures near the apical area of the clavus (the row of punctures on corial inner margin is represented only by a few sparse often irregularly placed punctures in apical 1/3 area of clavus in S. provisus).</p><p>There is considerable variation in the intensity of colour markings among the specimens examined.</p></div>	https://treatment.plazi.org/id/03D0F70EFFBAFFA9FF55FAECFA9FFAE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Malipatil, M. B.;Liu, Yingqi	Malipatil, M. B., Liu, Yingqi (2026): Additions to the knowledge of Stylogeocoris and Umbrageocoris of Australia (Hemiptera: Heteroptera: Geocoridae). Zootaxa 5768 (3): 370-380, DOI: 10.11646/zootaxa.5768.3.3, URL: https://doi.org/10.11646/zootaxa.5768.3.3
03D0F70EFFB8FFA8FF55FA13FC6AFB81.text	03D0F70EFFB8FFA8FF55FA13FC6AFB81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Umbrageocoris Kobor 2019	<div><p>Genus Umbrageocoris Kóbor, 2019</p><p>Umbrageocoris Kóbor, 2019: 140 . Type species: Umbrageocoris kondorosyi Kóbor, 2019, by monotypy.</p><p>Diagnosis. See Kóbor (2019).</p><p>Description and redescription. See Kóbor (2019, 2022).</p><p>Notes. This genus presently includes 5 species and 2 subspecies from the Australian and Indomalayan Regions, among them 3 Australian species: U. elegantulus (Distant, 1904), U. woodwardi (Malipatil, 1994) and U. malipatili Kóbor, 2022 (Kóbor 2022). An updated redescription of the genus, a key to included species, a differential diagnosis allowing separation of the genus from related genera ( Geocoris, Germalus Stål, 1862 and Stylogeocoris), and diagnoses for the included species were provided by Kóbor (2022).</p><p>Umbrageocoris can be readily distinguished from Stylogeocoris a nd Germalus by its labial segment II being shorter than III (segments II and III are subequal in the latter two genera (Malipatil 1994)), but not from the largest and the most widely distributed genus Geocoris, because the relative lengths of the labial segments in the latter genus are similar to the condition found in Umbrageocoris (e.g., Kóbor 2022).</p><p>Redescriptions of the genus Geocoris were provided by several workers, including Readio and Sweet (1982), Malipatil (1994) and Brailovsky (2016, 2025). In Geocoris the labial segment II is distinctly shorter than III; the eyes are semistylate, sometimes close to or in contact with the anterior angles of pronotum; and the ocelli are usually closer to the inner margin of eyes than to the medial line of vertex. Although the genus Umbrageocoris shares the above listed characters, Kóbor (2022) claimed that it can be distinguished from Geocoris distributed in the region by the combination of the following characters: (1) head pentagonal, posterior edge of eyes touching the anterior edges of pronotum (head lunulate or crescent-shaped, posterior edge of eyes encompassing anterior edges of pronotum in Geocoris); (2) pronotum is trapeziform with lateral margins slightly constricted medially (semicircular, lateral margins mostly not constricted in Geocoris); (3) trifurcate carina of scutellum is distinct, at most apically reduced (indistinct to completely reduced in Geocoris); (4) hamus of metathoracic wing is at most partly reduced, intervannals present (hamus completely reduced and intervannals absent in Geocoris); and (5) apex of suture of abdominal tergites V/VI obtuse (rounded in Geocoris).</p><p>It appears that based on the available literature as well as our own observations on specimens of the Australian species placed into Geocoris by Malipatil (1994), characters 1, 2 and 3 listed above are considerably variable amongst the specimens examined, sometimes even within the same species. In relation to character 4 (hind wing venation), Kóbor (2019b) believed his results supported the conclusions of Slater and Hurlbutt (1957) suggesting the status of hind wing venation in Geocorinae as systematic character should be examined extensively and carefully.</p><p>It should also be noted that several authors considered the genus Geocoris, the largest and the most widely distributed geocorine genus in all the major zoogeographic regions (Dellapé &amp; Henry 2025), as “an illdefined group of species belonging to perhaps several distinct genera” (e.g., Readio &amp; Sweet 1982, Malipatil 1994) which needs a thorough revision. In support of this, amongst the South American species of the genus Geocoris, three diverse morphological species groups, viz., flavilineus group, punctipes group and thoracicus group were proposed (Brailovsky 2025), and similarly amongst the 10 Mexican species, three species groups were recognised (Brailovsky 2016). For reasons explained above, the authors of the present study consider a thorough evaluation of morphological characters (including external and internal genitalia) along with molecular studies involving representative species from all major zoogeographic regions are needed for clearly defining the genus Geocoris (defined in the strict sense) and all other related genera. A detailed phylogenetic analysis of relationships among these genera would also be valuable towards a better understanding of the entire subfamily Geocorinae .</p></div>	https://treatment.plazi.org/id/03D0F70EFFB8FFA8FF55FA13FC6AFB81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Malipatil, M. B.;Liu, Yingqi	Malipatil, M. B., Liu, Yingqi (2026): Additions to the knowledge of Stylogeocoris and Umbrageocoris of Australia (Hemiptera: Heteroptera: Geocoridae). Zootaxa 5768 (3): 370-380, DOI: 10.11646/zootaxa.5768.3.3, URL: https://doi.org/10.11646/zootaxa.5768.3.3
03D0F70EFFB9FFA6FF55FB37FBFCF9E5.text	03D0F70EFFB9FFA6FF55FB37FBFCF9E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Umbrageocoris malipatili Kobor 2022	<div><p>Umbrageocoris malipatili Kóbor, 2022</p><p>(Figs. 12, 13, 17–19)</p><p>Umbrageocoris malipatili Kóbor, 2022: 142, 144, 145.</p><p>Geocoris provisus (non Bergroth, 1895): Malipatil (1994: 310). Misidentification.</p><p>The species that was misidentified and redescribed by Malipatil (1994: 310–314) as Geocoris provisus is recognized in the present study as Umbrageocoris malipatili Kóbor, 2022 . The latter species was described by Kóbor (2022) based on an incomplete female specimen (the holotype) collected from an unknown locality in New South Wales, Australia. Accordingly, the original description lacked details of several important diagnostic characters, among them the female genitalia. For this reason, an updated redescription and distribution records are provided here based on examination of specimens of both sexes from several localities in Queensland and New South Wales.</p><p>Specimens examined: AUSTRALIA. Queensland: 1 female, Pittsworth, Q., 21.viii.79, R.H. Broadley T4655 (QDPI); 1 male, Malanda Falls nr Malanda, 6.ii.1975, B.K Cantrell (QDPI) ; 1 male, Eungella, W. Mackay, 30. i.1975, B.K. Cantrell (QDPI) ; 1 female (dissected), Kingaroy, 24.iii.1982, D.J. Rogers, 1 male (dissected), Kingaroy, 20.ii.1987, D.J. Rogers, collected from soybean plants (QDPI); 2 males (1 dissected), Brisbane R., 4 km N. Harlin, 22.vii.1975, B. K Cantrell, D-Vac (QDPI); 1 female, Rex Range Lookout, via Julatten, 9.xi-2.xii.1981, Malaise trap (QDPI) ; 1 male, 1 female, Cairns Distr. (SAM); 1 female, Brisbane, 23.viii.1954, 1 female, 16.iii. 1965 (QM) . New South Wales: 1 female, Windsor, i.1999, D. James (MV) .</p><p>Diagnosis. See under Notes below.</p><p>Redescription</p><p>Colour generally yellowish brown; head dorsally and ventrally (except broad basal area behind eyes which is fuscous), anterior margin of pronotum and prosternum, pronotal lateral margins irregularly, lateral abdominal margins and connexiva, broad lateral margins of corium, legs and labium pale yellow, tip of labial segment IV fuscous; eyes red or fuscous red, ocelli hyaline; antennal segment IV and distal 1/2–2/3 of segment III brownish yellow; scutellum dark brown; hemelytra semi-hyaline with brownish punctation, inner basal area of membrane dirty white, median area of corium ferruginous (Fig. 12).</p><p>Body (Fig. 12). Generally shiny, lacking bristles; legs and antennae with short, yellow decumbent pubescence, in addition femora particularly fore femur, ventrally with widely spaced long bristles. Total length 4.30 (4.40); maximum width 1.72 (1.87).</p><p>Head shiny, almost impunctate, median furrow of vertex represented as a faint line extending from almost base of head to clypeus; jugae extending almost to tip of clypeus; bucculae not visible in dorsal view, extending caudad as low ridges to form Y with suture ending at about 1/2 length of head; head ventrally gradually swollen posterior to level of antennifers; length of head 0.64 (0.75); width across eyes 1.69 (1.70); interocular space 0.90 (0.80); interocellar space 0.43 (0.46); eye-ocellar space 0. 20 (0.20). Antennae slender; segment IV incrassate in middle and narrowed at ends; length of segments; I 0.26 (0.24); II 0.53 (0.60); III 0.45 (0.52); IV 0.55 (0.57). Labrum about as long as labial segment 1. Labium passing midcoxae; segment I ending well before base of head, slightly more incrassate than the following segments, basal 1/2 not distinctly narrowed; segment II shortest; length of segments: I 0.50 (0.64); II 0.36 (0.38); III 0.46 (0.53); IV 0.60 (0.60).</p><p>Thorax. Pronotum shiny; covered with coarse irregularly arranged punctures except on callosities and posterior margin area; thoracic pleura with closer punctures all over; length 0.98 (1.02); maximum width 1.60 (1.70). Scutellum with apex pointed; with coarse punctures at base and on sides; trifurcate carina feebly distinct; length 0.98 (1.02); maximum width 0.93 (0.95). Legs relatively long. Clavus narrowed towards scutellar apex; covered with a few punctures near base margining scutellum and an almost complete row of punctures along outer margin (Figs. 12, 13); commissure indistinct. Hemelytron slightly exceeding abdomen; corium covered with punctures as follows: two almost regular inner rows joining near base, separated and widened towards posterior margin, 1 row continuing along outer margin for basal 1/3, then curved inwards and ending before apex (Figs. 12, 13); exocorium slightly widened subapically; veins distinct on membrane (Fig. 12); length hemelytron 2.70 (3.04); length corium 1.90 (1.95).</p><p>Abdomen. Terga sclerotised; III–VI impunctate but coarsely rugose. Male tergum VII with several coarse punctures; median apodeme on anterior margin of sternum VII as long as sternum VI. Sternum IV with trichobothria arranged in linear series.</p><p>Female genitalia. Spermatheca with apical bulb, smaller sub-apical bulb and adjoining part of duct heavily sclerotised, remainder of duct weakly sclerotised; duct long, with 3–4 loose loops (Fig. 17).</p><p>Male genitalia. Pygophore with posterior margin rounded, not produced. Paramere with ventral and dorsal lobes distinct; blade long, narrow, gradually curved (Fig. 18). Aedeagus phallotheca with moderately sclerotised areas either side differentiated by a clear area in middle; wings of ejaculatory reservoir broad and narrowed laterally, gonoporal process slender, with 6–8 coils (Fig. 19).</p><p>Distribution. Originally recorded only from an unknown locality in New South Wales (Kóbor 2022), in the present study from Windsor (NSW) and several localities in southern, central and northern Queensland.</p><p>Notes. The species exhibits considerable colour variation amongst the specimens examined: in some the dorsum of head behind the eyes is fuscous only on the corners, but not in an entire band; in Brisbane (Queensland) specimens most of the pronotum, particularly the sides and large areas of the clavus and corium except the disc, are pale yellow. Among the other characters noted by Kóbor (2022: 148), the length of the median longitudinal furrow on dorsum of head appear to be variable even within one species, for instance one male (Kingaroy) and one female (Pittsworth) specimens have the furrow reaching as a faint line at most the middle of the vertex whereas in all other specimens (listed above) the furrow extending as a faint line to base of vertex. There is also slight variation in the wing venation, particularly the degree of development of hamus on metathoracic wing amongst the specimens examined. Hence the latter two characters are judged to be unreliable for species diagnosis.</p><p>This species can be distinguished from all known species except G. borealis that were placed into Geocoris by Malipatil (1994) by the buccula not projecting anteriorly and not visible as flaps on either side of the clypeus in dorsal view of the head. It differs from G. borealis by the overall body length more than two times as long as maximum body width, and the scutellum being narrowly pointed at the apex.</p><p>The remaining species of Umbrageocoris, including the two Australian species currently included in the genus by Kóbor (2022), U. elegantulus and U. woodwardi, have the buccula projecting anteriorly and visible as flaps on either side of the tylus in dorsal view of the head (= ‘cup-shaped labial trough’ of Kóbor (2022)). This condition is similar to that found in Cimex grylloides Linnaeus, 1761, the type species of Geocoris Fallén, 1814 by subsequent designation (Oshanin 1912). It is evident that the species currently included in the genus Umbrageocoris exhibit variation in this major diagnostic character (see Malipatil 1994). Therefore, a thorough review of this character across all species may be warranted to redefine the genera Geocoris and Umbrageocoris .</p></div>	https://treatment.plazi.org/id/03D0F70EFFB9FFA6FF55FB37FBFCF9E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Malipatil, M. B.;Liu, Yingqi	Malipatil, M. B., Liu, Yingqi (2026): Additions to the knowledge of Stylogeocoris and Umbrageocoris of Australia (Hemiptera: Heteroptera: Geocoridae). Zootaxa 5768 (3): 370-380, DOI: 10.11646/zootaxa.5768.3.3, URL: https://doi.org/10.11646/zootaxa.5768.3.3
