identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D7A111FFA6FFB5FA25D2E2FAA7FA4E.text	03D7A111FFA6FFB5FA25D2E2FAA7FA4E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syzygium bicolor Merr. & L. M. Perry	<div><p>2. Syzygium bicolor Merr. &amp; L.M.Perry — Fig. 1; Map 1</p><p>Syzygium bicolor Merr. &amp; L.M.Perry (1942) 286. — Type: Brass 13018 (holo A n.v.; iso BRI!, LAE!), Indonesia, Papua Province, Idenburg River, 6 km SW of Bernhard Camp, alt. 1200 m, Mar. 1939.</p><p>Tree to 30 m tall, to c. 60 cm dbh; bark reddish brown or grey, flaky or slightly rough. Vegetative branchlet terete, compressed or quadrangular; rounded, angled or weakly winged, 1–2.5 mm diam; bark dull, smooth to slightly striate; not glandular-verrucose, persistent. Leaf lamina 3–6 by 1.3–4 cm, 1.3–2.3 times as long as wide, elliptic to broadly elliptic; base cuneate, obtuse or narrowly cuneate; apex short acuminate, acuminate or acute; acumen flat or recurved; margin flat or revolute; cartilaginous; primary and secondary venation distinctly different with secondaries relatively little developed and not or rarely joining the intramarginal vein; primary veins 5–7 on each side of the mid-rib; in median part of the lamina at a divergence angle of 50–70° and 4–12 mm apart; intramarginal vein present, weakly or strongly arched, 1–4 mm from margin, secondary intramarginal vein present (rarely absent, in large leaves a tertiary intramarginal vein may be evident also). Petiole 2.5–5 mm long. Reproductive seasonal growth unit with a reproductive zone only. Inflorescence terminal, or terminal and distal axillary, few- to many-flowered, paniculate, up to 2– 5 by 1– 5 cm, major axis 1–1.5 mm thick at the midpoint, bark glandular-verrucose; bracts caducous or deciduous; bracteoles caducous. Flowers white, calyptrate (petals coherent and falling as a cap). Hypanthium dull, glandular-verrucose or striate (sometimes distinctly glandular-verrucose distally and striate for the remainder), visibly gland-dotted; stipitate; stipitately very narrowly obconic; 10–12 by 3–4 mm; stipe 2–4.5 mm long. Calyx lobes 4 or 5, transversely semielliptic or semicircular, 1 mm long. Petals 4–6, coherent and caducous, 3–3.5 mm long. Staminal disc flat (Fig. 1: 1.4, 1.5). Stamens numerous, 6–8.5 mm long. Style 6–8.5 mm long. Placentation axile-median; placenta oblong, narrowly oblong or narrowly oblong-triangular, the 2 distal lobes appressed, or subobtriangular and proximally rounded with the 2 distal lobes well separated. Ovules c. 12–22 per locule, ascending, arranged in two longitudinal rows (one row on each placental lobe). Fruit not seen.</p><p>Distribution — Indonesia (Papua), Papua New Guinea.</p><p>Habitat &amp; Ecology — Rainforest, moist mid-mountain forest. Altitude 1200–1800 m.</p><p>Note — Flowers occur in groups of 2, 3, 4 or 6 (rarely singly) on the terminal internodes and often the axis apex is ‘knobbly’ with two or more insertion points for flowers, perhaps indicating that a reduction in the number of internodes has occurred in the inflorescence. It may be that axis reduction has been a feature of inflorescence evolution in this species.</p></div>	https://treatment.plazi.org/id/03D7A111FFA6FFB5FA25D2E2FAA7FA4E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Craven, L. A.	Craven, L. A. (2019): Studies in Papuasian Syzygium (Myrtaceae): 1. Subgenus Perikion revised. Blumea 64 (2): 115-122, DOI: 10.3767/blumea.2019.64.02.03, URL: https://doi.org/10.3767/blumea.2019.64.02.03
03D7A111FFA6FFB4F96AD13AFE89FDC1.text	03D7A111FFA6FFB4F96AD13AFE89FDC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syzygium carrii T. G. Hartley & L. M. Perry	<div><p>3. Syzygium carrii T.G.Hartley &amp; L.M.Perry — Fig. 1; Map 1</p><p>Syzygium carrii T.G.Hartley &amp; L.M.Perry (1973) 209. — Type: Carr 13511 (holo A n.v.; iso CANB,LAE), Papua New Guinea, Central Province, Boridi, forest, c. 1525 m, 25 Nov. 1935.</p><p>Tree to c. 30 m tall. Vegetative branchlet quadrangular, winged, 0.7–1 mm diam; bark dull, smooth, slightly glandular-verrucose, persistent. Leaf lamina 1.5–2.8 by 0.7–1.7 cm, 1.9–2.1 times as long as wide, elliptic or obovate; base cuneate; apex acute or obtuse; margin revolute; coriaceous; primary and secondary venation distinctly different with secondaries relatively little developed and not or rarely joining the intramarginal vein (few secondary veins are present); primary veins 8–10 on each side of the mid-rib; in median part of the lamina at a divergence angle of 60–70° and 1–3 mm apart; intramarginal vein present, weakly arched, 0.5–1 mm from margin, secondary intramarginal vein absent. Petiole 0.7–1 mm long. Reproductive seasonal growth unit with a reproductive zone only. Inflorescence terminal to median axillary; 1- to few-flowered, racemose or paniculate, up to 1–2 by 0.2–0.7 cm wide, major axis 0.3–0.5 mm thick at the midpoint; bracts caducous or deciduous; bracteoles subtending each flower; deciduous. Flowers possibly calyptrate (petals coherent and probably falling as a cap). Hypanthium dull, glandular-verrucose, visibly gland-dotted; stipitate; stipitately very narrowly obconic or elongated goblet-shaped; 8 by 2 mm; stipe 2 mm long. Calyx lobes 4; transversely narrowly semielliptic, c. 0.5 mm long. Placentation axile-median. Ovules c. 10 per locule, pendulous, arranged in two longitudinal rows (one row on each placental lobe). Staminal disc and fruit not seen (see note).</p><p>Distribution — Papua New Guinea.</p><p>Habitat &amp; Ecology — Forest. Altitude 1520 m.</p><p>Note — This species is known only from one collection in late bud. The buds are not in good condition with some tissue breakdown evident, making observation difficult for determining some characters.</p></div>	https://treatment.plazi.org/id/03D7A111FFA6FFB4F96AD13AFE89FDC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Craven, L. A.	Craven, L. A. (2019): Studies in Papuasian Syzygium (Myrtaceae): 1. Subgenus Perikion revised. Blumea 64 (2): 115-122, DOI: 10.3767/blumea.2019.64.02.03, URL: https://doi.org/10.3767/blumea.2019.64.02.03
03D7A111FFA7FFB3FA25D58AFEB6FD83.text	03D7A111FFA7FFB3FA25D58AFEB6FD83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syzygium claviflorum (Roxb.) Steud.	<div><p>4. Syzygium claviflorum (Roxb.) Steud. — Fig. 1; Map 2</p><p>Syzygium claviflorum (Roxb.) Steud. (1841) 657. — Eugenia claviflora Roxb. (1832) 488. — Acmenosperma claviflorum (Roxb.) Kausel (1957) 609. — Type: Icones Roxburghianae no. 2499 (lecto K, selected by Soh &amp; Parnell 2015).</p><p>Eugenia leptantha Wight (1841) 14. — Syzygium leptanthum Nied. (1893) 85. — Eugenia leptalea Craib (1931) 649. — Eugenia claviflora Roxb. var. leptalea (Craib) M.R.Hend. (1949) 255. — Type: Griffith (holo, n.v.), Burma, Mergui.</p><p>Tree (sometimes flowering as a treelet) to 28 m tall, to 60 cm dbh; bark brown, dark brown, grey-brown, brownish, light-grey, creamy-grey, mottled grey, light fawn. Vegetative branchlet terete or compressed, rounded or angled, 0.8–2.1 mm diam; bark dull, smooth or cracked (occasionally slightly striate), not glandular-verrucose, flaking in relatively large pieces. Leaf lamina 5–19.3 by 1.6–6.6 cm, 1.6–3.9 times as long as wide, narrowly elliptic, elliptic, ovate, narrowly ovate, narrowly obovate or obovate; base narrowly cuneate, cuneate, obtuse or attenuate (sometimes approaching rounded); apex long acuminate or acuminate; acumen recurved (or occasionally flat); margin flat (or occasionally slightly revolute, often appears undulate but this may be an artefact of drying); primary and secondary venation distinctly different with secondaries relatively little developed and not or rarely joining the intramarginal vein; primary veins 12–27(–37) on each side of the mid-rib, in median part of the lamina at a divergence angle of 60–70° and 2–10 mm apart; intramarginal vein present, 0.5–3 mm from margin, secondary intramarginal vein present (sometimes difficult to discern). Petiole 2–7 mm long. Reproductive seasonal growth unit with a reproductive zone only. Inflorescence among the leaves or below the leaves and then ramuline or ramine, rarely terminal, 2- to many-flowered, spicate, umbellate or paniculate, up to 1.5–5 by 2–6 cm, major axis 0.3–1.8 mm thick at the midpoint, bark smooth or commonly granular-papillate; bracts deciduous; bracteoles subtending each flower, caducous. Flowers white, cream or green, calyptrate or not (petals coherent and falling as a cap or discrete and falling individually). Hypanthium dull, visibly gland-dotted, minutely (but distinctly) wrinkled; stipitate; elongated goblet-shaped or extremely narrowly funnel-shaped; 9.8–23 by 1.7–5.6 mm; stipe 6.5–17 mm long. Calyx lobes 4 or 5, very depressedly triangular, transversely narrowly semielliptic or transversely oblong, 0.3–0.8 mm long. Petals 6–8, coherent and then caducous and falling as a cap or deciduous and then falling individually, 2–3 mm long. Staminal disc flat (Fig. 1: 1.1). Stamens numerous, 3–8 mm long. Style 3–8 mm long. Placentation axile-median; placenta distinctly lobed, narrowly semiobovoid to semilinear-obovoid. Ovules c. 9–16 per locule, pendulous, arranged in two longitudinal rows (one row on each placental lobe). Mature fruit red, ellipsoid or very broadly obovoid and flat at the apex, 12–14 by 9–10 mm (excluding the calyx), the hypanthium rim not appreciably expanding in fruit; seed 3 mm across; cotyledons interlocked by an intrusive weakly ramifying tissue.</p><p>Distribution — Indonesia (Aru Islands, Papua Barat, Papua), Papua New Guinea.</p><p>Habitat &amp; Ecology — Oak forest, rainforest, swamp forest, edge of mangroves and riparian forest, riverine forest, lowland hill forest, tall open forest, forest on low stony hills, oak-beech forest. Altitude 0–2750 m.</p><p>Notes — A broad circumscription of S. claviflorum is taken in the present treatment. The species occurs from India and southern China southwards through Malesia to north-eastern Australia and it merits a detailed study throughout its range. I suspect that several distinct biological entities are included in the present circumscription. One specimen, Pleyte 1075, has leaves up to 35 by 11.4 cm wide but its flowers and inflorescences conform to variation within the species. It was recorded as being a shrub of 2 m tall. In Papuasia, flowers are seen to become consistently larger and leaves smaller in specimens from higher altitudes. In addition to being good-sized forest trees, plants may also be treelets or shrubs .</p><p>In S. claviflorum and the species most closely allied to it (e.g., S. suberosum), the hypanthium could be interpreted as being not stipitate (i.e., the hypanthium uniformly tapers from base to apex) but there is a point at which the parenchymatous tissue in the ovarian region stops and a darker-coloured proximal tissue begins. The darker tissue is interpreted as being the stipe. Syzygium claviflorum is similar to S. leptopodium but in general it differs in having larger leaves, a cuneate vs an often ± rounded leaf base, subcoriaceous vs chartaceous leaves (in montane material), larger flowers, more flowers per inflorescence, a tendency towards a lateral rather than terminal inflorescence, rounded or compressed branchlets vs ± tetragonous branchlets. These features all more or less break down. Anther size, however, appears to distinguish flowering material, the anthers being about 0.6–0.8 mm long in S. claviflorum and 0.2–0.4 mm long in S. leptopodium . Also, S. claviflorum usually has glandular, ± twisted staminal filaments; those in S. leptopodium are eglandular and ± straight.</p><p>The style articulates about 1 mm above its base, leaving a stub in mature fruit. Fruit sometimes develops without seeds, and then lacks the characteristic shape of fertile fruit, being narrowly obovoid.</p></div>	https://treatment.plazi.org/id/03D7A111FFA7FFB3FA25D58AFEB6FD83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Craven, L. A.	Craven, L. A. (2019): Studies in Papuasian Syzygium (Myrtaceae): 1. Subgenus Perikion revised. Blumea 64 (2): 115-122, DOI: 10.3767/blumea.2019.64.02.03, URL: https://doi.org/10.3767/blumea.2019.64.02.03
03D7A111FFA0FFB3FA25D54FFCF9FCB1.text	03D7A111FFA0FFB3FA25D54FFCF9FCB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syzygium kokomo Craven 2019	<div><p>5. Syzygium kokomo Craven, sp. nov. — Fig. 1; Map 3</p><p>From Syzygium claviflorum (Roxb.) Steud. it differs in the quadrangular vegetative branchlets; the flat leaf lamina acumen; and the stipitately very narrowly obconic hypanthium with a stipe 6–7.5 mm long. (In S. claviflorum the vegetative branchlets are terete or compressed, the leaf lamina acumen is recurved,and the hypanthium is funnel-shaped or elongated goblet-shaped with a stipe 6.5–17 mm long.) — Type: Henty &amp; Foreman NGF 42673 (holo L!; iso A!, BISH, BO, BRI, CANB, K, LAE, NSW,SING, all n.v.), Papua New Guinea, West Sepik (Sandaun) Province, Telefomin Subprovince, Kokomo Creek (a tributary of Frieda River), ridge forest, alt. c. 670 m, 28 June 1969.</p><p>Etymology. The specific epithet is a noun in apposition derived from the locality Kokomo Creek.</p><p>Tree to 27 m tall, to 25 cm dbh; bark dark brown outside, red brown inside. Vegetative branchlet quadrangular initially but at length terete, winged or angled, 0.8–1.4 mm diam; bark dull, smooth or cracked (cracking is fine and longitudinal), not glandular-verrucose, persistent or peeling in relatively thin strips. Leaf lamina 3.5–5 by 1.3–1.8 cm, 2.5–3.9 times as long as wide, narrowly ovate or ovate; base cuneate or narrowly cuneate; apex long acuminate, acumen flat and up to 1.5 cm long; margin flat; thickly coriaceous, approaching cartilaginous; primary and secondary venation distinctly different with secondaries relatively little developed and not or rarely joining the intramarginal vein (in some cases not even the primary veins reach the intramarginal vein); primary veins 3–6 on each side of the mid-rib, in median part of the lamina at a divergence angle of 40–60° and 3–6 mm apart; intramarginal vein present, weakly arched, 0.8–1 mm from leaf margin, secondary intramarginal vein present or absent. Petiole 3.5–5 mm long. Reproductive seasonal growth unit with a reproductive zone only. Inflorescence terminal, or distal or median axillary, few- to many-flowered, paniculate up to 1–3.5 by 1–2.5 cm, major axis 0.5–0.7 mm thick at the midpoint, bark glandular-verrucose; bracts deciduous; bracteoles apparently subtending each flow- er, deciduous (rarely a few persisting). Flowers (immature) with free calyx lobes . Hypanthium dull, smooth; stipitate; stipitately very narrowly obconic; 6–7.5 by 2 mm. Calyx lobes 5. Petals probably 5. Staminal disc, stamens and fruit not seen (see note).</p><p>Distribution — Papua New Guinea.</p><p>Habitat &amp; Ecology — Ridge forest . Altitude 670 m.</p><p>Notes — This species is known from a single collection only, this being in the bud stage.</p><p>The inflorescence is a simple paniculate structure, often with 2 pairs of lateral branches and then a terminal cluster of flowers at the apex of the final internode of the main axis. Flowers may terminate axes in groups of 3 or 7 (the latter presumably being 2 triads and 1 monad). Occasionally, perhaps through abortion, monads or dyads occur. There can be up to 7 inflorescences at the apex of a vegetative seasonal growth unit.</p><p>The figure of 6–7.5 mm given above for the hypanthium length is the total length of the buds at what is possibly the mid-bud stage.</p></div>	https://treatment.plazi.org/id/03D7A111FFA0FFB3FA25D54FFCF9FCB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Craven, L. A.	Craven, L. A. (2019): Studies in Papuasian Syzygium (Myrtaceae): 1. Subgenus Perikion revised. Blumea 64 (2): 115-122, DOI: 10.3767/blumea.2019.64.02.03, URL: https://doi.org/10.3767/blumea.2019.64.02.03
03D7A111FFA0FFB2F96AD419FF37F9A9.text	03D7A111FFA0FFB2F96AD419FF37F9A9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syzygium leptopodium Merr. & L. M. Perry	<div><p>6. Syzygium leptopodium Merr. &amp; L.M.Perry — Fig. 1; Map 3</p><p>Syzygium leptopodium Merr.&amp; L.M.Perry (1942) 284. — Type: Clemens 5326 (holo A n.v.; iso L!), Papua New Guinea, Morobe Province, Ogeramnang, forest, c. 1770–1830 m, 8 Feb. 1937.</p><p>Treelet, shrub or tree, usually to 8 m tall (once recorded as being c. 27 m tall), to 8 cm dbh. Vegetative branchlet terete, compressed or quadrangular, rounded or winged (if present, wings are narrow), 0.8–1.5 mm diam; bark dull, bark smooth or cracked (cracking is fine and approaching striate), not or only slightly glandular-verrucose, persistent or peeling in relatively thin strips. Leaf lamina (2–)4–7(–8.5) by (1.2–)1.7–2.4(–4) cm, 2–3.2 times as long as wide, narrowly elliptic, elliptic, narrowly ovate or ovate; base narrowly cuneate, cuneate, obtuse or occasionally rounded; apex long or short acuminate or acute (rarely abruptly acuminate); acumen flat; margin flat or slightly; chartaceous or coriaceous; primary and secondary venation distinctly different with secondaries relatively little developed and not or rarely joining the intramarginal vein; primary veins 6–17 on each side of the mid-rib, in median part of the lamina at a divergence angle of 50–90°, (1–)2–3(–5) mm apart; intramarginal vein present, weakly or strongly arched, (0.3–)0.5–1(–2) mm from margin; secondary intramarginal vein often present at the margin but not strongly developed. Petiole 0.5–4 mm long. Reproductive seasonal growth unit with a reproductive zone only. Inflorescence terminal, distal or median axillary, or lateral below the leaves (ramuline or ramine), 1- or 2- to few-flowered, paniculate or racemose up to 1.2–3.5 by 0.2–3 cm, major axis 0.4–0.6 mm thick at the midpoint; bark glandular-verrucose or smooth; bracts deciduous or caducous; bracteoles sometimes present but apparently not in all flowers, caducous (rarely a few persisting to anthesis). Flowers white, cream or mauve (once described as whitish lilac), calyptrate (petals coherent and falling as a cap). Hypanthium dull, glandular-verrucose, visibly gland-dotted, plane or weakly ribbed; stipitate; elongated goblet-shaped, very elongatedly stipitate-bowl-shaped, or thickly narrowly obconic (occasionally very narrowly obconic); 7.5–14.5 by 2–3 mm wide; stipe 1–3 mm long. Calyx lobes 2 –4, depressedly triangular, very depressedly triangular, oblong or triangular, (0.2–)0.3–0.5(–0.7) mm long. Petals c. 4–7; coherent and caducous, c. 1.5–2.3 mm long. Staminal disc raised (Fig. 1: 4.2). Stamens numerous, 1.75–7 mm long. Style 4–7 mm long. Placentation axile-median; placenta more or less flattened, narrowly obovoid. Ovules c. 6–12 per locule, pendulous, arranged in two longitudinal rows (one row on each placental lobe). Mature fruit wine red, orange red, dark or red, magenta, maroon or purplish black, glandular-verrucose, plane or weakly ribbed, subpyriform or ellipsoid, 10–12 by 6–9 mm (excluding the calyx), the hypanthium rim not appreciably expanding in fruit; seed a spheroid or broadly obovoid and deeply impressed at the apex, 5.5–6 mm across; cotyledons interlocked by an intrusive weakly ramifying tissue, collateral or superposed.</p><p>Distribution — Indonesia (Papua Barat, Papua); Papua New Guinea.</p><p>Habitat &amp; Ecology — Mossy mid-montane forest, montane rainforest with Castanopsis, oak forest, mixed degraded rainforest, forest remnant in river gorge, secondary forest, lower montane rainforest, Nothofagus forest, mossy fagaceous forest, primary forest, edge of Barringtonia - Leptospermum swamp, rocky open situation. Altitude 50–2200 m.</p><p>Notes — See under S. claviflorum for notes on differences between that species and S. leptopodium .</p><p>The basic floral units within the inflorescence are monads and triads. Sometimes an inflorescence consists of a single monad, or an axis with 2 proximal monads on short axes then 2 sessile monads with a terminal triad or 5-flowered cluster. Often it consists of an axis with 2 sterile monads and the terminal 3- or 5-flowered unit, or it may be a sessile 3- or 5-flowered unit without any peduncle.</p><p>Flowers with only 2 calyx lobes were observed in Van Royen 4945.</p></div>	https://treatment.plazi.org/id/03D7A111FFA0FFB2F96AD419FF37F9A9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Craven, L. A.	Craven, L. A. (2019): Studies in Papuasian Syzygium (Myrtaceae): 1. Subgenus Perikion revised. Blumea 64 (2): 115-122, DOI: 10.3767/blumea.2019.64.02.03, URL: https://doi.org/10.3767/blumea.2019.64.02.03
03D7A111FFA1FFB1F96AD784FF08FE6A.text	03D7A111FFA1FFB1F96AD784FF08FE6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syzygium saundersii Craven 2019	<div><p>7. Syzygium saundersii Craven, sp. nov. — Fig. 1; Map 4</p><p>From Syzygium claviflorum (Roxb.) Steud. it differs in the obtuse, rounded or truncate leaf lamina apex; the glossy, non-stipitate and 8–10 mm long hypanthium; the 7–11 mm long stamens; the 5–5.5 mm long style; and the c. 10 ovules per locule. (In S. claviflorum the leaf lamina apex is long acuminate or acuminate,the hypanthium is dull, stipitate and 9.8–23 mm long,the stamens are 3–8 mm long, and there are 9–16 ovules per locule.) — Type: Saunders 663 (holo CANB!; iso A, BISH, BM, BO, BRI, G, K, L, LAE, MEL, US, all n.v.), Papua New Guinea, Western Highlands Province, c. 800 m S of Tomba, forest, alt. c. 2440 m, 2 July 1957.</p><p>Etymology. The species is named in honour of John Campbell Saunders (1930–2001), forest botanist in the CSIRO Division of Land Research &amp; Regional Survey’s New Guinea survey team and its successors from 1954 to 1995. John’s herbarium collections are sometimes scanty as they were collected as vouchers for his many wood samples; the type of the present species is a notable exception.He encouraged CSIRO herbarium collectors such as R.D. Hoogland,R. Pullen,R. Schodde and LAC to also collect wood samples and is directly responsible for the significant CSIRO contribution to Papua New Guinean wood samples,deposited primarily in FPAw and PMPw.</p><p>Tree to 26 m tall (bole to 15 m), to 45 cm dbh; bark grey. Vegetative branchlet quadrangular to compressed, angled to slightly winged, 2– 6 mm diam; bark dull, smooth, not glandular-verrucose, persistent. Leaf lamina 4.5–13 by 2.7–8 cm, 1.3–1.7 times as long as wide, broadly obovate, obovate or broadly elliptic; base attenuate or cuneate; apex truncate, rounded or obtuse; margin revolute; cartilaginous; primary and secondary venation distinctly different with secondaries relatively little developed and not or rarely joining the intramarginal vein (secondary veins are few); primary veins 10–16 on each side of the mid-rib, in median part of the lamina at a divergence angle of 65–75°, 3–10 mm apart; intramarginal vein present, weakly arched, 1.5–3 mm from margin, secondary intramarginal vein absent. Petiole 12–27 mm long. Reproductive seasonal growth unit with distinct vegetative and reproductive zones, or with a reproductive zone only (the vegetative zone is weakly developed and consists at most of a few pairs of leaves (that often are reduced in size). Inflorescence terminal or distal axillary, few- to many-flowered, paniculate, up to 3.5–12 by 2.5–9 cm, major axis 2–3 mm thick at the midpoint; bark smooth; bracts deciduous; bracteoles subtending each flower, caducous. Flowers calyptrate (petals coherent and falling as a cap). Hypanthium glossy, smooth (in sicco strongly wrinkled but this presumably is an artefact of drying), visibly but not strongly gland-dotted; not stipitate (tapering evenly to the base or truncate or rounded); thickly narrowly obconic; 8–10 by 3.5–4.5 mm. Calyx lobes 4, transversely semielliptic or semielliptic, 1–2 mm long. Petals 4; caducous (falling at anthesis); coherent (outer petal apparently falls separately but inner 3 seemingly fall as a unit), 5.5 mm long. Staminal disc flat (Fig. 1: 1.9). Stamens c. 55, 7–11 mm long. Style 5–5.5 mm long. Placentation axile-median; placenta apparently more or less oblong and not very prominent. Ovules c. 10 per locule, ascending, arranged in two longitudinal rows (one row on each placental lobe) or possibly arranged irregu- larly. Fruit not seen.</p><p>Distribution — Papua New Guinea.</p><p>Habitat &amp; Ecology — Forest, mountain forest on steep slope, mixed montane forest. Altitude 2440–2600 m.</p><p>Note — The relationships of this species may lie with S. bicolor .</p></div>	https://treatment.plazi.org/id/03D7A111FFA1FFB1F96AD784FF08FE6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Craven, L. A.	Craven, L. A. (2019): Studies in Papuasian Syzygium (Myrtaceae): 1. Subgenus Perikion revised. Blumea 64 (2): 115-122, DOI: 10.3767/blumea.2019.64.02.03, URL: https://doi.org/10.3767/blumea.2019.64.02.03
03D7A111FFA2FFB1FA25D527FEE0F7E6.text	03D7A111FFA2FFB1FA25D527FEE0F7E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syzygium sleumeri Craven 2019	<div><p>8. Syzygium sleumeri Craven, sp. nov. — Fig. 1; Map 4</p><p>From Syzygium claviflorum (Roxb.) Steud. it differs in the winged branchlets; the obtuse, rounded or retuse leaf lamina apex; the glandular-verrucose, ribbed, and 10 mm long hypanthium; the 6–9 mm long stamens; and 8–10 ovules per locule. (In S. claviflorum the branchlets are rounded or angled; the leaf lamina apex is long acuminate or acuminate; the hypanthium is minutely wrinkled and generally plane, and 9.8–23 mm long; the stamens are minutely wrinkled, generally plane, and 3–8 mm long; and there are 9–16 ovules per locule.) — Type: Sleumer &amp; Vink BW 14247 (holo CANB!; iso L n.v., MAN n.v.), Indonesia, Papua Barat, Mt Gwamongga,Anggi Gigi Lake, fire-vegetation of Ericaceae and Baeckea on clay, alt. 2250 m, 21 Jan. 1962.</p><p>Etymology. The species is named in honour of Hermann Otto Sleumer (1906–1993), a prolific author on the taxonomy of several plant families and an expert in Ericaceae in particular.</p><p>Shrub to 2 m tall. Vegetative branchlet quadrangular, winged, 0.6–1.3 mm diam; bark dull-glossy, smooth, not glandular-verrucose, persistent. Leaf lamina 1.5–3.5 by 0.7–1.6 cm, 1.6–2.4 times as long as wide, elliptic, narrowly elliptic, broadly elliptic or obovate; base cuneate or narrowly cuneate; apex obtuse, rounded or occasionally retuse; margin flat to revolute; cartilaginous; primary and secondary venation obscure; primary veins 5–8 on each side of the mid-rib, in median part of the lamina at a divergence angle of c. 60°, 1–3 mm apart; intramarginal vein present (often obscure), weakly arched, 0.2–0.5 mm from margin. Petiole 0.5–0.7 mm long. Reproductive seasonal growth unit with a reproductive zone only. Inflorescence leafless, terminal or distal axillary (sometimes terminal and distal axillary), 1- to few-flowered, racemose, up to 1–3 by 1.5–2 cm, major axis 0.4–0.5 mm thick at the midpoint; bark glandular-verrucose; bracts caducous or deciduous, bracteoles subtending each flower; deciduous. Flowers white (noted as calyx lobes red, petals and filaments white), calyptrate (petals coherent and falling as a cap). Hypanthium dull, glandular-verrucose, visibly gland-dotted, weakly ribbed; stipitate; generally elongated goblet-shaped (at the apex curved slightly), 10 by 3–3.3 mm wide; stipe 2.5–3 mm long. Calyx lobes 4; transversely narrowly semielliptic, 0.5 mm long. Petals 7, coherent and caducous, 2.3 mm long. Staminal disc flat (intermediate between Fig. 1: 1.2, 1.6). Stamens c. 75, 6–9 mm long. Style 8.5 mm long. Placentation axile-median; placenta is an obscure, flattened and narrowly obovate structure (bilobed in the distal part). Ovules c. 8–10 per locule, pendulous, arranged in two longitudinal rows (one row on each placental lobe). Fruit not seen.</p><p>Distribution — Indonesia (Papua).</p><p>Habitat &amp; Ecology — Fire-vegetation of Ericaceae and Baeckea, steppe. Altitude 50–2250 m.</p><p>Additional specimen examined. INDONESIA, Papua, Misool, Sorong, between Fakal and Tip, Pleyte 897.</p><p>Note — The shrub habit is characteristic, if not diagnostic. The inflorescence is a racemose structure, 1- to c. 6-flowered, terminal and/or distal axillary. The flowers are in monads or triads, or sometimes clustered in 4’s and then always at the apex of the inflorescence.</p></div>	https://treatment.plazi.org/id/03D7A111FFA2FFB1FA25D527FEE0F7E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Craven, L. A.	Craven, L. A. (2019): Studies in Papuasian Syzygium (Myrtaceae): 1. Subgenus Perikion revised. Blumea 64 (2): 115-122, DOI: 10.3767/blumea.2019.64.02.03, URL: https://doi.org/10.3767/blumea.2019.64.02.03
03D7A111FFA2FFB1F96AD784FA8BFB69.text	03D7A111FFA2FFB1F96AD784FA8BFB69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syzygium subamplexicaule Merr. & L. M. Perry	<div><p>9. Syzygium subamplexicaule Merr. &amp; L.M.Perry — Fig. 1; Map 4</p><p>Syzygium subamplexicaule Merr. &amp; L.M.Perry (1942) 285. — Type: Brass 8218 (holo A n.v.; iso BRI!), Papua New Guinea, Western (Fly River) Province, Lower Fly River, E bank opposite Sturt Island, Oct. 1936.</p><p>Treelet to 2.5 m tall. Vegetative branchlet terete, rounded, 2.5 mm diam; bark dull-glossy, smooth, not glandular-verrucose, persistent. Leaf lamina 22–27 by 5–9 cm wide, 3.5 times as long as wide, narrowly ovate to narrowly oblong-ovate; base cordate or truncate; apex acuminate; margin flat; coriaceous (chartaceous fide M&amp;P), primary and secondary venation distinctly different with secondaries relatively little developed and not or rarely joining the intramarginal vein; primary veins 22–25 on each side of the mid-rib, in median part of the lamina at a divergence angle of 70–80°, 8–20 mm apart; intramarginal vein present, weakly arched, 2.5–4 mm from margin, secondary intramarginal vein absent. Petiole 1.5–3 mm long. Reproductive seasonal growth unit with a reproductive zone only. Inflorescence terminal, 2- to few-flowered, paniculate, up to c. 2 cm long, major axis 1 mm thick at the midpoint, bark smooth; bracts deciduous to persistent; bracteoles absent from each flower in the unit. Hypanthium dull, glandular-verrucose, visibly gland-dotted, ribbed; stipitate. Calyx lobes 4, triangular, 0.75–1 mm long. Staminal disc ascending (Fig. 1: 3.2). Stamens 4–6 mm long (fide M&amp;P). Mature fruit pink, glandular-verrucose, ribbed, stipitately narrowly ellipsoid, 14–17 by 6–7.5 mm wide (excluding the calyx), the hypanthium rim not appreciably expanding in fruit; seed narrowly ellipsoid-obovoid and deeply impressed at the apex, 3–5 mm across; cotyledons interlocked by an intrusive weakly ramifying tissue, collateral.</p><p>Distribution — Papua New Guinea.</p><p>Habitat &amp; Ecology — Rainforest. Altitude 10 m.</p><p>Note — The BRI sheet of the type collection is the only material seen. Flowers have not been seen and the data presented here includes information in Merrill &amp; Perry’s (1942) description. The species is closely related to S. claviflorum .</p></div>	https://treatment.plazi.org/id/03D7A111FFA2FFB1F96AD784FA8BFB69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Craven, L. A.	Craven, L. A. (2019): Studies in Papuasian Syzygium (Myrtaceae): 1. Subgenus Perikion revised. Blumea 64 (2): 115-122, DOI: 10.3767/blumea.2019.64.02.03, URL: https://doi.org/10.3767/blumea.2019.64.02.03
03D7A111FFA2FFB0F96BD227FEA1F9D6.text	03D7A111FFA2FFB0F96BD227FEA1F9D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Syzygium suberosum Craven 2019	<div><p>10. Syzygium suberosum Craven, sp. nov. — Fig. 1; Map 4</p><p>From Syzygium claviflorum (Roxb.) Steud. it differs in the often corky branchlets; the persistent calyx lobes; and the strongly ramifying chalazal tissue in the embryo. (In S. claviflorum the branchlets are always smooth; the calyx lobes are deciduous;and the chalazal tissue in the embryo is weakly ramifying.) — Type: Van der Sijde BW 5565 (holo CANB!; iso L n.v., MAN n.v.), Indonesia, Papua Barat, Sidei (c. 50 km W of Manokwari), primary forest, alt. 50 m, 28 Mar. 1958.</p><p>Etymology. The specific epithet is derived from the Latin suber, cork, in reference to the corky branchlet bark.</p><p>Shrub, treelet or tree; to 7 m tall; bark grey, fissured and flaking. Vegetative branchlet terete or quadrangular, rounded, or winged, 1.5–5 mm diam; bark dull, smooth and corky, not glandular-verrucose, persistent. Leaf lamina 8.5–19 by 2.5–7.4 cm, 2.6–4 times as long as wide, narrowly elliptic or elliptic; base cuneate, obtuse, narrowly cuneate or attenuate; apex long acuminate or acuminate; acumen flat or recurved; margin flat; chartaceous to coriaceous; primary and secondary venation distinctly different with secondaries relatively little developed and not or rarely joining the intramarginal vein; primary veins 10–17 on each side of the mid-rib, in median part of the lamina at a divergence angle of 60–70°, 4–17 mm apart; intramarginal vein present, weakly arched, 1.5–8 mm from margin, secondary intramarginal vein present or absent. Petiole 3–5 mm long. Reproductive seasonal growth unit with a reproductive zone only. Inflorescence leafless, median axillary or lateral (ramuline), 1- to few-flowered, cymose, up to 2 by 0.5–2.5 cm wide (0.5 cm when 1-flowered), major axis 1 mm thick at the midpoint, bark smooth; bracts persistent; bracteoles subtending each flower, persistent to deciduous. Flowers white, calyptrate (petals coherent and falling as a cap). Hypanthium dull, glandular-verrucose to smooth, visibly gland-dotted; stipitate; stipitately very narrowly obconic, or elongated-goblet-shaped to very narrowly elongated-goblet-shaped, 18–21 by 2.5–3 mm wide; stipe 6–8 mm long. Calyx lobes 5 (often irregular in size and sometimes nearly obsolete), transversely semielliptic or transversely narrowly semielliptic, 0.3–1 mm long, persistent. Petals c. 8, coherent and caducous, up to 2 mm long. Staminal disc flat (Fig. 1: 1.10). Stamens numerous, 9–10 mm long. Style c. 10.5 mm long. Placentation axile-median; placenta more or less linear and flattened. Ovules c. 10 per locule, pendulous, arranged in two longitudinal rows (one row on each placental lobe). Fruit red, yellow or orange (probably mature (but galled) fruit recorded as red, youngish fruit recorded as yellow-orange), glandular-verrucose, finely ribbed, very narrowly obconic or very narrowly pyriform, 20–28 mm by 5 mm wide (excluding the calyx), the hypanthium rim not appreciably expanding in fruit; seed narrowly obovoid and deeply impressed at the apex, 3 mm across; cotyledons interlocked by an intrusive strongly ramifying tissue, collateral.</p><p>Distribution — Indonesia (Papua Barat).</p><p>Habitat &amp; Ecology — Oak-dominated primary forest, primary forest. Altitude 5–50 m.</p><p>Note — The inflorescence is a short, cymose structure consisting of monads and/or triads. The smallest inflorescence seen consisted of a single monad, and the largest comprised two lateral monads and a terminal triad. The shape of the staminal disc is very distinctive.</p><p>Acknowledgments Studies of Papuasian Syzygium at CANB were commenced by Tom Hartley and LAC in the 1970s. It is a matter of regret they could not be pursued to fruition at that time;two papers only being produced. Tom’s enthusiasm for the genus and his excellent field knowledge then assisted the project considerably in refining species concepts; these have been further refined in recent years by LAC as the work has progressed. Cath Busby obtained geocode data for those collections for which the data was not recorded on labels. Nunzio Knerr generated the distribution maps ready for final editing. Siobhan Duffy, in good humour, printed numerous digital images of type specimens for study, prepared the staminal disc diagrams from LAC’s crude drawings, and prepared the final versions of the distribution maps. The curators and/or directors of the following herbaria have made available specimens and/or images that have been used in my ongoing studies of Papuasian Syzygium and I sincerely thank them for this assistance:A, B, BISH, BM, BO, BRI, CANB, E, K, L, LAE, LY, MEL, NY, P, WRSL.The facilitation of loans, provision of digital images,and/or examina- tion of specimens have been of great benefit to the research on Syzygium . Those who have assisted me variously in the foregoing and other matters include, in no particular sequence, Emily Wood, Melinda Peters, Wayne Takeuchi, Eve Lucas, Ana Claudia Araujo, Shelley James, Frédéric Danet, Gaëtan Guignard, Mélanie Thiébaut, Jan-Frits Veldkamp,Wim Vink, Gerard Thijsse,Nicolien Sol, Kirsten Cowley,Thomas Zanoni, Rusty Russell,Anton Igersheim, Adele Smith.</p></div>	https://treatment.plazi.org/id/03D7A111FFA2FFB0F96BD227FEA1F9D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Craven, L. A.	Craven, L. A. (2019): Studies in Papuasian Syzygium (Myrtaceae): 1. Subgenus Perikion revised. Blumea 64 (2): 115-122, DOI: 10.3767/blumea.2019.64.02.03, URL: https://doi.org/10.3767/blumea.2019.64.02.03
