taxonID	type	description	language	source
03D087DF991FFF8EFE22FC01FD136075.taxon	description	urn: lsid: zoobank. org: act: 2 F 134 DDB-AFE 1 - 4 B 24 - AB 32 - E 1926 A 8 AE 2 E 2 Figs 1 F, 2 – 7, 10, Table 1 – 2	en	Sendra, Alberto, Mayvan, Mahmood Mehrafrooz, Selfa, Jesus, Jiménez-Valverde, Alberto, Kováč, Ľubomír (2026): A new species of an interesting cave-adapted dipluran (Diplura, Campodeidae, Plusiocampinae) from the Kopet Dagh Mountains in Iran. European Journal of Taxonomy 1034: 85-102, DOI: 10.5852/ejt.2026.1034.3159, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3159/14063
03D087DF991FFF8EFE22FC01FD136075.taxon	diagnosis	Diagnosis Body robust, length 4 − 6 mm; epicuticle is smooth; 32 – 38 antennomeres; cupuliform organ with about 10 complex olfactory chemoreceptors compounded by a central column surrounded by a spiral fold of 5 – 6 turns; pronotum with 1 + 1 ma, 2 + 2 − 4 + 4 la 1 − 4, 2 + 2 lp 2, 3 macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la 2 − 3, 2 + 2 lp 2 − 3, 1 + 1 mp and 1 + 1 ma, 1 + 1 la, 2 + 2 lp 2 − 3, 1 + 1 mp long macrosetae; metathoracic legs reach the eighth abdominal segment; tibia is 1.1 – 1.3 times as long as the femur or tarsus; femora II – III have 3 long dorsal macrosetae; tibia II – III with 3 − 2 ventral macrosetae; calcars with long barbs almost all over; end part of the tarsus has two dorsal and one lateral long robust pubescens setae; claws are similar in size, both with large lateral crests; lateral processes have a basal laminar boarder that narrows along the distal half and all borders prolongated in long fringes; 1 + 1 post 1 macrosetae on I – III urotergites, 1 + 1 la 1, 3 + 3 post 1 – 3 macrosetae on IV urotergite, 1 + 1 la 1, 5 + 5 post 1 − 5 macrosetae on V urotergite; 2 + 2 − 3 + 3 la 1 − 3, 5 + 5 post 1 − 5 macrosetae on VI−VII urotergites, 7 + 7 − 8 + 8 post macrosetae on VIII urotergites and 11 + 11 post macrosetae on IX abdominal segment; urosternite I with 10 + 10 − 13 + 13 well-barbed macrosetae; urosternites II – VII with 7 + 7 macrosetae, urosternite VIII with 2 + 2 macrosetae; the stylus setae are well barbed with almost complete thin barbs; female urosternite I with subcylindrical appendages, each with 26 glandular a 1 setae and male with larger subcylindrical appendages with up to 34 glandular setae a 1.	en	Sendra, Alberto, Mayvan, Mahmood Mehrafrooz, Selfa, Jesus, Jiménez-Valverde, Alberto, Kováč, Ľubomír (2026): A new species of an interesting cave-adapted dipluran (Diplura, Campodeidae, Plusiocampinae) from the Kopet Dagh Mountains in Iran. European Journal of Taxonomy 1034: 85-102, DOI: 10.5852/ejt.2026.1034.3159, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3159/14063
03D087DF991FFF8EFE22FC01FD136075.taxon	etymology	Etymology The specific epithet ‘ pax ’ is a Latin noun meaning ‘ peace’. It is used here as a noun in apposition and does not change to agree in gender with the genus name. The name is intended as a symbolic tribute to peace, reflecting a personal aspiration for harmony in both nature and society; and a relevant desire in these times of war in the area studied.	en	Sendra, Alberto, Mayvan, Mahmood Mehrafrooz, Selfa, Jesus, Jiménez-Valverde, Alberto, Kováč, Ľubomír (2026): A new species of an interesting cave-adapted dipluran (Diplura, Campodeidae, Plusiocampinae) from the Kopet Dagh Mountains in Iran. European Journal of Taxonomy 1034: 85-102, DOI: 10.5852/ejt.2026.1034.3159, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3159/14063
03D087DF991FFF8EFE22FC01FD136075.taxon	materials_examined	Type material Holotype IRAN • ♀; Moghan Cave, Moghan village, Mashhad City, Razavi Khorasan Province; 36 ° 06 ʹ 59 ʺ N, 59 ° 22 ʹ 06 ʺ E; 2193 m a. s. l.; 29 Jul. 2024; Mahmood Mehrafrooz Mayvan leg.; ZMFUM-MoAn 101 IR. Paratypes IRAN • 1 ♂; Moghan Cave, Moghan village, Mashhad City, Razavi Khorasan province; 36 ° 06 ʹ 59 ʺ N, 59 ° 22 ʹ 06 ʺ E; 2193 m a. s. l.; 29 Jul. 2024; Mahmood Mehrafrooz Mayvan leg; ZMFUM-MoAn 104 IR. • 1 ♂; same collection data as for holotype; ZMFUM-MoAn 102 IR • 1 ♂; same collection data as for holotype; ZMFUM-MoAn 103 IR. • 1 ♂; same collection data as for holotype; MUVHN-ZE 6928 • 1 ♀; same collection data as for holotype; MUVHN-ZE 6976 • 1 ♀; same collection data as for holotype; MCNB- 2025 - 7270 • 1 ♂; same collection data as for holotype; MCNB- 2025 - 7271 • 1 ♂ same collection data as for holotype; UPJŠ- 01 - 25 • ♂ same collection data as for holotype; UPJŠ- 02 - 25. Other studied material IRAN • 2 ♂♂, 4 ♀♀; Moghan Cave, Moghan village, Mashhad City, Razavi Khorasan Province, 36 ° 06 ʹ 59 ʺ N, 59 ° 22 ʹ 06 ʺ E, 2193 m a. s. l.; 29 Jul. 2024; Mahmood Mehrafrooz Mayvan leg.; AS coll. Used for scanning electron microscopy.	en	Sendra, Alberto, Mayvan, Mahmood Mehrafrooz, Selfa, Jesus, Jiménez-Valverde, Alberto, Kováč, Ľubomír (2026): A new species of an interesting cave-adapted dipluran (Diplura, Campodeidae, Plusiocampinae) from the Kopet Dagh Mountains in Iran. European Journal of Taxonomy 1034: 85-102, DOI: 10.5852/ejt.2026.1034.3159, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3159/14063
03D087DF991FFF8EFE22FC01FD136075.taxon	description	Taxa description BODY. Body length 4 − 5 mm in males, 5.1 − 6 mm in females, and 3.1 mm in a juvenile (Fig. 1 F). The epicuticle is smooth even under high magnifications (Fig. 4 A−C); body robust and covered by clothing setae with thin barbs along half to one-third distal portion. HEAD. Antennae have 32 – 38 antennomeres in intact adult antennae, and 28 antennomeres in a female juvenile; antennae ~ 0.88 − 0.80 times as long as the body’s length, with medial antennomeres 1.5 times as long as wide and apical antennomere more than two times as long as wide. Two types of gouge sensilla and glandular setae cover the last antennomere: numerous long and slim gouge sensilla, scarce large and thick gouge sensilla, and an apical whorl of glandular setae around the cupuliform organ (Fig. 2 C−D). The cupuliform organ occupies ¹ ⁄ 11 of total length in the last antennomere and has about 10 complex olfactory chemoreceptors compounded by a central column surrounded by a spiral fold of five to six turns (Fig. 2 A−B). The distal and central antennomeres have three whorls of slightly barbed macrosetae, in addition to smooth, thin setae. They also have a distal whorl of 12 − 14 long, slim gouge sensilla (38 – 50 µm long) (Fig. 2 E−F). The proximal antennomeres have typical trichobothria, plus a small bacilliform sensillum on the third antennomere in ventral position. The frontal process has a slightly protrusion with one distal smooth macroseta, plus three shorter macrosetae with a few distal barbs; the three-insertion line macrosetae and x macrosetae bear a few distal barbs, and their length ratios are: a / i / p / x 43 / 49 / 41 / 62, respectively, in the holotype; occiput of the dorsal head with 3 + 3 ma, la, lp macrosetae bears a few distal barbs. The labial palps are large and suboval with large coniform latero-external sensillum, two guard setae, up to fourteen setae on anterior border, and up to 100 neuroglandular setae in the holotype. THORAX. Thoracic macroseta distribution: pronotum with 1 + 1 ma, 2 + 2 − 4 + 4 la 1 – 4, 2 + 2 lp 2 − 3 macrosetae; mesonotum with 1 + 1 ma, 2 + 2 la 2 − 3, 2 + 2 lp 2 − 3, 1 + 1 mp and 1 + 1 ma, 1 + 1 la, 2 + 2 lp 2 − 3, 1 + 1 mp long macrosetae (Figs 3, 4 A−C). All macrosetae are long and with thin barbs along ⅔ of their basal length; marginal setae are thin and barbed along two-thirds of the distal portion and double the length of the clothing setae. The legs are slightly elongated, and the metathoracic legs reach the eighth abdominal segment. The tibia is 1.1 − 1.3 times as long as the femur or tarsus. Femora II – III have 3 long dorsal macrosetae, 2 in a few specimens (femora I with 2 dorsal macrosetae); tibia II−III with 3 – 2 ventral macrosetae (Fig. 5 A−B, D). The calcars have long barbs almost all over (Fig. 5 C). The tarsus has two ventral rows of ventral setae with barbs on medial portion setae and a few setiform sensilla; the end part of the tarsus has two dorsal and one lateral long robust pubescens setae (Fig. 6 A−D). The claws are similar in size, both with large lateral crests. Lateral processes have a basal laminar boarder that narrows along the distal half and all borders prolongated in long fringes (Fig. 6 A−D). ABDOMEN. Distribution of abdominal macrosetae on tergites: 1 + 1 post 1 macrosetae on I – III urotergites, 1 + 1 la 1, 3 + 3 post 1 - 3 macrosetae on IV urotergite, 1 + 1 la 1, 5 + 5 post 1 – 5 macrosetae on V urotergite; 2 + 2 – 3 + 3 la 1 – 3, 5 + 5 post 1 - 5 macrosetae on VI – VII urotergites, 7 + 7 − 8 + 8 post macrosetae on VIII urotergites and 11 + 11 post macrosetae on IX abdominal segment (Fig. 7 A−D). All tergal abdominal macrosetae are long and well-differentiated with thin barbs along most of their length (Fig. 7 A−D). Urosternite I with 10 + 10 − 13 + 13 well-barbed macrosetae (Fig. 4 d), urosternites II – VII with 7 + 7 macrosetae (Fig. 4 E), urosternite VIII with 2 + 2 macrosetae; urosternal macrosetae barbed with thin barbs almost from the base to two-thirds. The stylus setae is well barbed with almost complete thin barbs. (Fig. 4 F). Two completed cerci with six (paratype ♂ 04) and seven (holotype) primary articles slightly longer from proximal to distal, one in addition to a basal article divided into three secondary articles, both slightly longer than the body length, 1.2 for the paratype ♂ 04 and 1.1 for holotype; primary article bears 6 − 12 whorls of long macrosetae with thin barbs along distal ⅓, one distal whorl of smooth setae, and one apical whorl of thin setae with thin barbs along distal ⅓. SECONDARY SEX CHARACTERS. Female urosternite I with subcylindrical appendages, each bearing up to 26 glandular a 1 setae in a distal field. Male with larger subcylindrical appendages than the female, each bearing up to 34 glandular setae a 1 (Fig. 4 D).	en	Sendra, Alberto, Mayvan, Mahmood Mehrafrooz, Selfa, Jesus, Jiménez-Valverde, Alberto, Kováč, Ľubomír (2026): A new species of an interesting cave-adapted dipluran (Diplura, Campodeidae, Plusiocampinae) from the Kopet Dagh Mountains in Iran. European Journal of Taxonomy 1034: 85-102, DOI: 10.5852/ejt.2026.1034.3159, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3159/14063
