identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D087EFFFB3FF8BFF3BFF2F8A1D1327.text	03D087EFFFB3FF8BFF3BFF2F8A1D1327.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capila jayadeva subsp. jayadeva Moore 1866	<div><p>Capila jayadeva jayadeva Moore, 1866</p><p>(Fig. 2: A, B; Fig. 4; Fig. 8: A–C)</p><p>Ismene jayadeva Moore, [1858]: 248. nomen nudum</p><p>Capila jayadeva Moore, 1866: 785, pl. XLII, fig. 3 for ♂ holotype, fig. 4 for ♀ [misidentified as the ♂ of C. zennara Moore, 1866], type locality: Darjeeling; Nicéville 1893: 349; Swinhoe 1912: 266, pl. 755, figs. 1, 1b for ♂, 1a for ♀; Evans 1949: 87; Miyazaki et al. 2007: 3, figs. H-15 for ♂ ♀; Monastyrskii &amp; Devyatkin 2016: 71.</p><p>Specimens examined. C Vietnam: Dambri waterfalls, Bao Loc, Lam Dong Province: 1♂, 30.IV.2004, leg. local catcher ; 1♂, 1.V.2004, leg. local catcher; 1♂, 12.IV.2009, leg. N.D. Khoa; 1♂, 12.IX.2009, leg. N.D. Khoa; 1♂, 2004, leg. N.D. Khoa; 1♂, 4.V.2006, leg. Tamamitsu Saito; 1♂, 7.IV.2009, leg. Tamamitsu Saito. 2♂♂, Loc Tan, Lam Dong Province, 850 m, 8.IV.2013, 11.IV.2013, leg. Takashi Miyagawa. 1♂, Hon Ba, Khanh Hoa Province, 1500 m, 8.V.2015, leg. Takashi Hasegawa. 2♂♂, Chu Dang Ya, Pleiku, Gia Lai Province, 850–1250 m, 29.IV.2018, 8.IV.2023, leg. Takashi Miyagawa. 3♀♀, Da Ton, Bao Lam, Lam Dong Province, IV.2016, leg. Dang Van. 1♀, Dambri waterfalls, Bao Loc, Lam Dong Province, 19.IV.2009, leg. N.D. Khoa. 1♀, Chu Dang Ya, Pleiku, Gia Lai Province, 850–1250 m, 27.IV.2018, leg. Takashi Miyagawa. S Laos: 1♂, Dak Cheung, Sekong Province, 10.V.2012, leg. local catcher (all in coll. Saito) .</p><p>Note. Capila Moore, 1866 was established as a monotypic genus based on C. jayadeva as the type species. In the original description, Moore (1866) stated that ‘(in both sexes of Capila) hind tibiae with a dense tuft of very long hairs at the side’. Nicéville (1893) pointed out that the dense hair tuft is a male secondary sexual character, and he believed that Moore (1866) misidentified the male of Pisola zennara (= Capila zennara) as the female of C. jayadeva while the female of C. jayadeva as the male of P. zennara . Swinhoe (1912) followed Nicéville’s idea and provided images of both sexes of the two species. Seitz (1922, pl. 163C) also presented figures of jayadeva (misspelled as jajadeva) and zennara, but Evans (1949: 87) noted ‘Seitz as jayadeva ’ for zennara and ‘figs in Seitz are zennara ’ for jayadeva in his work, meaning that he considered the figure of jayadeva in Seitz as zennara . This is rather confusing, because judging from the characters shown in Seitz (1922, pl. 163C) the figure of ‘ jajadeva ’ undoubtedly belongs to a male of jayadeva rather than zennara .</p><p>Since the wing patterns of the male and female are quite different, it is not clear with what evidence Nicéville (1893) and Swinhoe (1912) determined the sex-pairing of the two species, and until now, no proof, such as mating observation or rearing record supporting the sex-pairing has been produced.</p><p>In the present paper, the female of jayadeva was confirmed by the COI gene sequences, and both imagoes and genitalia of both sexes of this taxon were illustrated. However, the lectotype designation and sex-pairing of zennara require further study.</p><p>Distribution: NE India (Sikkim, Assam), N Myanmar, S Laos, C Vietnam.</p></div>	https://treatment.plazi.org/id/03D087EFFFB3FF8BFF3BFF2F8A1D1327	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Xue, Guo-Xi;Inayoshi, Yutaka;Saito, Kotaro;Li, Meng;Rao, Zhi	Xue, Guo-Xi, Inayoshi, Yutaka, Saito, Kotaro, Li, Meng, Rao, Zhi (2026): Capila jayadeva Moore, 1866 is a sexually dimorphic and male polymorphic species including six subspecies (Hesperiidae, Pyrginae). Zootaxa 5748 (1): 89-100, DOI: 10.11646/zootaxa.5748.1.4, URL: https://doi.org/10.11646/zootaxa.5748.1.4
03D087EFFFB4FF8CFF3BFBB28A1F1033.text	03D087EFFFB4FF8CFF3BFBB28A1F1033.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capila jayadeva subsp. mackwoodi (Evans 1914)	<div><p>Capila jayadeva mackwoodi (Evans, 1914)</p><p>(Fig. 3: A–F; Fig. 5; Fig. 8: D–F)</p><p>Orthopaetus mackwoodi Evans, 1914: 303, type locality: Anisakan, near Maymyo [in Mandalay Region, Myanmar].</p><p>Capila hainana mackwoodi: Evans 1949: 88.</p><p>Capila hainana: Yu et al. 2022 (checklist).</p><p>Specimens examined. Thailand: 1♂, Mae Hong Son, 19.IV.1984; 1♂, Fang District, Chiang Mai Province, IV.1991; 7♂♂, Pua District, Nan Province, IV.1992; 1♂, Wiang Papao, Chiang Rai Province, 2.V.2015, leg. Y. Inayoshi [Specimen ID D46]; 1♂, Chiang Mai District, Chiang Mai Province, 27–30.IV.1974, leg. Aroon Samruadkit; 6♂♂ 2♀♀, Omkoi District, Chiang Mai Province, III–IV.1994 .</p><p>Note. Judging from the male specimens of mackwoodi examined in this study, the size of the white spot at cell end on the forewing is variable: in northern Thailand (Fang district of Chiang Mai and Pua district of Nan) the spot is always well developed, while in the areas from north-western to western Thailand, namely from Mae Hong Son and around Chiang Mai City (Chiang Mai district to Wiang Pa Pao district of Chiang Rai), through Omkoi district of Chiang Mai to Kanchanaburi, the spot is smaller, exhibiting a trend towards the unmarked peninsular ssp. arooni .</p><p>Distribution: Myanmar, W &amp; N Thailand, N &amp; C Laos, China (S Yunnan).</p></div>	https://treatment.plazi.org/id/03D087EFFFB4FF8CFF3BFBB28A1F1033	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Xue, Guo-Xi;Inayoshi, Yutaka;Saito, Kotaro;Li, Meng;Rao, Zhi	Xue, Guo-Xi, Inayoshi, Yutaka, Saito, Kotaro, Li, Meng, Rao, Zhi (2026): Capila jayadeva Moore, 1866 is a sexually dimorphic and male polymorphic species including six subspecies (Hesperiidae, Pyrginae). Zootaxa 5748 (1): 89-100, DOI: 10.11646/zootaxa.5748.1.4, URL: https://doi.org/10.11646/zootaxa.5748.1.4
03D087EFFFB4FF8CFF3BFF2F8B1615DF.text	03D087EFFFB4FF8CFF3BFF2F8B1615DF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capila jayadeva subsp. translucida (Leech 1893)	<div><p>Capila jayadeva translucida (Leech, 1893)</p><p>(Fig. 2: C, D)</p><p>Rhopalocampta translucida Leech, 1893: 642, pl. XXXIX, fig. 11 for ♂ holotype, type locality: ‘ Omeishan’ [Emei Shan, Sichuan, China].</p><p>Pisola pussa Hering, 1918: 53, pl. 4, figs. 4, 4a for ♀ holotype, type locality: ‘Yüh-Sa-Tam’ [ Yushatan, a village in Guangzhou, China].</p><p>Capila translucida: Evans 1949: 87; Chou 1994: 700; Fan &amp; Wang 2004: 154; Yuan et al. 2015: 130; Monastyrskii &amp; Devyatkin 2016: 71; Wu &amp; Hsu 2017: 1289; Xue et al. 2018: 263; Xue et al. 2024a: 386.</p><p>Capila nigrilima Chou &amp; Gu, 1994, In: Chou 1994: 772, 700, fig. 10 for ♂ holotype, type locality: Hainan.</p><p>Capila hainana hainana Crowley, 1900: 510, type locality: Hainan.</p><p>Capila hainana sinorientalis Huang &amp; Ding, 1994, In: Chou 1994: 772, 700, fig. 7 for ♀ holotype and ♀ paratype, type locality: Fujian.</p><p>Specimens examined. As listed in Xue et al. (2024a: 385).</p><p>Note. Although the females of translucida and jayadeva have similar wing patterns, it is hard to imagine the two taxa are conspecific because their males bear obviously different appearances. However, the COI gene sequences revealed that they belong to the same species, and the morphological characters of the genitalia of both sexes (Fig. 4; Fig. 8: A–C; Xue et al. 2024a: fig. 2, fig. 3) support this result. Since the two taxa are allopatric and jayadeva has the priority, translucida is treated as a subspecies of jayadeva .</p><p>Evans (1949: 87) recorded the year of publication as 1894 for translucida, and subsequent authors followed this record. However, according to the original literature, the ‘Part V. Hesperiidae . Pages 557–642’ of Leech’s work was published in December, 1893. Therefore, we herein made a correction.</p><p>Distribution: China (Sichuan, Guangxi, Hunan, Guangdong, Jiangxi, Fujian, Hainan), N Vietnam.</p></div>	https://treatment.plazi.org/id/03D087EFFFB4FF8CFF3BFF2F8B1615DF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Xue, Guo-Xi;Inayoshi, Yutaka;Saito, Kotaro;Li, Meng;Rao, Zhi	Xue, Guo-Xi, Inayoshi, Yutaka, Saito, Kotaro, Li, Meng, Rao, Zhi (2026): Capila jayadeva Moore, 1866 is a sexually dimorphic and male polymorphic species including six subspecies (Hesperiidae, Pyrginae). Zootaxa 5748 (1): 89-100, DOI: 10.11646/zootaxa.5748.1.4, URL: https://doi.org/10.11646/zootaxa.5748.1.4
03D087EFFFB4FF8DFF3BF91E8AA41113.text	03D087EFFFB4FF8DFF3BF91E8AA41113.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capila jayadeva subsp. barroni (Riley & Godfrey 1921)	<div><p>Capila jayadeva barroni (Riley &amp; Godfrey, 1921)</p><p>(Fig. 3: G, H; Fig. 6)</p><p>Orthophaetus (sic!) barroni Riley &amp; Godfrey, 1921: 187, pl. VI, figs. 10 for ♂, 11 for ♀, type locality: Hup Bon, Sriracha [in Chonburi Province, SE Thailand].</p><p>Specimens examined. Thailand: 1♂, Nakhon Nayok Province, 16.IV.2024, leg. S. Sophonviwatkul [Specimen ID E80]; 1♂, Sa Kaeo Province, 15.VI.2013 [Specimen ID D45]; 1♂, Chanthaburi Province, 25.IV.2015, leg. Koichi Yano [Specimen ID D44] .</p><p>Note. This taxon was treated as a synonym of Capila mackwoodi by Godfrey (1930: 357), and then as a synonym of C. hainana hainana by Evans (1949: 88). The former is now a subspecies of C. jayadeva, while the latter is a synonym of translucida (Xue et al. 2024a) which is a subspecies of C. jayadeva as well. The male of barroni is similar to that of mackwoodi, bearing a white spot at cell end of the forewing, but it can be distinguished from mackwoodi by the darker ground color on both sides of the wings. Therefore, barroni is treated as a subspecies of C. jayadeva herein.</p><p>Distribution: C to SE Thailand (Nakhon Nayok, Chanthaburi, Sa Kaeo) to S Cambodia.</p></div>	https://treatment.plazi.org/id/03D087EFFFB4FF8DFF3BF91E8AA41113	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Xue, Guo-Xi;Inayoshi, Yutaka;Saito, Kotaro;Li, Meng;Rao, Zhi	Xue, Guo-Xi, Inayoshi, Yutaka, Saito, Kotaro, Li, Meng, Rao, Zhi (2026): Capila jayadeva Moore, 1866 is a sexually dimorphic and male polymorphic species including six subspecies (Hesperiidae, Pyrginae). Zootaxa 5748 (1): 89-100, DOI: 10.11646/zootaxa.5748.1.4, URL: https://doi.org/10.11646/zootaxa.5748.1.4
03D087EFFFB7FF8FFF3BFAA38D5E10FF.text	03D087EFFFB7FF8FFF3BFAA38D5E10FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capila jayadeva subsp. arooni Eliot 1987	<div><p>Capila jayadeva arooni Eliot, 1987</p><p>(Fig. 3: I, J; Fig. 7; Fig. 8: G)</p><p>Capila hainana arooni Eliot, 1987: 27, type locality: Muang Chone [in Ranong Province, P. Thailand].</p><p>Specimens examined. Thailand: Muang Chone, Ranong Province: 1♂, 12.IV.1983; 1♂, 10.III.1988; 1♂, 12.III.1989; 1♀, 15.IV.1983; 1♀, 20.III.1989. 1♂, Khao Poata, Surat Thani Province, 20.III.1983 .</p><p>Note. Although this taxon was not included in the molecular analysis of the present paper, the genitalia characters of both sexes identified its conspecificity with jayadeva . As the southernmost subspecies, the unmarked wings in the male make it easily distinguishable from other subspecies.</p><p>Distribution: Peninsular Thailand (Ranong, Surat Thani).</p></div>	https://treatment.plazi.org/id/03D087EFFFB7FF8FFF3BFAA38D5E10FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Xue, Guo-Xi;Inayoshi, Yutaka;Saito, Kotaro;Li, Meng;Rao, Zhi	Xue, Guo-Xi, Inayoshi, Yutaka, Saito, Kotaro, Li, Meng, Rao, Zhi (2026): Capila jayadeva Moore, 1866 is a sexually dimorphic and male polymorphic species including six subspecies (Hesperiidae, Pyrginae). Zootaxa 5748 (1): 89-100, DOI: 10.11646/zootaxa.5748.1.4, URL: https://doi.org/10.11646/zootaxa.5748.1.4
03D087EFFFB9FF81FF3BFF2F8CF9141C.text	03D087EFFFB9FF81FF3BFF2F8CF9141C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Capila jayadeva subsp. khambounei Uehara 2024	<div><p>Capila jayadeva khambounei Uehara, 2024</p><p>Capila jayadeva khambounei Uehara, 2024: 16, fig. 6 for ♂ holotype and ♂ ♀ paratypes, type locality: Ban Thateng, Sekong, S. Laos.</p><p>Specimens examined. The type series including 5♂♂ 3♀♀, examined from photos.</p><p>Note. Among the subspecies of jayadeva, this taxon bears the most similar wing pattern to the nominate subspecies, and geographically it is the closest to the latter. One male of the examined specimens of ssp. jayadeva in the present study was collected from Sekong Province of Laos, near the type locality of khambounei. According to Uehara &amp; Masui (2024), the following characters of khambounei can be used to distinguish the taxon from the nominate subspecies: in the male, the ground color on the dorsal side of both wings is lighter, the streaks between the veins are less distinct, the distal ends of the upper and middle streaks in the cell of the forewing are expanded or even conjoined into one spot; in the female, the black stripes on the dorsal side of the hindwing are less distinct.</p><p>Distribution: Only known from the type locality.</p></div>	https://treatment.plazi.org/id/03D087EFFFB9FF81FF3BFF2F8CF9141C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Xue, Guo-Xi;Inayoshi, Yutaka;Saito, Kotaro;Li, Meng;Rao, Zhi	Xue, Guo-Xi, Inayoshi, Yutaka, Saito, Kotaro, Li, Meng, Rao, Zhi (2026): Capila jayadeva Moore, 1866 is a sexually dimorphic and male polymorphic species including six subspecies (Hesperiidae, Pyrginae). Zootaxa 5748 (1): 89-100, DOI: 10.11646/zootaxa.5748.1.4, URL: https://doi.org/10.11646/zootaxa.5748.1.4
