identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C987DF3306FFF99ABE0B062986FAFB.text	03C987DF3306FFF99ABE0B062986FAFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lasianthus biflorus (Blume) M. Gangop. & Chakrab., J. Econ.	<div><p>Lasianthus biflorus (Blume) M.Gangop. &amp; Chakrab., J. Econ. Taxon.Bot. 16: 338. 1992;Zhu &amp;</p><p>Taylor, Fl. China 19: 189. 2011; Zhu et al., Blumea</p><p>57: 19. 2012; Naiki et al., Acta Phytotax. Geobot.</p><p>66: 156. 2015; Napiroon et al., Phytotaxa 439: 10.</p><p>2020; Dang &amp; Naiki, Phytotaxa 581: 27. 2023.</p><p>Litosanthes biflora Blume, Cat. Gew. Buitenzorg</p><p>(Blume): 22. 1823. Type: INDONESIA, Java, Blume</p><p>s.n. ( lectotype L! [L.0000722], designated by Zhu,</p><p>2002).</p><p>Lasianthus gracilis King &amp; Gamble, J. Asiat. Soc. Bengal, Pt. 2, Nat. Hist. 73: 132. 1904. Type: MALAYSIA, Perak, Tea Gardens, Ridley 2904 (lectotype SING! [SING0059255], designated by Wong, 1988).</p><p>Lasianthus gainii M.Gangop. &amp; Chakrab., J. Econ. Taxon. Bot. 16: 338. 1992. Syntype: Kusukusu, s.d., Hayata &amp; Sasaki s.n. (n.v.).</p></div>	https://treatment.plazi.org/id/03C987DF3306FFF99ABE0B062986FAFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Q. T., Pham;S., Tagane;P., Souladeth;T., Yamamoto;P., Phonepaseuth;T., Vongthavone;D., Kongxaisavath;B. V., Truong;Dang, V. S.	Q. T., Pham, S., Tagane, P., Souladeth, T., Yamamoto, P., Phonepaseuth, T., Vongthavone, D., Kongxaisavath, B. V., Truong, Dang, V. S. (2025): Two new records of Lasianthus (Rubiaceae) for the Flora of Laos. Rheedea 35 (1): 39-45, DOI: 10.22244/rheedea.2025.35.01.08
03C987DF3306FFFB9ABE0C002A59FC07.text	03C987DF3306FFFB9ABE0C002A59FC07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Litosanthes gracilis Hayata, Icon. Pl. Formosan.	<div><p>Litosanthes gracilis Hayata, Icon. Pl. Formosan. 2: 113. 1912. Syntype: Botanrosha, G.Nakahara s.n. (n.v.). Fig. 1</p><p>Shrubs, evergreen, up to 2 m high. Twigs terete, densely hirsute; internodes 1–1.5(–4) cm long. Leaves opposite; blades elliptic or subovate, 1–2.5 × 0.5–1.5 cm, chartaceous, greenish-brown when dry, adaxially glabrous, abaxially puberulent, apex acute, aristate with arista 1–2 mm long, base cuneate, margin entire, midrib slightly prominent adaxially, prominent abaxially, secondary veins 6–10 pairs, ascending at an angle of 60–80° from midrib, curved to the margin, flat adaxially, tertiary veins subparallel; petioles 1–2 mm long, puberulent. Stipules lanceolate, 1–1.5 mm long, puberulent. Inflorescences solitary or cymose, peduncles 0.5–1.5 cm long, puberulent; bracts small, inconspicuous. Flowers pedicellate, pedicels 0.5–1 mm long; calyx campanulate, 1–2 mm long, glabrous, lobes 4; corolla white, 2–4.5 mm long, glabrous outside, triangular-lanceolate, lobes ovate. Fruits drupaceous, obovoid, 3.5–5 mm in diam., glabrous, smooth, black when ripe; pyrenes 4.</p><p>Flowering &amp; fruiting: Flowers and fruits were collected in September.</p><p>Habitat: Lasianthus biflorus was collected in the semi-shaded understory of semi-evergreen forests at 510–540 m elev., where the species of Lauraceae, Rubiaceae, Myrtaceae, Fagaceae and Fabaceae are dominant.</p><p>Distribution: Cambodia, China, Indonesia, Malaysia, Philippines, Thailand, Taiwan and Vietnam. In Laos, this species occurs in the Nam Kading National Protected Area of Bolikhamxai Province in central Laos.</p><p>Specimens examined: LAOS, Bolikhamxai Province, Viengthong District, Ban Nadi, Nam Kading National Protected Area, at 515 m, N 18 O 19 ’ 49.401’’ E 104 O 31 ’ 27.655’’, 04.09.2023, Tagane et al. Z367 FOF [FOF005521], KAG [KAG186811], VMM ); same locality, 05.09.2023, Tagane et al. Z416 FOF [FOF005571], KAG [KAG186853], VNM [VNM00072084] ); Khamkeuth District, near Ban Kaengbid, at 537 m elev., N 18 O 13 ’ 9.859’’ E 104 O 30 ’ 21.09’’, 15.03.2024, Tagane et al. Z839 FOF [FOF005966], KAG [KAG187254], VNM [VNM00070633] .</p><p>Notes: Lasianthus biflorus is a new record for the Flora of Laos. This species is closely related to Lasianthus bifloroideus H.Zhu endemic to Borneo but differs in having twigs densely hirsute (vs. thinly puberulous), cymes with 0.5–1.5 cm long peduncles (vs. sessile or subsessile), and smooth fruit (vs. warty). Among the species of Lasianthus in Laos, L. biflorus is easily recognized by its small size of lamina, which is up to 2.5 cm long and the smallest among the Lasianthus species in Laos, long peduncles up to 1.5 cm long, and ripening black fruits.</p><p>Lasianthus hispidulus (Drake) Pit. In P.H. Lecomte, Fl. Indo-Chine 3: 393. 1924; Pham, Fl. Vietnam 3: 205, f. 8523. 2000; Zhu, Acta Phytotax. Sin. 39: 130. 2001; Zhu &amp; Taylor, Fl. China 19: 192. 2011; Zhu et al., Blumea 57: 48. 2012; Naiki et al., Acta Phytotax. Geobot. 66: 168. 2015; Napiroon et al., Phytotaxa 439: 17. 2020; Dang &amp; Naiki, Phytotaxa 581: 73. 2023. Mephitidia hispidula Drake, J. Bot. (Morot) 9: 239. 1895. Lectotype (designated by Zhu, 2001): VIETNAM, Thai Nguyen Province, Eberhardt 3852 (P).</p><p>Lasianthus brevidens Craib, Kew Bull. 1933: 19 . 1933. Type: THAILAND, Ratchburi, Prachuap, Khao Luang, 700 m, Kerr 10833 (holo K [K000777026!], isotypes BK [BK257382 image!], BM [BM000945605 image!]).</p></div>	https://treatment.plazi.org/id/03C987DF3306FFFB9ABE0C002A59FC07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Q. T., Pham;S., Tagane;P., Souladeth;T., Yamamoto;P., Phonepaseuth;T., Vongthavone;D., Kongxaisavath;B. V., Truong;Dang, V. S.	Q. T., Pham, S., Tagane, P., Souladeth, T., Yamamoto, P., Phonepaseuth, T., Vongthavone, D., Kongxaisavath, B. V., Truong, Dang, V. S. (2025): Two new records of Lasianthus (Rubiaceae) for the Flora of Laos. Rheedea 35 (1): 39-45, DOI: 10.22244/rheedea.2025.35.01.08
03C987DF3304FFFB9ABE0A8F2C19FA02.text	03C987DF3304FFFB9ABE0A8F2C19FA02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lasianthus bunzanensis Simizu, Trans. Nat. Hist. Soc.	<div><p>Lasianthus bunzanensis Simizu, Trans. Nat. Hist. Soc. Taiwan 34: 301. 1944. Type: CHINA, Taiwan, Taipei, Suzuki 16385 (holotype TI!). Fig. 2.</p><p>Shrubs, evergreen, up to 1.5 m high. Twigs terete, densely villous with yellowish brown hairs; internodes 2–4 cm long. Leaves opposite; blades elliptic to ovate-oblong, 6.5–10 × 2.5–5 cm, thinly coriaceous, yellowish brown when dry, adaxially glabrous, abaxially densely villous, yellowish brown, apex cuspidate-acuminate or acuminate, aristate with arista 1.5–2 mm long, base acute or obtuse, margins entire, midrib prominent adaxially, prominent abaxially, secondary veins 4–6 pairs, ascending at an angle of 45–70° from midrib, curved to the margins, slightly prominent adaxially, tertiary veins parallel; petioles 4–7.5 mm long, densely villous with yellowish brown hairs. Stipules subulate, 0.5–1 mm long, hirsute on both surfaces. Inflorescences solitary, subsessile; bracts absent. Flowers subsessile; calyx campanulate, 1–1.5 mm long, densely villous with yellowish brown hairs, tube c. 1 mm long, lobes 5, c. 0.5 mm long; corolla not seen. Fruits drupaceous, globose, 3.5–5 mm in diam., hirsute, blue when ripe; pyrenes 5.</p><p>Flowering &amp; fruiting: Fruits were collected in September and March, flowers not seen.</p><p>Habitat: Lasianthus hispidulus grows in secondary evergreen forests, at 450–520 m elevation.</p><p>Distribution: Cambodia, China, Indonesia, Japan, Malaysia, Myanmar, Papua New Guinea, Thailand, Taiwan and Vietnam. Our finding in Laos fills the gap of the distribution of this species.</p><p>Specimens examined: LAOS, Bolikhamxai Province, Viengthong District, Ban Nadi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.50586&amp;materialsCitation.latitude=18.219406" title="Search Plazi for locations around (long 104.50586/lat 18.219406)">Nam Kading National Protected Area</a>, at 515 m, N 18° 13’ 9.859’’, E 104° 30’ 21.09’’, 05.09.2023, Tagane et al. Z417 (FOF, KAG [KAG186854], VNM) . Khammouane Province, Khounkham District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.50306&amp;materialsCitation.latitude=18.219631" title="Search Plazi for locations around (long 104.50306/lat 18.219631)">Tad Nam Sanam</a>, at 458 m, N 18° 13’ 10.6674’’, E 104° 30’ 11.016’’, 15.03.2024, Tagane et al. Z828 (FOF, KAG [KAG187243], VNM [VNM00072869].</p><p>Notes: Lasianthus hispidulus is newly recorded in the Flora of Laos. This species is closely related to L. fordii Hance which is also distributed widely in tropical and subtropical regions in Southeast and East Asia, but distinguished by its branches, petioles, lower leaf surface and calyx densely covered with yellowish brown hairs (vs. subglabrous to thinly appressed hairy). It is also similar to L. curtisii King &amp; Gamble distributed widely in Southeast to East Asia but differs in having calyx lobes shorter than calyx tubes (vs. longer than calyx tubes).</p></div>	https://treatment.plazi.org/id/03C987DF3304FFFB9ABE0A8F2C19FA02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Q. T., Pham;S., Tagane;P., Souladeth;T., Yamamoto;P., Phonepaseuth;T., Vongthavone;D., Kongxaisavath;B. V., Truong;Dang, V. S.	Q. T., Pham, S., Tagane, P., Souladeth, T., Yamamoto, P., Phonepaseuth, T., Vongthavone, D., Kongxaisavath, B. V., Truong, Dang, V. S. (2025): Two new records of Lasianthus (Rubiaceae) for the Flora of Laos. Rheedea 35 (1): 39-45, DOI: 10.22244/rheedea.2025.35.01.08
