taxonID	type	description	language	source
03C987DF3306FFF99ABE0B062986FAFB.taxon	description	57: 19. 2012; Naiki et al., Acta Phytotax. Geobot. 66: 156. 2015; Napiroon et al., Phytotaxa 439: 10. 2020; Dang & Naiki, Phytotaxa 581: 27. 2023. Litosanthes biflora Blume, Cat. Gew. Buitenzorg (Blume): 22. 1823. Type: INDONESIA, Java, Blume s. n. (lectotype L! [L. 0000722], designated by Zhu, 2002). Lasianthus gracilis King & Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 132. 1904. Type: MALAYSIA, Perak, Tea Gardens, Ridley 2904 (lectotype SING! [SING 0059255], designated by Wong, 1988).	en	Q. T., Pham, S., Tagane, P., Souladeth, T., Yamamoto, P., Phonepaseuth, T., Vongthavone, D., Kongxaisavath, B. V., Truong, Dang, V. S. (2025): Two new records of Lasianthus (Rubiaceae) for the Flora of Laos. Rheedea 35 (1): 39-45, DOI: 10.22244/rheedea.2025.35.01.08
03C987DF3306FFFB9ABE0C002A59FC07.taxon	description	Flowering & fruiting: Flowers and fruits were collected in September. Habitat: Lasianthus biflorus was collected in the semi-shaded understory of semi-evergreen forests at 510 – 540 m elev., where the species of Lauraceae, Rubiaceae, Myrtaceae, Fagaceae and Fabaceae are dominant. Distribution: Cambodia, China, Indonesia, Malaysia, Philippines, Thailand, Taiwan and Vietnam. In Laos, this species occurs in the Nam Kading National Protected Area of Bolikhamxai Province in central Laos. Specimens examined: LAOS, Bolikhamxai Province, Viengthong District, Ban Nadi, Nam Kading National Protected Area, at 515 m, N 18 O 19 ’ 49.401 ’’ E 104 O 31 ’ 27.655 ’’, 04.09.2023, Tagane et al. Z 367 FOF [FOF 005521], KAG [KAG 186811], VMM); same locality, 05.09.2023, Tagane et al. Z 416 FOF [FOF 005571], KAG [KAG 186853], VNM [VNM 00072084]); Khamkeuth District, near Ban Kaengbid, at 537 m elev., N 18 O 13 ’ 9.859 ’’ E 104 O 30 ’ 21.09 ’’, 15.03.2024, Tagane et al. Z 839 FOF [FOF 005966], KAG [KAG 187254], VNM [VNM 00070633]. Notes: Lasianthus biflorus is a new record for the Flora of Laos. This species is closely related to Lasianthus bifloroideus H. Zhu endemic to Borneo but differs in having twigs densely hirsute (vs. thinly puberulous), cymes with 0.5 – 1.5 cm long peduncles (vs. sessile or subsessile), and smooth fruit (vs. warty). Among the species of Lasianthus in Laos, L. biflorus is easily recognized by its small size of lamina, which is up to 2.5 cm long and the smallest among the Lasianthus species in Laos, long peduncles up to 1.5 cm long, and ripening black fruits.	en	Q. T., Pham, S., Tagane, P., Souladeth, T., Yamamoto, P., Phonepaseuth, T., Vongthavone, D., Kongxaisavath, B. V., Truong, Dang, V. S. (2025): Two new records of Lasianthus (Rubiaceae) for the Flora of Laos. Rheedea 35 (1): 39-45, DOI: 10.22244/rheedea.2025.35.01.08
03C987DF3304FFFB9ABE0A8F2C19FA02.taxon	description	Flowering & fruiting: Fruits were collected in September and March, flowers not seen. Habitat: Lasianthus hispidulus grows in secondary evergreen forests, at 450 – 520 m elevation. Distribution: Cambodia, China, Indonesia, Japan, Malaysia, Myanmar, Papua New Guinea, Thailand, Taiwan and Vietnam. Our finding in Laos fills the gap of the distribution of this species. Specimens examined: LAOS, Bolikhamxai Province, Viengthong District, Ban Nadi, Nam Kading National Protected Area, at 515 m, N 18 ° 13 ’ 9.859 ’’, E 104 ° 30 ’ 21.09 ’’, 05.09.2023, Tagane et al. Z 417 (FOF, KAG [KAG 186854], VNM). Khammouane Province, Khounkham District, Tad Nam Sanam, at 458 m, N 18 ° 13 ’ 10.6674 ’’, E 104 ° 30 ’ 11.016 ’’, 15.03.2024, Tagane et al. Z 828 (FOF, KAG [KAG 187243], VNM [VNM 00072869]. Notes: Lasianthus hispidulus is newly recorded in the Flora of Laos. This species is closely related to L. fordii Hance which is also distributed widely in tropical and subtropical regions in Southeast and East Asia, but distinguished by its branches, petioles, lower leaf surface and calyx densely covered with yellowish brown hairs (vs. subglabrous to thinly appressed hairy). It is also similar to L. curtisii King & Gamble distributed widely in Southeast to East Asia but differs in having calyx lobes shorter than calyx tubes (vs. longer than calyx tubes).	en	Q. T., Pham, S., Tagane, P., Souladeth, T., Yamamoto, P., Phonepaseuth, T., Vongthavone, D., Kongxaisavath, B. V., Truong, Dang, V. S. (2025): Two new records of Lasianthus (Rubiaceae) for the Flora of Laos. Rheedea 35 (1): 39-45, DOI: 10.22244/rheedea.2025.35.01.08
