identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D2A373FFDAFFAFFF26F9ADFD822CD2.text	03D2A373FFDAFFAFFF26F9ADFD822CD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triozidus Li 1994	<div><p>Genus Triozidus Li, 1994 stat. rev.</p><p>Triozidus Li, 1994 . Type species: Triozidus acanthopanaicis Li, 1994: 85, by original designation.</p><p>Lunatrioza Kwon, Suh &amp; Kwon, 2022. Type-species: Trioza stackelbergi Loginova, 1967: 345, by original designation; nomen nudum [conference poster, not a published work in the sense of the International Code of Zoological Nomenclature], syn. nov.</p><p>Adult colour and structure. General body colour yellow-brown to dark brown, with light to mid-brown on areas of dorsum and legs, and darker brown to black on some or all of antennal flagellum, head and pronotum, and abdominal tergites. Forewing membrane clear or yellowish, with or without distinct dark patch around clavus. Medium to large bodied, 3.5–6.0 mm (including wings). Head narrower to slightly wider than thorax, inclined at about 45–50º from longitudinal body axis (in lateral view). Head with well developed, medium-long genal processes, vertex trapezoidal, moderately concave medially at posterior margin, lateral ocelli on small tubercles, medial epicranial suture distinct. Clypeus narrowly rounded ventrally, distal proboscis segment short.Antennae medium long, slender, 10-segmented, segment 3 longest, a single rhinarium subapically on segments 4, 6, 8 and 9. Thorax weakly to moderately arched. Pronotum deflexed from mesothorax at a 45° angle. Legs moderately short and robust, tibia longer than femur; hind leg with meracanthus well developed, horn-shaped, more or less straight, acute at apex; metatibia length equal to or longer than head width, with a single or cluster of small spines basally, inflated apically and bearing 3 (1+2) sclerotized apical spurs (not stalked) and a row of unsclerotized bristles; metabasitarsus without apical spurs, slightly longer than apical metatarsus. Forewing relatively large, length approximately 1.2–1.5 x body length, broadest in the middle or distal to the middle, with trifurcation of veins R+M+Cu strict; membrane lacking surface spinules except occasional small area with spinules at base of cell cu 2, a narrow cluster of marginal radular spines present at wing margin, more or less centrally positioned in cells m 2 and cu 1, and in the posterior half of cell m 1; size of cell m 1 subequal to, or notably larger than, cell cu 1; vein Rs either long and sinuous or medium short and evenly curving to fore margin; forewing apex subacute to acute. Hindwing medium-short, length less than 2/3 forewing length. Male terminalia with subgenital plate globose, posterior margin more or less rounded or more angular (in lateral view); proctiger short with slight to moderate posterior lobe medially, length subequal or shorter than paramere; paramere simple (in lateral profile), more or less parallel sided, either straight, moderately sinuous or with incision medially on anterior surface, tapering to blunt or pointed apex; distal aedeagus segment either distinctly angled or arched with a bipartite projection medially consisting of either thumb-like or semicircular projections, or less typically more or less straight and lacking such projections, apex inflated and developed into a shallow or widely curving hook. Female terminalia short, length approximately subequal to head width, proctiger slightly longer than subgenital plate, anal ring large and consisting of a single row of pores. Ovipositor dorsal valvulae relatively small and cuneate, ventral valvulae with one or a few shallow serrations.</p><p>Immature colour and structure. Fifth instar immature. Coloration. General body colour yellowish-brown to dark brown, with light to mid-brown areas on dorsum and legs, and darker brown to black on some sclerites and terminal antennal segments. Body shape in dorsal view elongate-oval with wing pads protruding and humeral lobes developed; sclerites present dorsally and ventrally. Body surface covered with medium dense simple setae or sectasetae. Antenna weakly curved, 8-segmented; scape, pedicel and segment 3 thicker than remaining flagellum; with a single subapical rhinarium on each of segments 4 and 6, and two on segment 8. Legs moderately long, femur shorter than tibiotarsus; tarsi with only one fully developed claw (outer), tarsal arolium longer than claws, triangular with a medial cleft, with short unguitractor, lacking pedicel. Humeral lobe of forewing pad relatively short, reaching hind margin of eye. Caudal plate broadly rounded posteriorly. Circumanal ring in ventral position, relatively wide and short; outer ring mostly composed of a single row of elongate pores, or with convolutions and intermittent multi-pore sections.</p><p>Egg. Only known for T. stackelbergi . Oblong-ovoid with a short, laterally positioned pedicel sub-basally on underside and short tail (illustrated in Loginova 1967).</p><p>Host plant. Eleutherococcus spp. ( Araliaceae).</p><p>Biology. All species for which biology is known produce enclosed galls on some or all of the following host organs: leaves, twigs, male and female flowers, petioles or petiolules.</p><p>Distribution. The genus occurs in China (Li 2011), Japan (Shinji 1940, Miyatake 1996), Russia (Loginova 1967, Konovalova 1988), South Korea (Cho et al. 2017, Kwon &amp; Kwon 2020), and Taiwan (this study).</p><p>Comment. The genus includes four described nominal species and two new species. Kwon &amp; Kwon (2020) synonymised Triozidus acanthopanaicis Li, 1994, the type species of Triozidus, with Heterotrioza stackelbergi . Species of Triozidus sensu novo have been variously referred to Heterotrioza Dobreanu &amp; Manolache, Heterotrioza (Dyspersa) Klimaszewski, Trioza or Triozidus (Klimaszewski 1973; Li 2011; Kwon &amp; Kwon 2020; Cho et al. 2022). Loginova (1967) discussed morphological differences between T. ukogi and T. stackelbergi, and Cho et al. (2017) pointed out that the latter has been misidentified repeatedly in the literature from Korea as either Trioza ukogi, Heterotrioza (Dyspersa) ukogi or Heterotrioza ukogi .</p><p>Unusual features of the genus typically, but not exclusively, include the bipartite projection on the shaft of the apical aedeagus segment (i.e., not on the apex as in Heterotrioza), and the immatures with uneven or asymmetrical development of tarsal claws; interestingly, it seems that the undeveloped claw becomes increasingly vestigial in later instars. Many triozid groups are known to be relatively morphologically homogeneous resulting in peripatetic taxonomic histories, and this is especially the case with taxa formerly placed within the catch-all genus Trioza . These challenges have increased the reliance on molecular data, and the reciprocal illumination these data offer, to help resolve morphologically challenging groups. Nevertheless, Triozidus has a number of morphological characters that collectively serve to help distinguish it from related groups. In particular, the large size of the forewing relative to body size is more exaggerated than in related genera (e.g., Dyspersa, Heterotrioza, Lauritrioza Conci &amp; Tamanini,</p><p>Trioza s.s.), the forewing cell m 1 is always larger than cu 1 (usually markedly so), the transition from vertex to genal cones is not as distinctly stepped, and the apex of the metatibia is more expanded. These, together with the biology and host plants in Eleutherococcus, and the immature vestigial claw, define Triozidus . Relationships with other</p><p>Trioza sensu lato taxa and other genera in Group B (as shown in Fig. 16), are still in flux taxonomically as these groups become more well defined, but the same assemblage of characters given above, together with the molecular data, provides the best diagnosis.</p></div>	https://treatment.plazi.org/id/03D2A373FFDAFFAFFF26F9ADFD822CD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.	Liao, Yi-Chang, Inoue, Hiromitsu, Percy, Diana M. (2025): The psyllid genus Triozidus Li, 1994 stat. rev. sensu novo (Hemiptera: Psylloidea: Triozidae) in East Asia is redefined with the addition of two new species from Taiwan inducing galls on the leaflet petiolules of Eleutherococcus trifoliatus (Araliaceae). Zootaxa 5727 (1): 113-139, DOI: 10.11646/zootaxa.5727.1.8, URL: https://doi.org/10.11646/zootaxa.5727.1.8
03D2A373FFDCFFAFFF26FDB8FA142A30.text	03D2A373FFDCFFAFFF26FDB8FA142A30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triozidus Li 1994	<div><p>Key to adults of Triozidus</p><p>1. Forewing with Rs vein long and sinuous (terminating distal to a line drawn through termination of vein Cu 1a and bifurcation of vein M); cell cu 1 higher resulting in cell m 1 not being markedly larger than cell cu 1 (Fig. 9B); on Eleutherococcus spinosus (or other Eleutherococcus spp. where alien................................... Triozidus ukogi (Shinji, 1940) comb. nov.</p><p>- Forewing with Rs short and more or less evenly curving to the fore margin (terminating at or proximal to a line drawn through termination of vein Cu 1a and bifurcation of vein M); cell cu 1 lower and cell m 1 markedly larger than cell cu 1 (Figs 2E, 5C, 13D); on other Eleutherococcus species (where host plant is known)................................................. 2</p><p>2. Antennae shorter than 1.5 x head width; apex of paramere (in lateral profile) tapering to a slender, anteriorly directed point (Fig. 16)............................................................ Triozidus ceratophorus (Li, 2005) comb. nov.</p><p>- Antennae longer than 1.5 x head width; apex of paramere (in lateral profile) more or less blunt and directed inward or posteriorly (Figs 3B, 6B, 14B, 16)................................................................................. 3</p><p>3. Body including forewing shorter than 4.5 mm; distal aedeagus segment lacking medial projections (Figs 3C, 16)......... 4</p><p>- Body including forewing longer than 4.5 mm; distal aedeagus segment with distinct medial projections (Figs 6C, 14C, 16).. 5</p><p>4. Head and antennae entirely black (Fig. 1A); paramere (in lateral profile) with distinct angular incision medially on anterior surface, apex rounded (Figs 3A, 3B, 16)............................................ Triozidus burckhardti sp. nov.</p><p>- Head mostly yellow-brown to orange-brown with probably only terminal antennal segments black; paramere (in lateral profile) without angular medial incision but with slight incision near the base on anterior surface, apex elongate and extended rearward (Fig. 16)............................................... Triozidus eleutherococci (Konovalova, 1980) comb. nov.</p><p>5. Genal processes shorter (length just over half vertex length, &lt;0.3 head width) and more blunt apically (Fig. 13A); forewing with distinct dark marginal patch around clavus, but without a small dark spot at trifurcation of R+M+Cu (Fig. 13D); forewing broader (ratio length:width &lt;2.6), widest distal to middle, length approximately 2 x Rs vein length (Fig. 13D); paramere broader (in lateral profile) (Figs 14A, 14B, 16); distal aedeagus segment with medial projections narrow and thumb-like (Figs 14C, 16); female terminalia shorter (length of proctiger less than head width and &lt;3 x anal ring length) (Fig. 14D); on Eleutherococcus trifoliatus ..................................................................... Triozidus yangorum sp. nov.</p><p>- Genal processes longer (length approximately equal to vertex length,&gt; 0.3 x head width) and more acute apically (Fig. 5A); forewing without distinct dark marginal patch around clavus, but with a small dark spot at trifurcation of R+M+Cu (in Japanese and South Korean material, see Fig. 5C, but possibly absent in eastern Russia material); forewing narrower (ratio length:width&gt; 2.6), widest in middle, length approximately 2.5 x Rs vein length; paramere narrower (in lateral profile) (Figs 6A, 6B, 16); distal aedeagus segment with medial projections semi-circular (Figs 6C, 16); female terminalia longer (length of proctiger greater than head width and&gt; 3 x anal ring length) (Fig. 6D); on Eleutherococcus sessiliflorus and E. divaricatus (or other Eleutherococcus spp. where introduced).......................... Triozidus stackelbergi (Loginova, 1967) comb. nov.</p><p>Key to 5 th instar immatures of Triozidus</p><p>(known for three species)</p><p>1. Body covered in narrow truncate sectasetae (Fig. 7C)............... Triozidus stackelbergi (Loginova, 1967) comb. nov.</p><p>- Body covered in long simple setae (Figs 11C, 15C).......................................................... 2</p><p>2. Circumanal ring heart-shaped (medial constriction mainly from anterior), outer ring composed of a mix of elongate and irregular shaped pores (Fig. 15D)......................................................... Triozidus yangorum sp. nov.</p><p>- Circumanal ring peanut-shaped (with antero-posterior medial constriction), outer ring composed of a single narrow row of elongate pores (Fig. 11D)............................................. Triozidus ukogi (Shinji, 1940) comb. nov.</p></div>	https://treatment.plazi.org/id/03D2A373FFDCFFAFFF26FDB8FA142A30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.	Liao, Yi-Chang, Inoue, Hiromitsu, Percy, Diana M. (2025): The psyllid genus Triozidus Li, 1994 stat. rev. sensu novo (Hemiptera: Psylloidea: Triozidae) in East Asia is redefined with the addition of two new species from Taiwan inducing galls on the leaflet petiolules of Eleutherococcus trifoliatus (Araliaceae). Zootaxa 5727 (1): 113-139, DOI: 10.11646/zootaxa.5727.1.8, URL: https://doi.org/10.11646/zootaxa.5727.1.8
03D2A373FFDDFFAEFF26FF0CFCB82AB9.text	03D2A373FFDDFFAEFF26FF0CFCB82AB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triozidus burckhardti Liao & Inoue & Percy 2025	<div><p>Triozidus burckhardti Liao &amp; Percy sp. nov.</p><p>(Figs 1A–B, 2, 3)</p><p>Type material. Holotype: TAIWAN • ♂; Pingtung Co., Manzhou, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.835976&amp;materialsCitation.latitude=22.08164" title="Search Plazi for locations around (long 120.835976/lat 22.08164)">Changle</a>; 22°04’53.9”N, 120°50’09.5”E; 29 Jan 2024; Y. C. Liao leg.; Eleutherococcus trifoliatus; NCHU, dry mounted . Paratypes: TAIWAN • 1♂, 1♀; same data as for holotype, but in ethanol and slide mounted . 1♂, 2♀; same data as for holotype, but DMPC, in ethanol . 1♂, 1♀; same data as for holotype, but NHMB, in ethanol . 3♀; same data as for holotype, but 26 Mar 2020 .</p><p>Description. Adult (Figs 1A, 2, 3). Coloration. Body color yellowish brown with entire head, compound eyes and antennae black (Fig. 1A). Abdominal tergites blackish brown. Legs yellow. Forewing pale yellow, transparent, slightly darker brown around clavus (Fig. 2E). Hindwing transparent.</p><p>Structure. Body median-sized, length from anterior head margin to tip of folded forewing 3.6–3.8 mm, female generally larger. Head (Fig. 2A) slightly narrower than thorax. Vertex width 1.7–1.8 x length, minutely pubescent. Genal processes prominent, length along mid-line 0.7–0.8 x vertex length, divergent, conical, acute at apex, covered in short hair. Antenna (Fig. 2B) length 1.8–2.0 x head width, antennal segment 3 approximately double length of segment 4, relative length of flagellar segments as 1.0: 0.5: 0.3: 0.4: 0.3: 0.3: 0.2: 0.2; two unequal terminal setae: longer seta 0.7 x, and shorter seta 0.3 x segment 10 length. Thorax moderately arched dorsally, minutely pubescent. Hind leg (Fig. 2C) metatibia length 1.0–1.1 x head width. Forewing moderately broad (Fig. 2E), length 5.8–5.9 x head width, and 2.9–3.0 x width, widest slightly distal to the middle; wing apex acute; vein Rs moderately long, gradually curved to fore margin; vein M evenly curved with bifurcation posterior to line connecting apices of veins Rs and Cu1 a; cell m 1 larger than cell cu 1; vein Cu 1a evenly curved, vein Cu 1b straight; veins minutely pubescent. Hindwing (Fig. 2D) 0.6 x as long and 0.5 x as wide as forewing.</p><p>Male terminalia (Figs 3A–C). Proctiger short, with moderate posterior lobes reaching maximum extension in the basal half (Fig. 3A). Subgenital plate subglobular, with medium long setae laterally and ventrally; posterior dorsal margin slightly convex. Paramere (Fig. 3B) slightly longer than proctiger; in profile angulate at base, with a deep incision anteriorly in the basal half, before narrowing to apex which is acute and directed inward; inner face and surface beset with median-long setae. Distal aedeagus segment (Fig. 3C) about as long as paramere (Figs 3A, 16), straight medially without projection, apical portion largely inflated into a hook with acute apex; sclerotized end tube of ductus ejaculatorius short, slightly sinuous. Female terminalia (Fig. 3D) cuneate, short; proctiger dorsal margin slightly sinuate with medial depression and acute apex, only slightly longer than subgenital plate, with long setae in the apical half, anal ring length approximately one third proctiger length, consisting of a single row of pores. Subgenital plate, in profile, triangular, apex acute, beset with long hairs laterally and ventrally. Ovipositor dorsal valvulae cuneate, ventral valvulae straight with several shallow serrations apically.</p><p>Measurements in mm (1 male, 1 female). Body length (including forewing) ♂ 3.56; ♀ 3.75. Head width ♂ 0.47; ♀ 0.52. Vertex width ♂ 0.27; ♀ 0.31. Vertex length ♂ 0.16; ♀ 0.17. Genal cone length ♂ 0.11; ♀ 0.13. Antenna length ♂ 0.93; ♀ 0.93. Metatibia length ♂ 0.50; ♀ 0.53. Forewing length ♂ 2.75; ♀ 3.0. Forewing width ♂ 0.93; ♀ 1.05. Aedeagus length ♂ 0.20. Paramere length ♂ 0.22. Proctiger length ♀ 0.62. Subgenital plate length ♀ 0.60.</p><p>Etymology. The specific name honours Dr Daniel Burckhardt for his contribution to our knowledge of the world’s psyllid fauna.</p><p>Distribution. Taiwan; only found in the southernmost part (Hengchun peninsula).</p><p>Host plant. Probably Eleutherococcus trifoliatus ( Araliaceae), but immature specimens are needed to confirm host and gall type.</p><p>Biology. This species apparently shares the same host plant as T. yangorum but the two psyllid species are allopatric, with T. burckhardti only found in the southernmost part of Taiwan and T. yangorum in central and northern Taiwan. They also appear to have a similar galling biology, galls and immatures were recorded for T. burckhardti but no immature specimens were preserved. Immatures (1 st instar) were observed to develop within fully enclosed round galls that are positioned on the base of leaflets, or just below the leaflets on the petiolules, or on the leaf petioles. Each gall consists of a single gall chamber with usually a single immature. Galls are found individually or in aggregates of two to four galls per leaf, often with a gall on each of the three petiolules (Fig. 1B). As is the case for T. yangorum, T. burckhardti is univoltine with adults emerging in January.</p><p>Genetic resources. Adult male and female sequences of COI and cytB were identical, unique haplotypes: PQ817990 and PQ817253 (for COI and cytB respectively).</p></div>	https://treatment.plazi.org/id/03D2A373FFDDFFAEFF26FF0CFCB82AB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.	Liao, Yi-Chang, Inoue, Hiromitsu, Percy, Diana M. (2025): The psyllid genus Triozidus Li, 1994 stat. rev. sensu novo (Hemiptera: Psylloidea: Triozidae) in East Asia is redefined with the addition of two new species from Taiwan inducing galls on the leaflet petiolules of Eleutherococcus trifoliatus (Araliaceae). Zootaxa 5727 (1): 113-139, DOI: 10.11646/zootaxa.5727.1.8, URL: https://doi.org/10.11646/zootaxa.5727.1.8
03D2A373FFD0FFA3FF26FCBDFB9929C4.text	03D2A373FFD0FFA3FF26FCBDFB9929C4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triozidus ceratophorus (Li 2005) Liao & Inoue & Percy 2025	<div><p>Triozidus ceratophorus (Li, 2005) comb. nov.</p><p>Heterotrioza ceratophora Li, 2005: 201 .</p><p>Material examined. None.</p><p>Description. Adult described from a single male (Li 2005, 2011).</p><p>Host plant. Unknown.</p><p>Distribution. China (Gansu) (Li 2005, 2011).</p><p>Comment. Li’s (2011) key to Heterotrioza separates T. ceratophorus and T. stackelbergi from other taxa that belong in Heterotrioza based on wing characters that highlight the much larger size of cell m 1 relative to cu 1 in the Triozidus taxa. Additionally, T. ceratophorus possesses the medial projections on the distal aedeagus segment that is typical for Triozidus, versus the terminal projections typical for Heterotrioza .</p></div>	https://treatment.plazi.org/id/03D2A373FFD0FFA3FF26FCBDFB9929C4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.	Liao, Yi-Chang, Inoue, Hiromitsu, Percy, Diana M. (2025): The psyllid genus Triozidus Li, 1994 stat. rev. sensu novo (Hemiptera: Psylloidea: Triozidae) in East Asia is redefined with the addition of two new species from Taiwan inducing galls on the leaflet petiolules of Eleutherococcus trifoliatus (Araliaceae). Zootaxa 5727 (1): 113-139, DOI: 10.11646/zootaxa.5727.1.8, URL: https://doi.org/10.11646/zootaxa.5727.1.8
03D2A373FFD0FFA3FF26FA83FCDC2A01.text	03D2A373FFD0FFA3FF26FA83FCDC2A01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triozidus eleutherococci (Konovalova 1980) Liao & Inoue & Percy 2025	<div><p>Triozidus eleutherococci (Konovalova, 1980) comb. nov.</p><p>Trioza eleutherococci Konovalova, 1980: 21 .</p><p>Material examined. None.</p><p>Description. Adult male and female in Konovalova (1980), with additional information in the key in Konovalova (1988).</p><p>Host plant. Given as Eleutherococcus senticosus in Konovalova (1980) but no mention of galls.</p><p>Distribution. Russia (Primorsky Krai, Russian Far East) (Konovalova 1980, 1988).</p><p>Comment. This species, like T. burckhardti, lacks the medial projections on the distal aedeagus segment. The paramere shape is diagnostic with the apex extended rearward more so than in other Triozidus (Fig. 16). Other distinguishing characters that also suggest this species may be most closely related to T. burckhardti are small size (overall body size less than 4.5 mm, forewing length less than 3.5 mm) (Fig. 16), relatively narrow forewing, and genal cones that are short and acute (Konovalova 1980, 1988).</p></div>	https://treatment.plazi.org/id/03D2A373FFD0FFA3FF26FA83FCDC2A01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.	Liao, Yi-Chang, Inoue, Hiromitsu, Percy, Diana M. (2025): The psyllid genus Triozidus Li, 1994 stat. rev. sensu novo (Hemiptera: Psylloidea: Triozidae) in East Asia is redefined with the addition of two new species from Taiwan inducing galls on the leaflet petiolules of Eleutherococcus trifoliatus (Araliaceae). Zootaxa 5727 (1): 113-139, DOI: 10.11646/zootaxa.5727.1.8, URL: https://doi.org/10.11646/zootaxa.5727.1.8
03D2A373FFD0FFA6FF26F8C8FAD32C42.text	03D2A373FFD0FFA6FF26F8C8FAD32C42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triozidus stackelbergi (Loginova 1967) Liao & Inoue & Percy 2025	<div><p>Triozidus stackelbergi (Loginova, 1967) comb. nov.</p><p>(Figs 4–7)</p><p>Trioza stackelbergi Loginova, 1967: 345 .</p><p>Heterotrioza (Dyspersa) stackelbergi: Klimaszewski (1973: 248).</p><p>Triozidus acanthopanaicis Li, 1994: 85, synonymised by Kwon &amp; Kwon (2020: 220).</p><p>Heterotrioza acanthopanaicis: Li (2011: 1521).</p><p>Heterotrioza stackelbergi: Kwon &amp; Kwon (2020: 220), Cho et al. (2022: 72).</p><p>Lunatrioza stckelbergi [sic] Kwon, Suh &amp; Kwon, 2022: nomen nudum [conference poster, not a published work in the sense of the International Code of Zoological Nomenclature].</p><p>Material examined. KOREA • 4 ♂, 3♀; Yeong wol-gun, Jucheon-myeon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.49081&amp;materialsCitation.latitude=37.435917" title="Search Plazi for locations around (long 128.49081/lat 37.435917)">Jucheon-ri</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=128.49081&amp;materialsCitation.latitude=37.435917" title="Search Plazi for locations around (long 128.49081/lat 37.435917)">Swimteo Park</a>; 37°26’09.3”N, 128°29’26.9”E; 3 Oct 2014; Jin-Yeong Choi leg.; NCHU, slide mounted, glycerol and ethanol . JAPAN • 6♀, 1 immature; Honshu, Nagano Pref., Kitasaku-gun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.46349&amp;materialsCitation.latitude=35.93515" title="Search Plazi for locations around (long 138.46349/lat 35.93515)">Minamimaki-mura</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.46349&amp;materialsCitation.latitude=35.93515" title="Search Plazi for locations around (long 138.46349/lat 35.93515)">Nobeyama</a>; 35.93515 N, 138.46349 E, 1370 m; 16 Oct. 2018; Akihide Koguchi leg.; Eleutherococcus divaricatus (twig galls, flower galls); HIC, dry, slide mounted . 15♂, 14♀, 4 immatures; same data but 6. Oct. 2019; HIC, NCHU, dry, slide mounted and in ethanol . 20♂, 27♀, 18 immatures; Honshu, Nagano Pref., Kitasaku-gun, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.4725&amp;materialsCitation.latitude=35.97757" title="Search Plazi for locations around (long 138.4725/lat 35.97757)">Minamimaki-mura</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=138.4725&amp;materialsCitation.latitude=35.97757" title="Search Plazi for locations around (long 138.4725/lat 35.97757)">Itabashi</a>; 35.97757 N, 138.47251 E, 1350 m; 6 Oct. 2019; Akihide Koguchi leg.; Eleutherococcus divaricatus (twig galls, flower galls); HIC, NCHU, dry, slide mounted and in ethanol .</p><p>Note. The redescription below is based on material examined from Japan and South Korea, as well as the description in Loginova (1967) of eastern Russia material and notes on Chinese specimens by Li (2011).Measurements given below are taken from the material from Japan and South Korean, with measurements, if different and where reported in Loginova (1967), shown in [].</p><p>Description. Adult (Figs 4A–B, 5, 6). Coloration. General body color brown (Japan and South Korean), or yellow-orange with parts brown (eastern Russia; Loginova 1967). Antennae with apical half of 1 st segment and the remaining antenna dark brown (Japan and South Korean), or only terminal segment dark (eastern Russia; Loginova 1967). Compound eyes dark brown. Ocelli orange. Legs brown. Forewing veins brown, membrane transparent, with a small dark spot at trifurcation of R+M+Cu 1 in Japanese and South Korean material, but possibly absent in eastern Russia material or omitted from Loginova’s (1967) description.</p><p>Structure. Body large-sized, length from anterior head margin to tip of folded forewing 5.3–5.8 mm [4.6–5.7 mm], female generally larger; covered in short fine setae. Head (Fig. 5A) slightly narrower than thorax. Vertex width 1.4–1.5 x [1.6 x] length, minutely pubescent. Genal processes prominent, length along mid-line 0.8–1.0 x vertex length, divergent, conical, subacute at apex, with long pubescence. Antenna (Fig. 5B) length 2.0–2.1 x [1.9–2.0 x] head width, antennal segment 3 approximately 1.4 x segment 4, relative length of flagellar segments as 1.0: 0.7: 0.3: 0.4: 0.3: 0.3: 0.2: 0.2; two unequal terminal setae: longer seta 1.1 x, and short, truncate seta 0.2 x segment 10 length. Thorax weakly arched dorsally, minutely pubescent. Hind leg (Fig. 5E) metatibia length 1.2–1.3 x head width. Forewing (Fig. 5C) relatively long and narrow, length 6.0–6.6 x head width, and 2.8–2.9 x width, widest in the middle; wing apex acute; vein Rs moderately long, gradually curved to fore margin; vein M evenly curved with bifurcation posterior to line connecting apices of veins Rs and Cu 1a; cell m 1 larger than cell cu 1; vein Cu 1a evenly curved, vein Cu 1b straight; veins minutely pubescent. Hindwing (Fig. 5D) 0.65 x as long and 0.5 x as wide as forewing.</p><p>Male terminalia (Fig. 6A–C). Proctiger short, with moderate posterior lobes reaching maximum extension in the basal half, covered in long setae except for basal third laterally (Fig. 6A). Subgenital plate subglobular, with medium long setae laterally and ventrally; posterior dorsal margin slightly convex. Paramere (Fig. 6B) about as long as proctiger; in profile more or less parallel sided and straight, slightly sinuous, irregularly narrowing to apex which is acute and directed inward and rearward; inner face beset with long setae, outer surface with shorter setae. Distal aedeagus segment (Fig.6C) slightly shorter than paramere (Figs6A, 16),curved medially with semi-circle projections, apical portion inflated, crescent-shaped, with acute apex [eastern Russian specimens have distal aedeagus segment slightly longer, less curved medially and the semi-circular projections are less extended]; sclerotized end tube of ductus ejaculatorius short, sinuous. Female terminalia (Fig. 6D) cuneate, slender; proctiger abruptly narrowing in apical fourth with acute apex, dorsal margin more or less straight, slightly longer than subgenital plate, with long setae in the apical half, anal ring length approximately one fourth proctiger length, consisting of a single row of pores. Subgenital plate, in profile, triangular, apex acute, beset with medium long hairs laterally and ventrally. Ovipositor dorsal valvulae cuneate, ventral valvulae straight with a single shallow serration.</p><p>Fifth instar immature [based on specimens from Japan] (Fig. 7). Coloration. General color brown. Forewing pad and thorax pale brown. Body (Fig. 7A) length 1.3–1.4 x width. Body surface, including margin of head, wing pads (Fig. 7C), caudal plate and legs covered with medium dense, narrow truncate sectasetae. Antenna (Fig. 7E) relative length of antennal segments 3 to 8 as 1.0: 0.5: 0.5: 0.4: 0.6: 1.4. Legs (Fig. 7B) as for generic description. Forewing pad length 2.9 x width, and 2.6 x antenna length. Caudal plate length 0.7 x width. Circumanal ring (Fig. 7D) heart-shaped (medial constriction mainly from anterior); width 0.3 x caudal plate width; outer ring mostly composed of a single row of elongate pores with a few intermittent irregular pore shapes; inner ring composed of a mixture of short and elongate pores.</p><p>Host plant. Eleutherococcus sessiliflorus in Russia (Loginova 1967) and South Korea (Cho et al. 2017), Eleutherococcus divaricatus in Japan (this study), and Eleutherococcus senticosus in China (Li 2011) and South Korea (according to Kwon &amp; Kwon 2020).</p><p>Biology. Induces round galls on the leaf surface in South Korea (illustrated in Cho et al. 2017; Kwon &amp; Kwon 2020), but in Japan produces rounded galls, sometimes at high densities, on twigs, stems and leaf petioles, as well as on male and female flowers (Fig. 4C–D). In Japan, it appears to be bivoltine, with the first generation adults emerging in early summer (June–July) and the second generation around October (A. Koguchi, personal communication).</p><p>Distribution. China (Li 2011), Eastern Russia (Loginova 1967) and South Korea (Cho et al. 2017, Kwon &amp; Kwon 2020), and reported here for the first time for Japan.</p><p>Comment. Specimens examined from Japan and South Korea conform closely to the original description and illustrations by Loginova (1967) based on material from eastern Russia. Loginova (1967) reports a slightly smaller body size (4.6–5.7 mm) compared to the Japanese and Korean material (5.3–5.8 mm), and a forewing shape that is slightly longer and narrower, with the relative size difference between cells cu 1 and m 1 more marked; Loginova (1967) also does not mention a dark spot at trifurcation of R+M+Cu, although this may simply be omitted from her description. We also note that specimens from South Korea are marginally larger than Japanese material. Altogether, the differences equate to moderate natural variation across the species range. Kwon &amp; Kwon (2020) list this species as a pest on medicinal Eleutherococcus spp. in South Korea.</p><p>Genetic resources. Adult and immature sequences of COI were identical, unique haplotype: PQ817986.</p></div>	https://treatment.plazi.org/id/03D2A373FFD0FFA6FF26F8C8FAD32C42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.	Liao, Yi-Chang, Inoue, Hiromitsu, Percy, Diana M. (2025): The psyllid genus Triozidus Li, 1994 stat. rev. sensu novo (Hemiptera: Psylloidea: Triozidae) in East Asia is redefined with the addition of two new species from Taiwan inducing galls on the leaflet petiolules of Eleutherococcus trifoliatus (Araliaceae). Zootaxa 5727 (1): 113-139, DOI: 10.11646/zootaxa.5727.1.8, URL: https://doi.org/10.11646/zootaxa.5727.1.8
03D2A373FFD5FFA4FF26FE09FE9F2AF9.text	03D2A373FFD5FFA4FF26FE09FE9F2AF9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triozidus ukogi (Shinji 1940) Liao & Inoue & Percy 2025	<div><p>Triozidus ukogi (Shinji, 1940) comb. nov.</p><p>(Figs 8–11)</p><p>Trioza ukogi Shinji, 1940: 66 .</p><p>Heterotrioza (Dyspersa) ukogi: Klimaszewski (1973: 248).</p><p>Heterotrioza ukogi: Kwon &amp; Kwon (2020: 221); Cho et al. (2022: 72).</p><p>Lunatrioza ukogi Kwon, Suh &amp; Kwon, 2022: nomen nudum [conference poster, not a published work in the sense of the International Code of Zoological Nomenclature].</p><p>Material examined. JAPAN • 6♂ 7♀; Honshu, Tochigi Pref., Utsunomiya City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.949&amp;materialsCitation.latitude=36.525" title="Search Plazi for locations around (long 139.949/lat 36.525)">Kamikuwajima-machi</a>; 36.525 N, 139.949 E, 92 m; 28 Oct 2014; Rikio Sonobe leg.; Eleutherococcus spinosus (leaf petiole and petiolule galls); HIC, NHCU, dry, slide mounted and in ethanol . 29 immatures; same data but 10 Oct 2023; NHCU, slide mounted and in ethanol . 18♂ 7♀; Honshu, Tochigi Pref., Utsunomiya City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.846&amp;materialsCitation.latitude=36.527" title="Search Plazi for locations around (long 139.846/lat 36.527)">Shimotokami-cho</a>; 36.527 N, 139.846 E, 100 m; 16 Oct. 2015; Rikio Sonobe leg.; Eleutherococcus spinosus (leaf petiole and petiolule galls); HIC, dry mounted and in ethanol . 46 immatures, same data but 10 Oct. 2023; HIC, in ethanol . 1♂; Honshu, Tochigi Pref., Utsunomiya City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=139.881&amp;materialsCitation.latitude=36.593" title="Search Plazi for locations around (long 139.881/lat 36.593)">Nagaoka-cho</a>; 36.593 N, 139.881 E, 150 m; 13 Oct. 2015; Rikio Sonobe leg.; Eleutherococcus spinosus; HIC, in ethanol .</p><p>Description. Adult (Figs 8A, 9, 10). Coloration. Body color yellowish.Antennae yellow with apices of segments 4 to 6 dark brown, and entire segments 7 to 10 black. Compound eyes dark brown. Ocelli orange. Legs brown. Forewing pale yellow, transparent, with pale brown patch around clavus. Hindwing transparent, except clavus brown.</p><p>Structure. Body medium-sized, length from anterior head margin to tip of folded forewing 4.8–5.1 mm, female generally larger (Fig. 8A); covered in short fine setae. Head (Fig. 9A) slightly narrower than thorax. Vertex width 1.7 x length, with short pubescence. Genal processes prominent, length along mid-line 0.8 x vertex length, divergent, conical, subacute at apex, medium-long pubescence. Antenna (Fig. 9A) length approximately 1.5 x head width, antennal segment 3 approximately double length of segment 4, relative length of flagellar segments as 1.0: 0.5: 0.3: 0.3: 0.3: 0.3: 0.2: 0.2; two unequal terminal setae: longer seta 1.1 x, and short truncate seta 0.2 x, segment 10 length. Thorax moderately arched dorsally, minutely pubescent. Hind leg (Fig. 9D) metatibia length 1.0–1.1 x head width. Forewing broad (Fig. 9B), length 5.7 x head width, and 2.5 x width, widest slightly distal to the middle; wing apex subacute; vein Rs very long, sinuous and curved to apical margin; vein M evenly curved with bifurcation anterior to line connecting apices of veins Rs and Cu 1a; cell m 1 slightly larger than cell cu 1; vein Cu 1a evenly curved, vein Cu 1b more or less straight; veins minutely pubescent. Hindwing (Fig. 9C) 0.7 x as long and 0.5 x as wide as forewing.</p><p>Male terminalia (Fig. 10A–C). Proctiger short, with slight posterior lobes reaching maximum expansion in the basal half, covered in long setae except for basal third laterally (Fig. 10A). Subgenital plate subglobular, with medium long setae laterally; posterior dorsal margin slightly convex. Paramere (Fig. 10B) longer than proctiger; in profile more or less parallel sided and straight, irregularly narrowing to apex which is acute and directed inward and rearward; inner face beset with long setae. Distal aedeagus segment (Fig. 10C) distinctly shorter than paramere (Fig. 16), markedly curved medially with broad semi-circle projections, apical portion inflated into a broadly curved hook with bluntly rounded apex; sclerotized end tube of ductus ejaculatorius short, slightly sinuous. Female terminalia (Fig. 10D) cuneate, slender; proctiger narrowing in apical fourth with acute apex, dorsal margin straight, only slightly longer than subgenital plate, with long setae in the apical half, anal ring length one fourth proctiger length, consisting of a single row of pores. Subgenital plate, in profile, triangular, apex acute, beset with long hairs laterally and ventrally. Ovipositor dorsal valvulae cuneate, ventral valvulae straight apparently without serrations apically.</p><p>Fifth instar immature (Fig. 11). Coloration. General color yellow. Forewing pad and thorax also yellow. Body (Fig. 11A) length 1.3 x width. Body surface, including margin of head, wing pads (Fig. 11C), caudal plate and legs covered with sparsely distributed medium-short to long simple setae. Antenna (Fig. 11E) relative length of antennal segments 3 to 8 as 1.0: 0.7: 0.3: 0.6: 0.4: 1.8. Legs (Fig. 11A–B) as for generic description. Forewing pad length 2.7 x width, and 2.5–2.7 x antenna length. Caudal plate length 0.8 x width. Circumanal ring (Fig. 11D) peanut-shaped (medial constriction from anterior and posterior), width 0.3 x caudal plate width; outer ring composed of a single narrow row of elongate pores, inner ring composed of elongate pores.</p><p>Host plant. Eleutherococcus spinosus in Japan (Shinji, 1940); several other Eleutherococcus spp. ( E.divaricatus, E. sieboldianus, E. nodiflorus) have been mentioned for South Korea (Kwon 1983; Kwon &amp; Kwon 2020) but both hosts and distribution records remain doubtful (see Cho et al. 2017) or need to be confirmed.</p><p>Biology. Induces multi-chambered, spindle-shaped galls on the leaf petiole, petiolules and rarely on young twigs (Shinji 1940; Miyatake 1996).</p><p>Distribution. Japan (Shinji 1940; Miyatake 1996), South Korea (according to Kwon &amp; Kwon 2020, but needs confirmation; see Cho et al. 2017).</p><p>Comment. We only examined material from Japan. Occurrences reported for South Korea were partly misidentification of Triozidus stackelbergi (see Cho et al. 2017; Kwon &amp; Kwon 2020). Kwon &amp; Kwon (2020) list this species as a pest on medicinal Eleutherococcus spp. in South Korea.</p><p>Genetic resources. Adult sequences of COI were all slightly divergent, three haplotypes: PQ817987– PQ817989.</p></div>	https://treatment.plazi.org/id/03D2A373FFD5FFA4FF26FE09FE9F2AF9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.	Liao, Yi-Chang, Inoue, Hiromitsu, Percy, Diana M. (2025): The psyllid genus Triozidus Li, 1994 stat. rev. sensu novo (Hemiptera: Psylloidea: Triozidae) in East Asia is redefined with the addition of two new species from Taiwan inducing galls on the leaflet petiolules of Eleutherococcus trifoliatus (Araliaceae). Zootaxa 5727 (1): 113-139, DOI: 10.11646/zootaxa.5727.1.8, URL: https://doi.org/10.11646/zootaxa.5727.1.8
03D2A373FFC8FFBFFF26F886FE9F2A85.text	03D2A373FFC8FFBFFF26F886FE9F2A85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triozidus yangorum Liao & Inoue & Percy 2025	<div><p>Triozidus yangorum Liao &amp; Percy sp. nov.</p><p>(Figs 1C–F, 12–15)</p><p>Type material. Holotype: TAIWAN • ♂; Taipei City, Shilin, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.59114&amp;materialsCitation.latitude=25.13511" title="Search Plazi for locations around (long 121.59114/lat 25.13511)">Guzhen</a>; 25°08’06.4”N, 121°35’28.1”E; 17 Jan 2023; Y. C. Liao leg.; Eleutherococcus trifoliatus; NCHU, dry mounted . Paratypes: TAIWAN • 15♂, 19♀, 15 immatures; same data as for holotype . 2♂, 2♀; same data as for holotype, but NHMB . 7 immatures; Nantou Co., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=121.17305&amp;materialsCitation.latitude=24.089167" title="Search Plazi for locations around (long 121.17305/lat 24.089167)">Meifeng</a>; 24°05’21”N, 121°10’23”E; 23 Sep. 2008; M. M. Yang leg.; Eleutherococcus trifoliatus; NCHU, dry, slide mounted and in ethanol . Other material examined: TAIWAN • 1♂, 14 immatures; Nantou Co., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.79718&amp;materialsCitation.latitude=23.66513" title="Search Plazi for locations around (long 120.79718/lat 23.66513)">Sitou Forest Recreation Area</a>; 23.66513 N, 120.79718 E; 1295 m; 26 Jan. 2010; D. M. Percy leg.; Eleutherococcus trifoliatus; DMPC, slide mounted .</p><p>Description. Adult (Figs 1C, 12–14). Coloration. Body color yellowish brown (Figs 1C, 12). Newly emerged individuals light green. Entire head and abdominal tergites mid-brown. Antennae yellow with apices of segments 4, 6 and 8 dark brown, and entire segments 9 and 10 black. Compound eyes dark brown. Ocelli orange. Legs brown. Forewing pale yellow, transparent, with dark brown patch around clavus. Hindwing transparent, except clavus dark brown.</p><p>Structure. Body large-sized, length from anterior head margin to tip of folded forewing 5.3–6.1 mm, female generally larger (Fig. 12); covered in short fine setae. Head (Figs 12, 13A) slightly narrower than thorax. Vertex width 1.8–2.0 x length, minutely pubescent. Genal processes prominent, length along mid-line 0.7-0.8 x vertex length, divergent, conical, blunt at apex, medium-long pubescence. Antenna (Figs 13B–C) length 1.8–1.9 x head width, antennal segment 3 approximately double length of segment 4, relative length of flagellar segments as 1.0: 0.5: 0.3: 0.3: 0.3: 0.3: 0.2: 0.2; two unequal terminal setae: longer seta 1.1 x, and shorter, truncate seta 0.2 x segment 10 length. Thorax moderately arched dorsally, minutely pubescent. Hind leg (Fig. 13F) metatibia length 1.0–1.1 x head width. Forewing broad (Fig. 13D), length 6.1–6.8 x head width, and 2.4–2.6 x width, widest slightly distal to the middle; wing apex acute; vein Rs moderately long, gradually curved to fore margin; vein M evenly curved with bifurcation at or slightly anterior to line connecting apices of veins Rs and Cu 1a; cell m 1 larger than cell cu 1; vein Cu 1a evenly curved, vein Cu 1b more or less straight; veins minutely pubescent. Hindwing (Fig. 13E) 0.6 x as long and 0.5 x as wide as forewing.</p><p>Male terminalia (Figs 14A–C). Proctiger short, with moderate posterior lobes reaching maximum extension in the apical half, covered in long setae except for basal third laterally (Fig. 14A). Subgenital plate subglobular, with medium long setae laterally and ventrally; posterior dorsal margin slightly convex. Paramere (Figs 14A–B) about as long as proctiger; in profile broader at base, sinuous, before narrowing to apex which is acute and directed inward and rearward; inner face beset with long setae, outer surface with shorter setae. Distal aedeagus segment (Fig. 14C) about as long as paramere (Figs 14A, 16), angled medially with thumb-shaped projections, apical portion inflated into a shallow hook with blunt apex; sclerotized end tube of ductus ejaculatorius short, slightly sinuous. Female terminalia (Fig. 14D) cuneate, short; proctiger dorsal margin slightly sinuate with medial depression and acute apex, only slightly longer than subgenital plate, with long setae in the apical half, anal ring length approximately one third proctiger length, consisting of a single row of pores. Subgenital plate, in profile, triangular, apex acute, beset with long hairs laterally and ventrally. Ovipositor dorsal valvulae cuneate, ventral valvulae straight with several shallow serrations apically.</p><p>Measurements (range, mean±SD) in mm (5 males, 5 females). Body length (including forewing) ♂ 5.38–5.81, 5.63±0.17; ♀ 5.25–6.06, 5.81±0.32. Head width ♂ 0.74–0.78, 0.76±0.02; ♀ 0.76–0.80, 0.77±0.02. Vertex width ♂ 0.41–0.43, 0.42±0.01; ♀ 0.38–0.43, 0.41±0.02 Vertex length ♂ 0.23–0.30, 0.26±0.03; ♀ 0.28–0.29, 0.28±0.01. Genal cone length ♂ 0.19–0.22, 0.20±0.01; ♀ 0.20–0.23, 0.21±0.01. Antenna length ♂ 1.40–1.50, 1.44±0.04; ♀ 1.33–1.35, 1.34±0.01. Metatibia length ♂ 0.80–0.85, 0.82±0.03; ♀ 0.73–0.88, 0.81±0.06. Forewing length ♂ 4.58–4.93, 4.76±0.20; ♀ 4.85–5.56, 5.18±0.38. Forewing width ♂ 1.89–1.98, 1.93±0.04; ♀ 1.96–2.19, 2.06±0.11. Aedeagus length 0.30 (n =1). Paramere length 0.30 (n =2). Proctiger length ♀ 0.45–0.50 (n =2). Subgenital plate length ♀ 0.46–0.50 (n =2).</p><p>Fifth instar immature (Figs 1D, 15). Coloration. General color yellow. Forewing pad and thorax pale brown. Body (Fig. 15A) length 1.3–1.4 x width. Body surface, including margin of head, wing pads (Fig. 15C), caudal plate and legs covered with medium dense, long simple setae. Antenna (Fig. 15E) relative length of antennal segments 3 to 8 as 1.0: 0.85: 0.3: 0.4: 0.3: 1.8. Legs (Figs 15A–B) as for generic description. Forewing pad length 2.8–3.2 x width, and 2.4–2.8 x antenna length. Caudal plate length 0.8–1.0 x width. Circumanal ring (Fig. 15D) heart-shaped (medial constriction mainly from anterior); width 0.3 x caudal plate width; outer ring mostly composed of a single row of elongate pores, with occasional convolutions and intermittent multi-pore sections; inner ring composed of a mixture of small and irregular pores.</p><p>Measurements (range, mean±SD) in mm (5 immatures). Body length 2.53–2.88, 2.70±0.15. Head width 0.80– 0.85, 0.81±0.02. Antenna length 0.53–0.61, 0.57±0.03. Metatibiotarsus length 0.71–0.80, 0.76±0.04. Forewing pad length 1.36–1.60, 1.48±0.11. Caudal plate length 0.90–1.25, 1.01±0.14. Caudal plate width 1.08–1.18, 1.16±0.06. Circumanal ring width 0.30–0.35, 0.32±0.02.</p><p>Etymology. The specific name honours both Prof. Chung-Tu Yang and Dr. Man-Miao Yang for their enormous contribution to the systematics of Psylloidea .</p><p>Distribution. Taiwan; only found in the central and northern part.</p><p>Host plant. Eleutherococcus trifoliatus ( Araliaceae).</p><p>Biology. Univoltine with adults emerging December to January. Immatures develop within large, fully enclosed round galls that are positioned just below the base of leaflets on the petiolules. Occasionally galls are seen on the leaflet base, but this seems to be a secondary oviposition preference. Each gall consists of a single gall chamber with usually a single immature (Fig. 1D). Galls are found individually or in aggregates of two to four galls per leaf, often with a gall on each of the three petiolules (Figs 1E–F). The host plant, E. trifoliatus, is found in montane zones (500 to 1,500 m elevation) as an understory forest plant with prickly stems and a sprawling or scandent habit.</p><p>Genetic resources. Adult and immature sequences were identical: annotated mitochondrial genome: Genbank PQ826428.</p></div>	https://treatment.plazi.org/id/03D2A373FFC8FFBFFF26F886FE9F2A85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Liao, Yi-Chang;Inoue, Hiromitsu;Percy, Diana M.	Liao, Yi-Chang, Inoue, Hiromitsu, Percy, Diana M. (2025): The psyllid genus Triozidus Li, 1994 stat. rev. sensu novo (Hemiptera: Psylloidea: Triozidae) in East Asia is redefined with the addition of two new species from Taiwan inducing galls on the leaflet petiolules of Eleutherococcus trifoliatus (Araliaceae). Zootaxa 5727 (1): 113-139, DOI: 10.11646/zootaxa.5727.1.8, URL: https://doi.org/10.11646/zootaxa.5727.1.8
