identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03CC1E69A337FFD7B924F93DB77EFED7.text	03CC1E69A337FFD7B924F93DB77EFED7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alycaeinae W. T. Blanford 1864	<div><p>Subfamily Alycaeinae W.T. Blanford, 1864</p><p>Alycaeinae W.T. Blanford, 1864: 465 .</p><p>Remarks</p><p>In our previous publications (Páll-Gergely et al. 2016, 2017, 2020, 2021; Páll-Gergely &amp; Asami 2017; Páll-Gergely &amp; Auffenberg 2019; Páll-Gergely &amp; Hunyadi 2022; Páll-Gergely 2023), we treated this group as a family of its own right. However, the recent molecular phylogeny (Páll-Gergely et al. 2024) suggested treating Alycaeinae as a family would result in numerous paraphyletic groups, and therefore, Cyclophoridae should be maintained at the family rank, and Alycaeinae is to be treated as one of its subfamilies.</p></div>	https://treatment.plazi.org/id/03CC1E69A337FFD7B924F93DB77EFED7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A337FFD5B971F9B2B406F90C.text	03CC1E69A337FFD5B971F9B2B406F90C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclophoridae Gray 1847	<div><p>Family Cyclophoridae Gray, 1847</p><p>Cyclophoridae Gray, 1847: 181 .</p></div>	https://treatment.plazi.org/id/03CC1E69A337FFD5B971F9B2B406F90C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD7B92BFC7CB4C2FBFB.text	03CC1E69A335FFD7B92BFC7CB4C2FBFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus mizoramensis Pall-Gergely & Aravind 2025	<div><p>Chamalycaeus mizoramensis species group</p><p>Remarks</p><p>Chamalycaeus mizoramensis sp. nov. is classified in its own species group, as this species is not similar to any other species of Alycaeinae in the region.</p></div>	https://treatment.plazi.org/id/03CC1E69A335FFD7B92BFC7CB4C2FBFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD7B915FE75B74CFCCF.text	03CC1E69A335FFD7B915FE75B74CFCCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus Mollendorff 1897	<div><p>Genus Chamalycaeus Möllendorff, 1897</p><p>Alycaeus (Chamalycaeus) Möllendorff, 1897b: 93 .</p><p>Remarks</p><p>Chamalycaeus is characterised by a smooth (or pitted) protoconch and a spirally striated R1. Most species classified in Chamalycaeus here are transferred from the genus Dicharax (see Páll-Gergely et al. 2020).</p><p>Although no spiral striation was found on the protoconch of Chamalycaeus oglei (Godwin-Austen, 1914) (see Páll-Gergely et al. 2020), it is moved to the genus Metalycaeus Pilsbry, 1900 due to its similarity with M. brahma (Godwin-Austen, 1886) and other Himalayan species of Metalycaeus . Furthermore, Chamalycaeus perplexus (Godwin-Austen, 1914) is moved to Metalycaeus due to its similarity with M. polygonoma .</p></div>	https://treatment.plazi.org/id/03CC1E69A335FFD7B915FE75B74CFCCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A335FFD9BA94FB88B43FFDF7.text	03CC1E69A335FFD9BA94FB88B43FFDF7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus mizoramensis Pall-Gergely & Aravind 2025	<div><p>Chamalycaeus mizoramensis Páll-Gergely &amp; Aravind sp. nov.</p><p>urn:lsid:zoobank.org:act: 734649E7-5CAC-4462-9578-CC323E60FE85</p><p>Fig. 4</p><p>Diagnosis</p><p>A relatively large species of Chamalycaeus with a ca half whorl-long R2+R3, and R3 with a low central swelling and a fringed aperture.</p><p>Etymology</p><p>This species is named after the state of Mizoram, where the Blue Mountain is situated.</p><p>Type material examined</p><p>Holotype</p><p>INDIA – Mizoram • empty shell (D: 4.8 mm, H: 3 mm); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; AR15; NZSI LM1715.</p><p>Description</p><p>Shell off-white/ivory in colour, not translucent; shell outline slightly oval in dorsal view; spire slightly elevated, low conical; body whorl rounded; protoconch low, finely granular, spiral striae lacking, consisting of nearly 2 whorls; R1 of 1.75 whorls, with rather irregular, dense, low ribs (last half whorl of R1 with ca 55–60 ribs) and weaker, denser spiral striation; boundary between R1 and R2 not conspicuous because R2 ribs are only slightly higher and sharper than R1 ribs, but R2 area more inflated than R1; sutural tube conspicuously shorter than R2, probably due to teratological condition; R2 with ca 50– 52 straight, sharp but low ribs; R2+R3 ca half whorl, R3 shorter than R2, boundary between R2 and R3 clearly visible due to a slight constriction; R3 with long and low central swelling; R3 ribs stand approximately as densely as R1 ribs, with the exception of the beginning of R3 with more widely-spaced ribbing; aperture strongly oblique to shell axis, rounded with 5 small bays on the palatal side, resulting in a fringed peristome; boundary between inner and outer peristomes is not conspicuous but clearly visible, inner peristome thickened, slightly protruding and expanded; outer peristome thin, sharp, slightly expanded, especially towards umbilicus; umbilicus relatively narrow, ca one fourth of shell width, rounded.</p><p>MEASUREMENTS. D: 4.8 mm, H: 3 mm.</p><p>Distribution</p><p>Known only from the Blue Mountain region.</p><p>Differential diagnosis</p><p>No other species of Chamalycaeus resembles Chamalycaeus mizoramensis sp. nov. The relatively large size, the fringed aperture, the dense, fine ribbing on R1 and R3, and the long R2+R3 (ca half whorl) are characteristic of this species.</p></div>	https://treatment.plazi.org/id/03CC1E69A335FFD9BA94FB88B43FFDF7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFD9B916FD94B565FCA0.text	03CC1E69A33BFFD9B916FD94B565FCA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus armillatus (Benson 1856)	<div><p>Chamalycaeus armillatus species group</p><p>Remarks</p><p>Three tiny species from Myanmar belong here, which are characterized by widely-spaced R1 ribs, a short R2, a low and blunt R3 swelling, and a circular aperture.</p></div>	https://treatment.plazi.org/id/03CC1E69A33BFFD9B916FD94B565FCA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFD9B91CFCA1B7B0FAF1.text	03CC1E69A33BFFD9B91CFCA1B7B0FAF1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus armillatus (Benson 1856)	<div><p>Chamalycaeus armillatus (Benson, 1856)</p><p>Fig. 5A–D</p><p>Alycaeus armillatus Benson, 1856: 227 .</p><p>Alycaeus armillatus – Sowerby 1877: pl. 5, species 38. — Godwin-Austen 1914: 406, pl. 151 figs 3, 3a.</p><p>Alycaeus (Dicharax) armillatus – Kobelt 1902: 365. — Gude 1921: 236–237.</p><p>Chamalycaeus armillatus – Páll-Gergely et al. 2020: 36; 2021: 14, fig. 10b. — Preece et al. 2022: 79, fig. 32b.</p><p>Diagnosis</p><p>See under C. humilis (W.T. Blanford, 1862) and C. irmatallus Páll-Gergely, 2021 .</p><p>Type locality</p><p>“ad Thyet-Mio”.</p></div>	https://treatment.plazi.org/id/03CC1E69A33BFFD9B91CFCA1B7B0FAF1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33BFFDBB937FA92B2AEFDD3.text	03CC1E69A33BFFDBB937FA92B2AEFDD3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus humilis (W. T. Blanford 1862)	<div><p>Chamalycaeus humilis (W.T. Blanford, 1862)</p><p>Fig. 5E–I</p><p>Alycaeus humilis W.T. Blanford, 1862: 136–137 .</p><p>Alycaeus humilis – Sowerby 1877: pl. 5, species 40. — Godwin-Austen 1914: 408–409, pl. 151 fig. 8. Alycaeus (Dicharax) humilis – Kobelt 1902: 372. — Gude 1921: 255–256.</p><p>Dicharax humilis – Páll-Gergely et al. 2020: 64.</p><p>Type material examined</p><p>MYANMAR • holotype (single shell mentioned in the original description: Fig. 5E–I); River Bank, Myanoung, Pegu; NHMUK 1906.4.4.69 .</p><p>Type locality</p><p>“ad Akouktoung, ad ripas fluminis Irawaddi, in provincia Burmana Pegu ”.</p><p>Differential diagnosis</p><p>This species has a comparatively smaller aperture than C. armillatus . Chamalycaeus irmatallus has denser and finer R1 ribs, more expanded outer peristome and a less inflated R3 swelling.</p><p>Remarks</p><p>Only a single, corroded shell is known, therefore the important traits (presence or absence of spiral striation on the teleoconch) could not be examined without doubt. We classified this species in the genus Dicharax in our previous catalogue, but during the revision of Himalayan Dicharax, we had to reconsider our decision. Since the shell shape of A. humilis is most similar to the geographically closely occurring Chamalycaeus armillatus and C. irmatallus, we move this species to Chamalycaeus .</p></div>	https://treatment.plazi.org/id/03CC1E69A33BFFDBB937FA92B2AEFDD3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDBB916FA27B347F8A6.text	03CC1E69A339FFDBB916FA27B347F8A6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus asaluensis (Godwin-Austen 1914)	<div><p>Chamalycaeus asaluensis species group</p><p>Remarks</p><p>The species belonging here are characterized by a low conical, colourless shell, a strongly ribbed R1 (rib density relatively low), and an upper incision in the aperture. The aperture shape (fringed to circular) and the shape of R3 swelling are highly variable.</p><p>Some species of this species group have been classified in the genus Dicharax (see Páll-Gergely et al. 2020), because spiral striation could not be observed on them, partly because the examined shells were corroded, and partly because the spiral striation on the shells are naturally weak and difficult to observe.</p></div>	https://treatment.plazi.org/id/03CC1E69A339FFDBB916FA27B347F8A6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDBB930FD77B78DFA23.text	03CC1E69A339FFDBB930FD77B78DFA23.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus irmatallus Pall-Gergely 2021	<div><p>Chamalycaeus irmatallus Páll-Gergely, 2021</p><p>Fig. 5J–N</p><p>Chamalycaeus irmatallus Páll-Gergely in Páll-Gergely et al., 2021: 14–15, figs 9, 10a.</p><p>Type material examined</p><p>THAILAND • holotype; Surat Thani Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.39805&amp;materialsCitation.latitude=8.659717" title="Search Plazi for locations around (long 99.39805/lat 8.659717)">Limestone Mt</a>, 6 km S of Na San; 8°39.583′ N, 99°23.883′ E; 100 m a.s.l.; 3 Jun. 1987; F.G. Thompson leg.; UF 279508 • 11 paratypes; same data as for preceding; UF 551216 .</p><p>Additional material examined</p><p>MYANMAR • 6 shells (see Páll-Gergely et al. 2021: fig. 10); Thayet-myo, Pegu; Blanford coll.; NHMUK 1906.4.4.71 .</p><p>Type locality</p><p>“ Thailand, Surat Thani Province, limestone mtn., 6 km S of Na San, 8°39.583′ N, 99°23.883′ E; 100 m a.s.l.”.</p><p>Differential diagnosis</p><p>Chamalycaeus armillatus has a more rounded body whorl, longer R2 and R3, and a thicker peristome with greater distance between the edges of the inner and outer peristomes (Páll-Gergely et al. 2021). See also under C. humilis .</p></div>	https://treatment.plazi.org/id/03CC1E69A339FFDBB930FD77B78DFA23	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A339FFDDBAD2F8DBB4A1FA62.text	03CC1E69A339FFDDBAD2F8DBB4A1FA62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus asaluensis (Godwin-Austen 1914)	<div><p>Chamalycaeus asaluensis (Godwin-Austen, 1914)</p><p>Fig. 6</p><p>Alycaeus asaluensis Godwin-Austen, 1914: 385–386, pl. 145 figs 2, 2a–b.</p><p>Alycaeus barowliensis Godwin-Austen, 1914: 352, pl. 141 fig. 4. Syn. nov.</p><p>Alycaeus crispatus var. – Godwin-Austen 1874: 93, pl. 4 fig. 2.</p><p>Alycaeus barowliensis – Gude 1921: 205.</p><p>Alycaeus (Dicharax) asaluensis – Gude 1921: 237.</p><p>Alycaeus (Alycaeus) burowliensis [sic] – Ramakrishna et al. 2010: 46.</p><p>Chamalycaeus (Dicharax) asaluensis – Ramakrishna et al. 2010: 56.</p><p>Dicharax asaluensis – Páll-Gergely et al. 2020: 54.</p><p>Dicharax (?) barowliensis – Páll-Gergely et al. 2020: 77.</p><p>Diagnosis</p><p>Differs from the congeners by the long R3 swelling having straight anterior part.</p><p>Type material examined</p><p>INDIA • 2 syntypes of A. asaluensis (Fig. 6A–E); Dihung, N. Cachar; Godwin-Austen coll.; NHMUK 1903.7.1.2636 • holotype of A. barowliensis (single shell mentioned in the original description: Fig. 6F– J); Barowli R. Durrang, Assam; Godwin-Austen coll.; NHMUK 1903.7.1.2723 .</p><p>Type localities</p><p>“Barowli River, Akha Hills, Durrang, Assam ” ( A. barowliensis); “Neuglo”, “Phulong” and “Dihung River, N. Cachar, north of Asalu” ( A. asaluensis).</p><p>Differential diagnosis</p><p>Dicharax aspidentatus Gittenberger, Choki Gyeltshen &amp; Sherub Sherub, 2024 has a higher spire and larger distance between the inner and outer peristomes. Chamalycaeus crispatus (Godwin-Austen, 1871) has a constriction anterior to the R3 swelling, is glossier and less densely ribbed. Dicharax rugosus (Godwin-Austen, 1914) has a higher spire, a shorter R3 swelling and more strongly fringed peristome.</p><p>Remarks</p><p>Only the corroded holotype of A. barowliensis is known, which is very similar to C. asaluensis, but has a slightly more globose shell (and narrower umbilicus) and more widely-spaced ribs on R1. Nevertheless, given their geographic proximity and that all other traits are identical, these differences are best explained by intraspecific variability.</p><p>This species was included in the genus Dicharax due to the absence of spiral striation (Páll-Gergely et al. 2020). However, the shell shape and other characters agree with this species group, and therefore, it is reclassified in the genus Chamalycaeus .</p></div>	https://treatment.plazi.org/id/03CC1E69A339FFDDBAD2F8DBB4A1FA62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33FFFDFBACAF9E6B518F9F3.text	03CC1E69A33FFFDFBACAF9E6B518F9F3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus crispatus (Godwin-Austen 1871)	<div><p>Chamalycaeus crispatus (Godwin-Austen, 1871)</p><p>Fig. 7</p><p>Alycaeus crispatus Godwin-Austen, 1871: 91–92, pl. 4 fig. 1.</p><p>Alycaeus crispatus var. rywukensis Godwin-Austen, 1914: 373–374, pl. 154 figs 3, 3a. Syn. nov. Alycaeus crispatus var. minimus Godwin-Austen, 1914: 373, pl. 148 figs 5, 5a. Syn. nov.</p><p>Alycaeus crispatus – Godwin-Austen 1875: 8, pl. 4 fig. 3; 1914: 371–372, 389, pl. 145 figs 1, 1a–b. Alycaeus (Dicharax) crispatus – Kobelt 1902: 367. — Gude 1921: 242–243.</p><p>Alycaeus (Dicharax) crispatus var. minima – Gude 1921: 243–244.</p><p>Alycaeus (Dicharax) crispatus var. rywukensis – Gude 1921: 244.</p><p>Chamalycaeus (Dicharax) crispatus – Ramakrishna et al. 2010: 58.</p><p>Dicharax (?) crispatus – Páll-Gergely et al. 2020: 82.</p><p>Dicharax (?) crispatus minimus – Páll-Gergely et al. 2020: 83.</p><p>Dicharax (?) crispatus rywukensis – Páll-Gergely et al. 2020: 83.</p><p>Dicharax crispatus – Gittenberger et al. 2024: 205, figs 24, 27.</p><p>Diagnosis</p><p>The narrow and high R3 swelling distinguishes this species from other congeners of the same species group.</p><p>Type material examined</p><p>INDIA • 5 syntypes of A. crispatus (Fig. 7A–E); Shibak, Habiang Garo Hills; NHMUK 1903.7.1.2635 • 11 syntypes of A. crispatus; same container as previous (probably same locality); NHMUK 1903.7.1.2759 • holotype of A. crispatus var. minimus (single shell mentioned in the original description: Fig. 7K–O); Habiang Garo Hills, W. Khasi; Godwin-Austen leg.; NHMUK 1906.4.4.176 • 2 syntypes of A. crispatus var. rywukensis (Fig. 7F–J); Rywuk, Garo Hills, South base of; Godwin-Austen coll.; NHMUK 1903.7.1.2637 .</p><p>Type localities</p><p>“Khasia, Jaintia and N. Cachar Hills” ( A. crispatus); “Habiang Garo Hills, West Khasi” ( A. crispatus var. minimus); “Rywuk Valley of the Garo Hills” ( A. crispatus var. rywukensis).</p><p>Differential diagnosis</p><p>Dicharax rugosus has a higher spire, and a more strongly and densely ribbed R1. See also under C. asaluensis .</p><p>Remarks</p><p>Godwin-Austen (1914: 372) stated in his previous paper (Godwin-Austen 1871: 93) that the Alycaeus crispatus variety from north Cachar was renamed A. asaluensis . The originally figured sample (Godwin-Austen 1871: pl. 4 fig. 1) is from Shibak, Gabir Valley (Godwin-Austen 1914: 372).</p><p>Alycaeus crispatus and A. crispatus var. minimus do not differ in any important shell characters, and therefore, the latter is a junior synonym of the former. Alycaeus crispatus var. rywukensis is smaller than the previous two and has a ribbed instead of glossy R3. Nevertheless, we interpret this as intraspecific variability.</p><p>Gittenberger et al. (2024) reported this species from Bhutan. We agree with their identification based on the photos they provided.</p><p>This species was included in the genus Dicharax due to the absence of spiral striation (Páll-Gergely et al. 2020). However, the shell shape and other characters agree with this species group, and therefore, it is reclassified in the genus Chamalycaeus . Furthermore, re-examination of the syntypes of A. crispatus var. makarsae revealed spiral striation on the teleoconch.</p></div>	https://treatment.plazi.org/id/03CC1E69A33FFFDFBACAF9E6B518F9F3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A33DFFC1BAD7F997B556FBFC.text	03CC1E69A33DFFC1BAD7F997B556FBFC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus logtakensis (Godwin-Austen 1914)	<div><p>Chamalycaeus logtakensis (Godwin-Austen, 1914)</p><p>Fig. 8</p><p>Alycaeus logtakensis Godwin-Austen, 1914: 394–395, pl. 155 fig. 6.</p><p>Alycaeus tanghali Godwin-Austen, 1914: 401, pl. 137 figs 3, 3a–b. Syn. nov.</p><p>Alycaeus logtakensis – Gude 1921: 209–210.</p><p>Alycaeus (Alycaeus) logtakensis – Ramakrishna et al. 2010: 48.</p><p>Dicharax (?) logtakensis – Páll-Gergely et al. 2020: 94.</p><p>Chamalycaeus tanghali – Páll-Gergely et al. 2020: 46.</p><p>Diagnosis</p><p>The low R3 without conspicuous swelling distinguishes this species from all congeners of this species group.</p><p>Type material examined</p><p>INDIA • 1 syntype of A. logtakensis (Fig. 8A–E); on a low hill, Logtak Lake, Munipur; Godwin-Austen coll.; “figured”; NHMUK 1903.7.1.2639 • 6 syntypes of A. tanghali; Munipur; figured by Godwin-Austen; NHMUK 1903.7.1.2671 (Fig. 8F–J) .</p><p>Type localities</p><p>“Logtak Lake, Munipur (No. 2639 B.M. Coll.), on a low hill near the northern shore” ( A. logtakensis); “Munipur. Exact locality not recorded; somewhere on the northern side of the valley” ( A. tanghali).</p><p>Differential diagnosis</p><p>The most similar species is C. sculpturus, but that species has a less fringed aperture and a more rounded body whorl.</p><p>Remarks</p><p>There are no conchological differences between A. logtakensis and A. tanghali, and they are both from central Manipur State. Their descriptions were published in the same work but A. tanghali appeared a few pages later, therefore A. logtakensis is chosen to be the valid name. Godwin-Austen (1914) compared A. tanghali only with A. sculptilis .</p><p>Alycaeus logtakensis was included in the genus Dicharax because no spiral striation was observed on the corroded syntype (Páll-Gergely et al. 2020). However, re-examination of the syntype of A. logtakensis revealed spiral striation on the teleoconch. Chamalycaeus tanghali was already classified in Chamalycaeus due to spiral striation on the teleoconch.</p></div>	https://treatment.plazi.org/id/03CC1E69A33DFFC1BAD7F997B556FBFC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A323FFC5BAF4FB8DB591FCA9.text	03CC1E69A323FFC5BAF4FB8DB591FCA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus reflectus Pall-Gergely & Aravind 2025	<div><p>Chamalycaeus reflectus Páll-Gergely &amp; Aravind sp. nov.</p><p>urn:lsid:zoobank.org:act: E606B321-913F-409B-BCD7-B47531BB01DD</p><p>Figs 9–10, 12A</p><p>Diagnosis</p><p>A small species of Chamalycaeus with a ca quarter whorl-long R2+R3, an R3 with a low central swelling (situated slightly closer to the peristome), a strongly expanded outer peristome, and a rounded inner peristome.</p><p>Etymology</p><p>The specific epithet refers to the reflected outer peristome, which is characteristic of this species.</p><p>Type material examined</p><p>Holotype</p><p>INDIA – Mizoram • 1 empty shell (D: 1.9 mm, H: 1.4 mm, Figs 9, 12A); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 55; NZSI LM1723.</p><p>Paratype</p><p>INDIA – Mizoram • 1 empty shell; same data as for holotype; NZSI LM1724 .</p><p>Additional material examined</p><p>INDIA – Mizoram • 2 empty shells; same data as for holotype; Ar56; NZSI LM1725 • 1 empty shell (Fig. 10); same data as for holotype; AR44; NZSI LM1913 . – Tripura • 1 empty shell (Fig. 11); South Tripura, Purba Patichhari, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.5049&amp;materialsCitation.latitude=23.403784" title="Search Plazi for locations around (long 91.5049/lat 23.403784)">Towards Garji</a> from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=91.5049&amp;materialsCitation.latitude=23.403784" title="Search Plazi for locations around (long 91.5049/lat 23.403784)">Santirbazar</a>; 23°24.227′ N, 91°30.294′ E; 31 Jan. 2021; Nipu Kumar Das leg.; NZSI LM 1906 .</p><p>Description</p><p>Shell small, white, fresh shell possibly translucent; shell outline rather rounded in dorsal view; spire slightly elevated, low conical; body whorl rounded; protoconch low, finely granular, spiral striae lacking, consisting of ca 1.5 whorls; R1 of slightly under 1.5 whorls, with strong, sharp, but relatively low radial ribs that become more widely-spaced towards R2; last half whorl of R1 with ca 18 ribs, space between last two R1 ribs twice as large as distance between two ribs half whorl behind the end of R2; spiral striation absent; boundary between R1 and R2 conspicuous due to the sudden increase in rib density; R2 with ca 16 straight, sharp but low ribs; R2+R3 ca quarter whorl or slightly more; R3 and R2 of comparable length, boundary between R2 and R3 clearly visible due to a slight constriction; R3 with moderately long swelling that is situated closer to R3; R3 nearly smooth, with ca 3 weak ribs between swelling and outer peristome; aperture oblique to shell axis, rounded; boundary between inner and outer peristomes conspicuous; inner peristome strongly thickened, protruding and expanded; outer peristome thin, sharp, strongly reflected; umbilicus narrow, ca one fifth of shell width, rounded.</p><p>MEASUREMENTS. D: 1.9–2.5 mm, H: 1.4–1.9 mm.</p><p>Distribution</p><p>Known only from the Blue Mountain region.</p><p>Differential diagnosis</p><p>We have not found any similar species in the historical collection other than C. sculpturus, with which this species is sympatric. Chamalycaeus sculpturus is similar in shell shape and sculpture, and the ratios of regions. However, its peristome is dominated by the inner peristome, which is fringed and thickened, while the outer peristome is not expanded. In contrast, C. reflectus sp. nov. has a smooth (not fringed) inner, and a strongly expanded outer peristome. Furthermore, the upper apertural incision on C. crispatus is stronger and the R1 ribs (most clearly visible in the area just preceding R2) are denser. The peristome of C. sculptilis (Fig. 13A–D) is similar to that of C. reflectus, but that species is larger and has a strongly keeled body whorl.</p><p>Remarks</p><p>The single shell Ar44 (Fig. 10) is larger than the two type specimens and has a longer R2, while the sculpture and the morphology of the inner and outer peristomes of the type specimens and the Ar 44 specimen is identical. We need more extensive material to understand whether the observed differences represent an intraspecific diversity or differences between species. The two Ar 56 specimens are identical with the Ar 44 specimen but are strongly corroded. The single shell from Tripura (Fig. 11) is similar to the holotype, but has a less expanded outer peristome, and denser R1 and R2 ribs. Nevertheless, for the time being, we consider it conspecific with C. reflectus sp. nov.</p></div>	https://treatment.plazi.org/id/03CC1E69A323FFC5BAF4FB8DB591FCA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A326FFC4B914FEECB791FA91.text	03CC1E69A326FFC4B914FEECB791FA91.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus sculptilis (Benson 1856)	<div><p>Chamalycaeus sculptilis (Benson, 1856)</p><p>Fig. 13A–D</p><p>Alycaeus sculptilis Benson, 1856: 226–227 .</p><p>Alycaeus margarita Theobald in Hanley &amp; Theobald, 1874: 39, pl. 97 fig. 7 (renamed A. microstoma by Sowerby 1877).</p><p>Alycaeus microstoma Sowerby, 1877: pl. 4, species 28.</p><p>Alycaeus sculptilis – Sowerby 1877: pl. 4, species 32, figs a–b. — Godwin-Austen 1914: 398, 412, pl. 139 figs 7, 7a, pl. 155 fig. 8.</p><p>Alycaeus (Chamalycaeus) sculptilis – Kobelt 1902: 362. — Gude 1921: 233.</p><p>Chamalycaeus (Chamalycaeus) sculptilis – Ramakrishna et al. 2010: 55.</p><p>Chamalycaeus sculptilis – Páll-Gergely et al. 2020: 44–45. — Preece et al. 2022: 79, fig. 32c (neotype designation).</p><p>Diagnosis</p><p>Differs from all other species of this genus by the keeled body whorl and the finely beaded peristome. See also under Dicharax bhucarinatus Gittenberger &amp; Sherub Sherub, 2024 .</p><p>Type material examined</p><p>MYANMAR • neotype (designated by Preece et al. 2022: fig. 13a–d); Thyet Myo; Bens. coll.; UMZC I.102845.A • 2 shells; same data as for preceding; UMZC I.102845.B, I.102845.C</p><p>Additional material examined</p><p>MYANMAR • 1 shell; Pegu, Thayet-myo; “typical”, “aperture figured”; NHMUK 1906.4.4.70 .</p><p>Type locality</p><p>“raro ad Thyet-Mio prope fluvium Irawadi, non procul a finibus provinciae Burmanicae Britannicae” [rarely at Thyet-Mio near the river Irawadi, not far from the borders of British Burma] (translation by Preece et al. 2022).</p></div>	https://treatment.plazi.org/id/03CC1E69A326FFC4B914FEECB791FA91	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A326FFCBBAD3FAB2B4C4FD1F.text	03CC1E69A326FFCBBAD3FAB2B4C4FD1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus sculpturus (Godwin-Austen 1875)	<div><p>Chamalycaeus sculpturus (Godwin-Austen, 1875)</p><p>Figs 14–16</p><p>Alycaeus sculpturus Godwin-Austen 1875: 8, pl. 4 fig. 2.</p><p>Alycaeus crispatus var. makarsae Godwin-Austen, 1914: 372, pl. 158 fig. 13. Syn. nov.</p><p>Alycaeus (Alycaeus) sculpturus – Kobelt 1902: 351. — Ramakrishna et al. 2010: 51.</p><p>Alycaeus sculpturus – Godwin-Austen 1914: 398–399, pl. 145 figs 6, 6a–b. — Gude 1921: 218. Alycaeus (Dicharax) crispatus var. makarsae – Gude 1921: 243.</p><p>Dicharax (?) sculpturus – Páll-Gergely et al. 2020: 103.</p><p>Dicharax (?) crispatus makarsae – Páll-Gergely et al. 2020: 83.</p><p>Type material examined</p><p>INDIA • 1 syntype of A. sculpturus (Fig. 14A–E); Sikhami, NE Munipur; NHMUK 1903.7.1.2666 • 4 syntypes of A. sculpturus (Fig. 14F–J); Mungching, Munipur; NHMUK 1903.7.1.2667 • 8 syntypes of A. crispatus var. makarsae (Fig. 14K–O); Makarsa, Khasi; Godwin-Austen leg.; NHMUK 1903.7.1.2638 .</p><p>Additional material examined</p><p>INDIA – Mizoram • 1 empty shell (Fig. 15); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 68; NZSI LM1720 • 1 empty shell; same data as for preceding; Ar61; NZSI LM1721 • 2 complete shells (Fig. 16) + 2 broken shells; same data as for preceding; Ar74; NZSI LM1722 .</p><p>Type localities</p><p>“on the hill ranges from near Tellizo Peak to the eastward, and on Mungching Hill in Munipur” ( A. sculpturus); “Makarsa, N. Khasi Hills (or more correct, Maokarsa; the common Khasi prefix “Mao” meaning a stone)” ( A. crispatus var. makarsae).</p><p>Remarks</p><p>The type specimens of A. sculpturus and A. crispatus var. makarsae show no notable differences; therefore, the latter is a junior synonym of the former. The specimens collected in the Blue Mountain (Figs 15–16) are slightly smaller than typical C. sculpturus and A. crispatus var. makarsae shells and have multiple tiny knobs along the peristome but otherwise are very similar to those species. These knobs are probably homologous with the tips of triangle-shaped structures (termed chevrons in Gittenberger et al. 2024) in the aperture of typical C. sculpturus shells and many other alycaeine species (i.e., two tiny knobs of one shell are homologous with the two tips of a single triangle in the other). A similar case was observed in a Chinese species; typical Dicharax fimbriatus (Bavay &amp; Dautzenberg, 1912) shells had triangular thickenings along the peristome, while its synonym, Chamalycaeus plicilabris multidentatus Yen, 1939, had multiple tiny knobs along the peristome (Páll-Gergely et al. 2017).</p><p>This species was included in the genus Dicharax due to the absence of spiral striation (Páll-Gergely et al. 2020). However, re-examination of the syntypes of A. sculpturus and Alycaeus crispatus var. makarsae revealed some spiral striation on the teleoconch.</p></div>	https://treatment.plazi.org/id/03CC1E69A326FFCBBAD3FAB2B4C4FD1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A329FFCBB93EFD2CB4F3F800.text	03CC1E69A329FFCBB93EFD2CB4F3F800.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus solidus (Godwin-Austen 1914)	<div><p>Chamalycaeus solidus (Godwin-Austen, 1914)</p><p>Fig. 17</p><p>Alycaeus theobaldi var. solidus Godwin-Austen, 1914: 383–384, pl. 155 fig. 10.</p><p>Alycaeus kezamaensis Godwin-Austen, 1914: 393, pl. 149 fig. 1. Syn. nov.</p><p>Alycaeus (Dicharax) kezamaensis – Gude 1921: 258.</p><p>Alycaeus (Dicharax) theobaldi var. solida – Gude 1921: 273–274.</p><p>Chamalycaeus (Dicharax) kezamaensis – Ramakrishna et al. 2010: 62.</p><p>Dicharax kezamaensis – Páll-Gergely et al. 2020: 66.</p><p>Dicharax theobaldi solidus – Páll-Gergely et al. 2020: 75.</p><p>Diagnosis</p><p>This species can be distinguished from other species of this species group by the long and inflated R3 swelling. Chamalycaeus sylheticus is the most similar species; for comparisons, see under that species.</p><p>Type material examined</p><p>INDIA • 1 syntype of A. kezamaensis (Fig. 17F–J); Kezama, Naga Hills; Godwin-Austen coll.; NHMUK 1903.7.1.2556 • 4 syntypes of A. theobaldi var. solidus (Fig. 17A–E); Garo Hills; NHMUK 1903.7.1.2560 .</p><p>Type localities</p><p>“Kezama, Anghami-Naga Hills” ( A. kezamaensis); “Garo Hills” ( A. theobaldi solidus).</p><p>Remarks</p><p>Alycaeus kezamaensis is very slightly smaller than the syntypes of A. theobaldi solidus, but all other traits (shell and aperture shape, morphology and ratios of shell regions, sculpture) are identical. Therefore, they represent the same species. Alycaeus kezamaensis was introduced later in the same publication, therefore the valid name is Chamalycaeus solidus . Chamalycaeus solidus is a species of its own right, because it differs in important traits from D. theobaldi, namely the latter is more depressed, its R2 is longer, and the R3 swelling is lower and shorter (and they belong to different genera).</p><p>This species was included in the genus Dicharax due to the absence of spiral striation on the corroded syntypes of Alycaeus theobaldi var. solidus and Alycaeus kezamaensis (Páll-Gergely et al. 2020) . Nevertheless, the other conchological characters agree with species of this species group, and therefore, this species is reclassified in Chamalycaeus .</p></div>	https://treatment.plazi.org/id/03CC1E69A329FFCBB93EFD2CB4F3F800	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32FFFCDBACFFEECB7FBFBC1.text	03CC1E69A32FFFCDBACFFEECB7FBFBC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chamalycaeus sylheticus (Godwin-Austen 1914)	<div><p>Chamalycaeus sylheticus (Godwin-Austen, 1914)</p><p>Fig. 13F–I</p><p>Alycaeus sylheticus Godwin-Austen, 1914: 382, pl. 154 figs 4, 4a.</p><p>Alycaeus sylheticus – Gude 1921: 220.</p><p>Dicharax sylheticus – Páll-Gergely et al. 2020: 74; 2021: 23, fig. 16.</p><p>Type material examined</p><p>BANGLADESH • holotype (single shell mentioned in the original description: Fig. 13F–I); S Sylhet Hills; W. Channel leg.; NHMUK 1903.7.1.55 .</p><p>Type locality</p><p>“South Sylhet Hills”.</p><p>Differential diagnosis</p><p>The most similar species is C. solidus in terms of shell shape and sculpture, but that species is slightly larger, has a more elevated spire, and a more bulging R3 swelling.</p><p>Remarks</p><p>This species was included in the genus Dicharax in spite of the presence of spiral striation, because the R2 ribs were typical of that genus (Páll-Gergely et al. 2020). However, the critical revision of Himalayan species of Chamalycaeus and Dicharax revealed that C. sylheticus belongs to this species group of Chamalycaeus due to other conchological traits (colourless shells, strong and widely-spaced R1 ribs, upper apertural notch).</p></div>	https://treatment.plazi.org/id/03CC1E69A32FFFCDBACFFEECB7FBFBC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32FFFCDB998F8ACB30AF86C.text	03CC1E69A32FFFCDB998F8ACB30AF86C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx Godwin-Austen 1914	<div><p>Elongated Cycloryx</p><p>Remarks</p><p>The species belonging here usually have a conical or elongated conical shell and a thin peristome.</p></div>	https://treatment.plazi.org/id/03CC1E69A32FFFCDB998F8ACB30AF86C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32FFFCDB908F9B1B597F89E.text	03CC1E69A32FFFCDB908F9B1B597F89E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx Godwin-Austen 1914	<div><p>Genus Cycloryx Godwin-Austen, 1914</p><p>Alycaeus (Cycloryx) Godwin-Austen, 1914: 334 .</p><p>Type species</p><p>Cyclostoma constrictum Benson, 1851, by original designation.</p></div>	https://treatment.plazi.org/id/03CC1E69A32FFFCDB908F9B1B597F89E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32EFFCEB969F95CB7AAFD25.text	03CC1E69A32EFFCEB969F95CB7AAFD25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx bembex (Benson 1859)	<div><p>Cycloryx bembex (Benson, 1859)</p><p>Fig. 19A–H</p><p>Alycaeus bembex Benson, 1859: 178–179 .</p><p>Alycaeus bembex – Sowerby 1877: pl. 5, species 42. — Godwin-Austen 1884: pl. 51 fig. 5.</p><p>Alycaeus (Alycaeus) bembex – Kobelt 1902: 342.</p><p>Alycaeus (Cycloryx) bembex – Godwin-Austen 1914: 346–347, pl. 147 figs 1, 1a. — Gude 1921: 275– 276.</p><p>Cycloryx bembex – Ramakrishna et al. 2010: 70.</p><p>Pincerna bembex – Páll-Gergely et al. 2020: 170.</p><p>Diagnosis</p><p>This species can be distinguished from others due to the practically smooth, conical shell, and the thin peristome.</p><p>Type material examined</p><p>INDIA • 1 syntype (Fig. 19A–D); UMZC I.102730 • 6 possible syntypes (one of them separated in a different vial and marked with a “T”); Darjiling; NHMUK 1906.4.4.44 (Fig. 19E–H). The latter lot was not considered type by Preece et al. (2022) .</p><p>Type locality</p><p>“in valle Rungun”.</p></div>	https://treatment.plazi.org/id/03CC1E69A32EFFCEB969F95CB7AAFD25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32CFFCEB97DFD25B4B3F9E5.text	03CC1E69A32CFFCEB97DFD25B4B3F9E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx constrictus (Benson 1851)	<div><p>Cycloryx constrictus (Benson, 1851)</p><p>Fig. 20A–D</p><p>Cyclostoma constrictum Benson, 1851: 184–195 .</p><p>Alycaeus constrictus var. minor Benson, 1859: 181 .</p><p>Alycaeus constrictus – Sowerby 1877: pl. 5, species 41.</p><p>Alycaeus (Alycaeus) constrictus – Kobelt 1902: 343.</p><p>Alycaeus (Cycloryx) constrictus – Godwin-Austen 1914: 347–349, pl. 147 figs 4, 4a, pl. 154 figs 1, 1a. — Gude 1921: 277–278.</p><p>Alycaeus (Cycloryx) constrictus var. minor – Godwin-Austen 1914: 348.</p><p>Cycloryx constrictus – Ramakrishna et al. 2010: 70.</p><p>Pincerna constricta – Páll-Gergely 2017: 218, fig. 1a. — Páll-Gergely et al. 2020: 171.</p><p>Diagnosis</p><p>This species can be recognized based on the high conical shell shape, widely-spaced ribs, and thin peristome. R2 with several (ca 10) low ribs.</p><p>Type material examined</p><p>INDIA • holotype (see Fig. 20A–D and Preece et al. 2022); UMZC I.103745 .</p><p>Additional material examined</p><p>INDIA • 5 shells; Rungun Valley, Darjiling; NHMUK 1906.4.4.41.</p><p>Type locality</p><p>“ad Darjiling Himalayae Sikkimensis”.</p></div>	https://treatment.plazi.org/id/03CC1E69A32CFFCEB97DFD25B4B3F9E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A32CFFF0B91DF966B284FE01.text	03CC1E69A32CFFF0B91DF966B284FE01.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx costatus (Godwin-Austen 1914)	<div><p>Cycloryx costatus (Godwin-Austen, 1914)</p><p>Fig. 20E–I</p><p>Alycaeus (Cycloryx) costatus Godwin-Austen, 1914: 360–361, pl. 154 figs 2, 2a.</p><p>Alycaeus (Cycloryx) costatus – Gude 1921: 278.</p><p>Cycloryx costatus – Ramakrishna et al. 2010: 71.</p><p>Pincerna costata – Páll-Gergely 2017: 214. — Páll-Gergely et al. 2020: 171.</p><p>Type material examined</p><p>INDIA • 18 syntypes (Fig. 20E–I); Toruputu Peak, Dafla Hills, Dikrang Valley, Dafla; Godwin-Austen leg.; NHMUK 1903.7.1.2596 .</p><p>Type locality</p><p>“Dikrang Valley, Dafla Hills”.</p><p>Differential diagnosis</p><p>In size, shape and R1 sculpture it is similar to C. constrictus, but has only 4–5 R2 ribs.</p></div>	https://treatment.plazi.org/id/03CC1E69A32CFFF0B91DF966B284FE01	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A312FFF0B91EFE02B7E3F9A1.text	03CC1E69A312FFF0B91EFE02B7E3F9A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx elegans (Godwin-Austen 1914)	<div><p>Cycloryx elegans (Godwin-Austen, 1914)</p><p>Fig. 21A–E</p><p>Alycaeus (Cycloryx) elegans Godwin-Austen, 1914: 361, pl. 147 fig. 9.</p><p>Alycaeus (Cycloryx) khunhoensis Godwin-Austen, 1914: 403, pl. 147 fig. 8. syn. nov.</p><p>Alycaeus (Cycloryx) elegans – Gude 1921: 279.</p><p>Alycaeus (Cycloryx) khunhoensis – Gude 1921: 282.</p><p>Cycloryx elegans – Ramakrishna et al. 2010: 71.</p><p>Cycloryx khunhoensis – Ramakrishna et al. 2010: 72.</p><p>Pincerna elegans – Páll-Gergely 2017: 214. — Páll-Gergely et al. 2020: 175.</p><p>Pincerna khunhoensis – Páll-Gergely 2017: 214. — Páll-Gergely et al. 2020: 178.</p><p>Diagnosis</p><p>This species is characterized by a conical shell and dense ribbing on the entire surface. The peristome is thin, but the umbilicus is slightly covered by a reflected outer peristome.</p><p>Type material examined</p><p>INDIA • 1 syntype of C. elegans (Fig. 21A–E); Shengorh Pk, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7.1.2594 • 1 syntype of A. khunhoensis (Fig. 21F–J); Khunho Peak, Naga Hills; Godwin-Austen leg.; NHMUK 1903.7.1.2520 .</p><p>Type localities</p><p>“Shengorh Peak, Dafla Hills” ( C. elegans); “Khunho Peak, Naga Hills” ( C. khunhoensis).</p><p>Remarks</p><p>This species is similar to C. constrictus, but is characterized by a thinner peristome and finer ribbing. Cycloryx granum has similar sculpture, but it is more globular in shape.</p><p>Cycloryx khunhoensis has slightly less dense ribbing than C. elegans, which we interpret as part of the intraspecific variability.</p></div>	https://treatment.plazi.org/id/03CC1E69A312FFF0B91EFE02B7E3F9A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A312FFF2BAFAF9A1B267FDBB.text	03CC1E69A312FFF2BAFAF9A1B267FDBB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx pemaledai Gittenberger & Sherub Sherub 2022	<div><p>Cycloryx pemaledai Gittenberger &amp; Sherub Sherub, 2022</p><p>Fig. 19I–Q</p><p>Cycloryx pemaledai Gittenberger &amp; Sherub Sherub in Gittenberger et al., 2022: 76, figs 2, 9.</p><p>Cycloryx cf. pemaledai – Aravind &amp; Páll-Gergely 2023: 355, fig. 1.</p><p>Diagnosis</p><p>This species can be recognized based on the short R3.</p><p>Type material examined</p><p>Photos of the holotype (Fig. 19I–L) examined.</p><p>Additional material examined</p><p>INDIA • 2 shells (Fig. 19M–Q); North Sikkim, 2 km N of Lachen, towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=88.54439&amp;materialsCitation.latitude=27.74487" title="Search Plazi for locations around (long 88.54439/lat 27.74487)">Gurudongmar Lake</a>; 27.74487° N, 88.54439° E; 2749 m a.s.l.; 27 Nov. 2019; N.A. Aravind leg.; ATREE/2019/LS6001, ATREE/2019/LS6002 .</p><p>Type locality</p><p>“ Gasa Dzongkhag, 9 km S of Gasa, 2400 m a.s.l., 27°50′N 89°43′E ” (Bhutan).</p></div>	https://treatment.plazi.org/id/03CC1E69A312FFF2BAFAF9A1B267FDBB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF2B991F9C3B370F92C.text	03CC1E69A310FFF2B991F9C3B370F92C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx Godwin-Austen 1914	<div><p>Globular Cycloryx</p><p>Remarks</p><p>The species belonging here usually have a globular shell and a strongly expanded outer peristome.</p></div>	https://treatment.plazi.org/id/03CC1E69A310FFF2B991F9C3B370F92C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF2B91CFDCFB531F9BF.text	03CC1E69A310FFF2B91CFDCFB531F9BF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx tenellus (Godwin-Austen 1914)	<div><p>Cycloryx tenellus (Godwin-Austen, 1914)</p><p>Fig. 20J–N</p><p>Alycaeus (Cycloryx) tenellus Godwin-Austen, 1914: 417, pl. 155 figs 3–4, 4a.</p><p>Alycaeus (Cycloryx) tenellus – Gude 1921: 285.</p><p>Pincerna tenella – Páll-Gergely et al. 2020: 183.</p><p>Diagnosis</p><p>A species with somewhat questionable status (see Remarks). The only known shell is characterized by a somewhat conical-ovoid shell, smooth initial whorls, widely-spaced ribbing in the lower whorls and thin peristome.</p><p>Type material examined</p><p>MYANMAR • holotype (single shell mentioned in the original description: Fig. 20J–N); Shan Hills, in the same slide as N. 22; Fedden leg.; NHMUK 1906.5.5.87 .</p><p>Type locality</p><p>“ Shan States”.</p><p>Remarks</p><p>It is larger than any known species in this group. It is most similar to C. constrictus but has weaker ribbing (upper whorls smooth, although the holotype is strongly corroded). The more obvious separation between the inner and outer peristomes of this species compared to the figured syntype of C. constrictus is not a stable character, because in the other available lot of C. constrictus (NHMUK 1906.4.4.41) there are shells with similar peristome to that of the holotype of C. tenellus . A single shell is known of this species and very few of C. constrictus . If more material becomes available, it may happen that C. tenellus will be moved to the synonymy of C. constrictus .</p></div>	https://treatment.plazi.org/id/03CC1E69A310FFF2B91CFDCFB531F9BF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A310FFF4B935F95CB5C7FB5D.text	03CC1E69A310FFF4B935F95CB5C7FB5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx burrailensis (Godwin-Austen 1914)	<div><p>Cycloryx burrailensis (Godwin-Austen, 1914)</p><p>Fig. 22A–E</p><p>Alycaeus (Cycloryx) burrailensis Godwin-Austen, 1914: 403, pl. 147 figs 6, 6a.</p><p>Alycaeus (Cycloryx) burrailensis – Gude 1921: 276.</p><p>Cycloryx burrailensis – Ramakrishna et al. 2010: 70.</p><p>Pincerna burrailensis – Páll-Gergely et al. 2020: 170.</p><p>Type material examined</p><p>Lectotype (hereby selected: Fig. 22A–E)</p><p>INDIA • Japvo Peak, Naga Hills; elev. 9890 f; Godwin-Austen leg.; acc. no. 1830, NHMUK 1903.7.1.2591.1.</p><p>Paralectotypes INDIA • 5 specs; same data as for lectotype; NHMUK 1903.7.1.2591.2 to 1903.7.1.2591.6 • several specs in two vials, one of the vials with a single shell and a pink wool; Munipur side of the Burrail;</p><p>NHMUK 1903.7.1.2606.</p><p>Type localities</p><p>“Japvo Peak, Naga Hills, 9890 ft ”; “Munipur side of the Burrail”; Angaoluo Peak.</p><p>Differential diagnosis</p><p>Densely ribbed, similar to C. granum, but has wider spaces between R3 ribs.</p><p>Remarks</p><p>In the type collection in the same box we found the three vials (1: Japvo peak, labelled as “var. minor ”, 2591; 2: several shells with label “2606”; 3: single shell, also with label “2606” and a pink wool [which usually indicates that it is the “ type ”]). Besides the size, the difference between shells from the two localities is that R3 is much more densely ribbed in “var. minor “ (from Japvo Peak).</p><p>In the original description, the type locality was given as “Japvo Peak, Naga Hills, 9890 ft ”, and it is later added that “obtained also on the Munipur side of the Burrail, No. 2606 B.M.; and Angaoluo Peak, No. 2749 B.M.”. One understanding of Godwin-Austen’s concept is to accept only the lot from Japvo Peak as syntypes, and none of the shells from the other two sites. However, in that case specimens from Burrail would be excluded from the type series, which would be strange for a species named “ burrailensis ”. Moreover, although not mentioned in the original description, the shells from Japvo Peak were labelled as “var. minor, 2591”, suggesting, that the ones from Burrail should be considered “typical”. Lastly, the description states that there is a “distant fine costulation between sutural tube and aperture”, which unambiguously refers to the larger form from Burrail (NHMUK 1903.7.1.2606). Therefore, we accept shells belonging to all three lots as syntypes.</p><p>The single shell from Angaoluo Peak was also found in the general collection, and it is similar to the ones from Burrail in size and formation of the aperture, but the R1 ribbing is approximately half as dense.</p><p>Since the syntypes were collected at three different localities, and are somewhat different in shell traits, we hereby select a lectotype to avoid further nomenclatural confusion.</p></div>	https://treatment.plazi.org/id/03CC1E69A310FFF4B935F95CB5C7FB5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A316FFF6B936FAEDB70EFE26.text	03CC1E69A316FFF6B936FAEDB70EFE26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx dihingensis (Godwin-Austen 1914)	<div><p>Cycloryx dihingensis (Godwin-Austen, 1914)</p><p>Fig. 23A–E</p><p>Alycaeus (Cycloryx) graphicus var. dihingensis Godwin-Austen, 1914: 363–364, 404, pl. 146 figs 6, 6a.</p><p>Alycaeus (Cycloryx) graphicus var. dihingensis – Gude 1921: 281–282.</p><p>Pincerna graphica dihingensis – Páll-Gergely et al. 2020: 178.</p><p>Diagnosis</p><p>This species can be recognized by its large size, the reflected outer peristome in the direction of the umbilicus, and the thick peristome (protruding inner peristome). See also under Cycloryx rohiniae sp. nov.</p><p>Type material examined</p><p>INDIA • 2 syntypes (Fig. 23A–E); Sadia, Assam; NHMUK 1903.7.1.2516 .</p><p>Type locality</p><p>“Sadia, Assam, 350 ft ”.</p><p>Remarks</p><p>Similar to C. major, but it is larger and has a thicker peristome with more protruding inner peristome. Similar in size to A. otiphorus, but has a more globular shell, stronger ribbing on the apical whorls, and ribs on R3.</p></div>	https://treatment.plazi.org/id/03CC1E69A316FFF6B936FAEDB70EFE26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A314FFF6BA84FE5AB43BFCDA.text	03CC1E69A314FFF6BA84FE5AB43BFCDA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx globhutanus Gittenberger & Sherub Sherub 2022	<div><p>Cycloryx globhutanus Gittenberger &amp; Sherub Sherub, 2022</p><p>Fig. 24A–D</p><p>Cycloryx globhutanus Gittenberger &amp; Sherub Sherub in Gittenberger et al., 2022: 76, figs 2, 4.</p><p>Diagnosis</p><p>This species can be recognized based on the very widely-spaced R1 ribs and the smooth R2 with numerous slender microtunnels.</p><p>Type locality</p><p>“ Samdrup Jongkhar Dzongkhag, 9 km N of Samdrup Jongkhar, 1008 m a.s.l., warm broadleaf forest, 26°52′N 91°29′E ” (Bhutan).</p></div>	https://treatment.plazi.org/id/03CC1E69A314FFF6BA84FE5AB43BFCDA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A314FFF9B91DFC6FB599FE05.text	03CC1E69A314FFF9B91DFC6FB599FE05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx granum (Godwin-Austen 1893)	<div><p>Cycloryx granum (Godwin-Austen, 1893)</p><p>Fig. 22F–O</p><p>Alycaeus otiphorus var. Godwin-Austen, 1871: 93, pl. 5 fig. 6.</p><p>Alycaeus (Dioryx) granum Godwin-Austen, 1893: 593–594 .</p><p>Alycaeus (Cycloryx) multicostulatus Godwin-Austen, 1914: 404, pl. 147 fig. 7. Syn. nov.</p><p>Alycaeus (Cycloryx) burrailensis Godwin-Austen, 1914: 403 (in part).</p><p>Alycaeus (Dioryx) granum – Godwin-Austen 1897: 4–5, pl. 63 fig. 6.</p><p>Alycaeus (Alycaeus) granum – Kobelt 1902: 345.</p><p>Alycaeus (Cycloryx) granum – Godwin-Austen 1914: 364. — Gude 1921: 279–280. Alycaeus (Cycloryx) multicostulatus – Gude 1921: 282–283.</p><p>Cycloryx granum – Ramakrishna et al. 2010: 71.</p><p>Cycloryx multicostulatus – Ramakrishna et al. 2010: 73.</p><p>Pincerna granum – Páll-Gergely et al. 2020: 177.</p><p>Pincerna multicostulata – Páll-Gergely et al. 2020: 181.</p><p>Diagnosis</p><p>Shell globular, ribs dense on the entire shell.</p><p>Type material examined</p><p>INDIA • 1 syntype of A. granum (Fig. 22F–J); Margarita, near Sadia, Assam; Godwin-Austen coll.; NHMUK 1903.7.1.2511 • 1 syntype of A. multicostulatus (Fig. 22K–O); Munipur, head of Lanier Rr; Godwin-Austen leg.; NHMUK 1903.7.1.2557 .</p><p>Type localities</p><p>“Margarita, foot of Eastern Naga Hills” ( A. granum); “Head of the Lanier River, Lahupa Naga Hills, N.E. Manipur ” ( A. multicostulatus).</p><p>Remarks</p><p>Types of C. granum are much smaller than C. multicostulatus, but there is no difference between those shells other than shell size. Specimens from the Japvo Peak, Naga Hills (labelled as “ A. burrailensis var. minor ”, see also A. burrailensis) are intermediate in size between C. granum and C. multicostulatus . Therefore, we treat the latter as a junior synonym of the former.</p></div>	https://treatment.plazi.org/id/03CC1E69A314FFF9B91DFC6FB599FE05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31BFFF8BAA1FE06B5F4FE7F.text	03CC1E69A31BFFF8BAA1FE06B5F4FE7F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx haumbiclausus Gittenberger & Choki Gyeltshen 2022	<div><p>Cycloryx haumbiclausus Gittenberger &amp; Choki Gyeltshen, 2022</p><p>Fig. 24E–H</p><p>Cycloryx haumbiclausus Gittenberger &amp; Choki Gyeltshen in Gittenberger et al., 2022: 78, figs 1, 6.</p><p>Diagnosis</p><p>This species can be recognized by the widely-spaced R1 ribs and the smooth R2 with numerous slender microtunnels.</p><p>Type locality</p><p>“ Haa Dzongkhag, Gakiling, Rangtse Nye, 27°05′N 89°09′E, 890 m a.s.l.” (Bhutan).</p><p>Differential diagnosis</p><p>Differs from C. sajumbiclausus by the presence of spiral striation.</p></div>	https://treatment.plazi.org/id/03CC1E69A31BFFF8BAA1FE06B5F4FE7F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFF8B923FE0CB510F901.text	03CC1E69A31AFFF8B923FE0CB510F901.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx graphicus (W. T. Blanford 1862)	<div><p>Cycloryx graphicus (W.T. Blanford, 1862)</p><p>Fig. 25</p><p>Alycaeus graphicus W.T. Blanford, 1862: 137–138 .</p><p>Alycaeus (Dioryx) graphicus var. minor Godwin-Austen, 1874: 149, pl. 3 fig. 4.</p><p>Alycaeus (Cycloryx) difficilis Godwin-Austen, 1914: 415–416, pl. 155 figs 2, 2a. Syn. nov.</p><p>Alycaeus graphicus var. – Theobald 1870: 398, pl. 18 fig. 3.</p><p>Alycaeus graphicus – Sowerby 1877: pl. 4, species 34.</p><p>Alycaeus (Alycaeus) graphicus – Kobelt 1902: 345.</p><p>Alycaeus (Cycloryx) graphicus – Godwin-Austen 1914: 361, 419–420, pl. 146 figs 1, 1a–b. — Gude 1921: 280–281.</p><p>Alycaeus (Cycloryx) graphicus var. minor – Godwin-Austen 1914: 403, pl. 144 figs 9, 9a.</p><p>Alycaeus (Cycloryx) difficilis – Gude 1921: 278–279.</p><p>Cycloryx graphicus – Ramakrishna et al. 2010: 72.</p><p>Pincerna graphica – Páll-Gergely et al. 2020: 177.</p><p>Pincerna difficilis – Páll-Gergely et al. 2020: 175.</p><p>Diagnosis</p><p>This species can be recognized based on the dense R1 ribs, the R3 with widely-spaced ribs, the thick peristome (protruding inner peristome) and that the outer peristome is not reflected abruptly towards umbilicus (i.e., it does not form a sharp angle below the umbilicus).</p><p>Type material examined</p><p>MYANMAR • 4 syntypes of A. graphicus (Fig. 25A–E); Moditoung, Tongoop Pass, Arakan; NHMUK 1906.4.4.42 • 2 syntypes of A. difficilis (Fig. 25F–J); Shan States; Fedden leg.; NHMUK 1906.5.5.22 .</p><p>Type localities</p><p>“in montibus Arakanensibus provinciam Burmanam Pegu ad Arakan secernentibus” ( A. graphicus); “ Shan Hills” ( A. difficilis).</p><p>Remarks</p><p>Alycaeus graphicus and A. difficilis are identical in all shell traits (except minor difference in rib density), therefore they are treated as synonyms. The ribs of this species are more widely-spaced than those of C. major, although no other differences were found.</p></div>	https://treatment.plazi.org/id/03CC1E69A31AFFF8B923FE0CB510F901	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31AFFFCB910F902B4C5FD2E.text	03CC1E69A31AFFFCB910F902B4C5FD2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx major (Godwin-Austen 1893)	<div><p>Cycloryx major (Godwin-Austen, 1893)</p><p>Figs 26, 27F–J</p><p>Alycaeus (Dioryx) granum var. major Godwin-Austen, 1893: 594 .</p><p>Alycaeus (Cycloryx) mangutensis Godwin-Austen, 1914: 377–378, pl. 146 figs 5, 5a.</p><p>Alycaeus (Cycloryx) graphicus var. variabilis Godwin-Austen, 1914: 403–404, pl. 146 fig. 4. Syn. nov. Alycaeus (Cycloryx) thompsoni Godwin-Austen, 1914: 404, pl. 146 figs 3, 3a. Syn. nov.</p><p>Alycaeus (Cycloryx) mangutensis – Gude 1921: 282.</p><p>Alycaeus (Cycloryx) thompsoni – Gude 1921: 285.</p><p>Cycloryx mangutensis – Ramakrishna et al. 2010: 72.</p><p>Cycloryx thompsoni – Ramakrishna et al. 2010: 74.</p><p>Pincerna major – Páll-Gergely et al. 2020: 179.</p><p>Pincerna graphica variabilis – Páll-Gergely et al. 2020: 179.</p><p>Pincerna thompsoni – Páll-Gergely et al. 2020: 183.</p><p>Diagnosis</p><p>This species can be recognized by the dense R1 ribbing, the regular ribbing on R3, and the angle below the umbilicus, formed by the reflected outer peristome.</p><p>Type material examined</p><p>INDIA • 8 syntypes of A. major (Fig. 26A–E); Mangat Valley, Jiantia Hills; Godwin-Austen leg.; NHMUK 1903.7.1.2518 • 17 syntypes of A. graphicus var. variabilis (Fig. 26F–J); Lhota Naga; NHMUK 1903.7.1.2607 • 1 syntype of A. thompsoni (Fig. 26K–O); Munipur; Godwin-Austen coll.; NHMUK 1903.7.1.2550 .</p><p>Additional material examined</p><p>INDIA – Mizoram • 1 empty shell (Fig. 27F–J); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 53; NZSI LM1716 .</p><p>Type localities</p><p>“Mangut Valley, Jaintia Hills” ( A. major); “Lhota Naga ” ( A. graphicus var. variabilis); “Munipur” ( A. thompsoni).</p><p>Differential diagnosis</p><p>Cycloryx graphicus has more widely-spaced ribs than C. major, and the outer peristome does not form a sharp angle below the umbilicus in that species.</p><p>Remarks</p><p>The specimen from Blue Mountain is practically identical to the type specimens of P. major in all shell characters (size, shape, rib density). Specimens labelled as “ A. graphicus var.” from the Burroi Gorge, Dafla Hills (NHMUK 1903.07.1.2598) are also identical, although most specimens are larger.</p><p>Although slightly larger, A. thompsoni and A. graphicus var. variabilis also have similar traits. Therefore, they are treated as synonyms of Cycloryx major .</p></div>	https://treatment.plazi.org/id/03CC1E69A31AFFFCB910F902B4C5FD2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31EFFFCB907FD52B335F836.text	03CC1E69A31EFFFCB907FD52B335F836.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx margarita (Theobald 1874)	<div><p>Cycloryx margarita (Theobald, 1874)</p><p>Fig. 28</p><p>Alycaeus margarita Theobald in Hanley &amp; Theobald, 1874: 39, pl. 95 fig. 10.</p><p>Alycaeus (Cycloryx) graphiarius Godwin-Austen, 1914: 416, pl. 146 figs 7, 7a. Syn. nov.</p><p>Alycaeus margaritus – Sowerby 1877: pl. 1, species 5.</p><p>Alycaeus (Alycaeus) margarita – Kobelt 1902: 347.</p><p>Alycaeus (Cycloryx) margarita – Godwin-Austen 1914: 416.</p><p>Alycaeus margarita – Gude 1921: 211–212.</p><p>Alycaeus (Cycloryx) graphiarius – Gude 1921: 280.</p><p>Pincerna margarita – Páll-Gergely et al. 2020: 180.</p><p>Pincerna graphiaria – Páll-Gergely et al. 2020: 177.</p><p>Diagnosis</p><p>This species is characterized by a globular shell, and widely-spaced, strong ribs on the entire shell surface.</p><p>Type material examined</p><p>MYANMAR • 1 syntype of A. margarita (Fig. 28A–D); Upper Salwin; NHMUK 1888.12.4.892 • 1 syntype of A. margarita; Upper Salwin; NHMUK 1888.12.4.893 • 3 syntypes of A. graphiarius (Fig. 28E–I, mentioned as 1 syntype in Páll-Gergely et al. 2020); Shan States; Fedden leg.; Blanford coll.; NHMUK 1906.4.4.21 (two of them are A. margarita, the third shell belongs to another species, see remarks) .</p><p>Type locality</p><p>“ Shan Provinces” ( A. margarita); not specified for A. graphiarius .</p><p>Remarks</p><p>Two syntypes of A. graphiarius agreed with the original description (Godwin-Austen, 1914) by having widely-spaced ribs, but the third shell had much denser ribbing, and therefore, it clearly belongs to a different species.</p><p>The two sparsely ribbed syntypes of A. graphiarius are identical to those of A. margarita in all shell characters (size, shape, density of ribs), therefore the former is a junior synonym of the latter.</p></div>	https://treatment.plazi.org/id/03CC1E69A31EFFFCB907FD52B335F836	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A31DFFE1B975F9C5B2A5FD95.text	03CC1E69A31DFFE1B975F9C5B2A5FD95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx otiphorus (Benson 1859)	<div><p>Cycloryx otiphorus (Benson, 1859)</p><p>Fig. 29</p><p>Alycaeus otiphorus Benson, 1859: 178–179 .</p><p>Alycaeus (Cycloryx) summus Godwin-Austen, 1914: 349–350, pl. 147 figs 3, 3a. Syn. nov.</p><p>Alycaeus otiphorus – Sowerby 1877: pl. 4, species 30.</p><p>Alycaeus (Alycaeus) otiphorus – Kobelt 1902: 347.</p><p>Alycaeus (Cycloryx) otiphorus – Godwin-Austen 1914: 349, pl. 147 figs 2, 2a–b. — Gude 1921: 283. Alycaeus (Cycloryx) summus – Gude 1921: 284–285.</p><p>Cycloryx otiphorus – Ramakrishna et al. 2010: 73.</p><p>Pincerna otiphorus – Páll-Gergely et al. 2020: 182.</p><p>Pincerna summa – Páll-Gergely et al. 2020: 182.</p><p>Diagnosis</p><p>This species can be recognized based on the large, conical shell, and the weak sculpture. See also under Cycloryx rohiniae sp. nov.</p><p>Type material examined</p><p>INDIA • 1 syntype of A. otiphorus (see Preece et al. 2022: fig. 33e); no locality data; UMZC I.102555 • 12 syntypes of A. summus (Fig. 29F–J); Rechila Pk, W. Bhutan; Godwin-Austen leg.; NHMUK 1903.7.1.2573 .</p><p>Additional material examined</p><p>INDIA • single shell from the R. McAndrew collection (Fig. 29A–D); UMZC I.102560 • 1 shell of A. otiphorus; Darjiling; “compared with typical sp in Museum Cambridge”; NHMUK 1903.7.1.2565.</p><p>Type localities</p><p>“ad Pankabari (1000 ped. alt.) et in valle Rungun (4000 ped.) prope Darjiling Himalayanum” ( A. otiphorus); “Rechila Peak, Western Bhutan ” ( A. summus).</p><p>Remarks</p><p>Alycaeus summus is identical to A. otiphorus, therefore the former is a junior synonym of the latter. Unlike other members of this species group, A. otiphorus has a rather conical shell (instead of conical-globular), but it is classified here due to the expanded and reflected outer peristome.</p></div>	https://treatment.plazi.org/id/03CC1E69A31DFFE1B975F9C5B2A5FD95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE1BAC3FDB5B26DFAA6.text	03CC1E69A303FFE1BAC3FDB5B26DFAA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx paucicostatus (Godwin-Austen 1914)	<div><p>Cycloryx paucicostatus (Godwin-Austen, 1914)</p><p>Fig. 23F–J</p><p>Alycaeus paucicostatus Godwin-Austen, 1914: 361, pl. 147 figs 5, 5a.</p><p>Alycaeus (Cycloryx) paucicostatus – Gude 1921: 284.</p><p>Cycloryx paucicostatus – Ramakrishna et al. 2010: 73.</p><p>Pincerna paucicostata – Páll-Gergely et al. 2020: 182.</p><p>Diagnosis</p><p>This species can be distinguished from all congeners based on the lamella-like ribs on R3.</p><p>Type material examined</p><p>INDIA • 2 syntypes (Fig. 23F–J); Toruputu Pk, Dafla Hills; NHMUK 1903.7.1.2595 .</p><p>Type locality</p><p>“Torúpútú Peak, Dafla Hills”.</p><p>Differential diagnosis</p><p>The most similar species is C. burrailensis, but that species has denser R1 ribs.</p></div>	https://treatment.plazi.org/id/03CC1E69A303FFE1BAC3FDB5B26DFAA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A303FFE0BAD1FADBB348FB7A.text	03CC1E69A303FFE0BAD1FADBB348FB7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx rohiniae Pall-Gergely & Aravind 2025	<div><p>Cycloryx rohiniae Páll-Gergely &amp; Aravind sp. nov.</p><p>urn:lsid:zoobank.org:act: 80122B22-6AEE-4698-8E7A-62068B3CAE76</p><p>Fig. 27A–E</p><p>Diagnosis</p><p>A large species of Cycloryx with smooth R3.</p><p>Etymology</p><p>The new species is named after Ms Rohini Nilekani, author, environmentalist and acclaimed philanthropist.</p><p>Type material examined</p><p>Holotype</p><p>INDIA – Mizoram • empty shell (D: 3.7 mm, H: 4.1 mm, Fig. 27A–E); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 37; NZSI LM1717.</p><p>Paratype</p><p>INDIA – Mizoram • 1 empty shell; same data as for holotype; Ar 34; NZSI LM1718 • 1 empty shell; same data as for holotype; Ar 35; NZSI LM1719 .</p><p>Description</p><p>Shell large for the genus, all available shells slightly corroded, original colour may be greenish-yellowish, semitranslucent; shell conical or conical-globular, height and width approximately equivalent; body whorl rounded; protoconch smooth, glossy, consisting of 1.5 or slightly fewer whorls; R1 of ca 2–2.25 whorls, regularly ribbed, rib density gradually decreases towards end of R2 (last half whorl with ca 16 ribs); ribs thread-like, low; fine, dense spiral striation discernible between ribs; boundary between R1 and R2 not conspicuous, but clearly visible due to increase in rib density (R2 ribs twice as dense as ribs at the end of R1); R2 practically smooth, its “ribs” (6–7 in number) are not elevated from the surface, represented as slender, light stripes; sutural tube short, cylindrical; R2+R3 ca quarter whorl, R3 ca 3 times as long as R2; boundary between R2 and R3 clearly visible due to a shallow constriction; R3 with a low central, long swelling, smooth, only fine growth lines visible but no ribs; aperture slightly oblique to shell axis, rounded; inner and outer peristomes clearly separated; inner peristome slightly protruding and expanded; outer peristome strongly expanded, sharp, and reflected in direction of umbilicus; umbilicus open, narrow, the reflected outer peristome curves into umbilicus, terminating at its centre.</p><p>OPERCULUM. A single operculum of the Ar35 shells was examined. The inner side is smooth, without central swelling, outer side multispiral, without elevated lamina.</p><p>MEASUREMENTS. D: 3.7 mm, H: 3.5–4.1 mm.</p><p>Distribution</p><p>Known only from the Blue Mountain region.</p><p>Differential diagnosis</p><p>The only species with similar size and lack of R3 ribs is C. otiphorus, which has a more pointed (more triangular) shell, and has denser ribs on the apical part. Cycloryx dihingensis, which is similar in shell shape, has strong R3 ribs. All these species inhabit Assam, which geographically is distant from Mizoram.</p></div>	https://treatment.plazi.org/id/03CC1E69A303FFE0BAD1FADBB348FB7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A302FFE0BAA6FB0FB71DF92E.text	03CC1E69A302FFE0BAA6FB0FB71DF92E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cycloryx sajumbiclausus Gittenberger & Choki Gyeltshen 2022	<div><p>Cycloryx sajumbiclausus Gittenberger &amp; Choki Gyeltshen, 2022</p><p>Fig. 24I–L</p><p>Cycloryx sajumbiclausus Gittenberger &amp; Choki Gyeltshen in Gittenberger et al., 2022: 78, figs 1, 5.</p><p>Diagnosis</p><p>This species can be recognized based on the widely-spaced R1 ribs and the smooth R2 with numerous slender microtunnels. It differs from C. haumbiclausus based on the absence of spiral striation.</p><p>Type locality</p><p>“ Samdrup Jongkhar Dzongkhag, 4 km NW of Samdrup Jongkhar, 260 m a.s.l., 26°49′N 91°28′E ” (Bhutan).</p></div>	https://treatment.plazi.org/id/03CC1E69A302FFE0BAA6FB0FB71DF92E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A302FFE0B930F95CB20DF877.text	03CC1E69A302FFE0B930F95CB20DF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Genus Dicharax Kobelt &amp; Möllendorff, 1900</p><p>Charax Benson, 1859: 177.</p><p>Dicharax Kobelt &amp; Möllendorff, 1900: 186 (replacement name for Charax Benson, 1859, non Charax Scopoli, 1777 [Pisces]).</p><p>Type species</p><p>Alycaeus hebes Benson, 1857, subsequent designation by Gude (1921: 236).</p></div>	https://treatment.plazi.org/id/03CC1E69A302FFE0B930F95CB20DF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3BA71FD67B552FB2D.text	03CC1E69A301FFE3BA71FD67B552FB2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax aspidentatus Gittenberger, Choki Gyeltshen & Sherub Sherub 2024	<div><p>Dicharax aspidentatus Gittenberger, Choki Gyeltshen &amp; Sherub Sherub, 2024</p><p>Fig. 30A–D</p><p>Dicharax aspidentatus Gittenberger, Choki Gyeltshen &amp; Sherub Sherub in Gittenberger et al., 2024: 199, figs 18, 21, 45.</p><p>Type locality</p><p>“between Duenmang Tsachu and Gongphu Zero Point, 24 km SE of Zhemgang, 335 m a.s.l., 27°02’N 90°48’E, scree in warm broadleaf forest” (Bhutan).</p><p>Differential diagnosis</p><p>Probably the most similar species is D. rugosus due to the similar shell and aperture shape as well as rib density. However, that species has a shorter distance between the inner and outer peristomes and has a wider umbilicus. For the differences with other Bhutanese species see Gittenberger et al. (2024). See also under Chamalycaeus asaluensis and D. umbiclausus .</p></div>	https://treatment.plazi.org/id/03CC1E69A301FFE3BA71FD67B552FB2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3BACBFB5DB5BBF886.text	03CC1E69A301FFE3BACBFB5DB5BBF886.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax floridus Pall-Gergely & Hunyadi 2022	<div><p>Dicharax floridus Páll-Gergely &amp; Hunyadi, 2022</p><p>Fig. 31A–E</p><p>Dicharax floridus Páll-Gergely &amp; Hunyadi, 2022: 95, fig. 1.</p><p>Type material examined</p><p>MYANMAR • holotype (Fig. 31A–E); Shan State, Hopong centre 7.4 km towards Namsang, rd. #4 N 5 km, near Parpant Cave; 20°850.9630′ N, 97°814.2670′ E; 1170 m a.s.l.; 6 Oct. 2018; A. Hunyadi, K. Okubo and J.U. Otani, leg.; HA (to be deposited in the HNHM) .</p><p>Type locality</p><p>“ Myanmar, Shan State, Hopong centre 7.4 km towards Namsang, rd. #4 N 5 km, near Parpant Cave, 20°850.9630’N, 97°814.2670’E, 1170 m a.s.l.”</p><p>Differential diagnosis</p><p>This species differs from its congeners in this species group by the strongly expanded outer peristome and the smaller distance between the two peristomes. Some species of Chamalycaeus are also similar to Dicharax floridus, but their outer peristome is less expanded.</p></div>	https://treatment.plazi.org/id/03CC1E69A301FFE3BACBFB5DB5BBF886	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE3B9BFFEECB216FDE3.text	03CC1E69A301FFE3B9BFFEECB216FDE3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 1</p><p>Remarks</p><p>This species group is characterized by a usually strongly ribbed shell, a fringed aperture, and most species (exception: D. floridus) have a large space between the inner and outer peristomes.</p><p>The R3 swelling of D. obscurus is similar to that of the species belonging to the species group 5, but it is included in this group due to the fringed aperture and the large space between the two peristomes.</p><p>See also remarks under Dicharax species group 2 regarding D. bicrenatus .</p></div>	https://treatment.plazi.org/id/03CC1E69A301FFE3B9BFFEECB216FDE3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A301FFE7B92FF8BBB50DFB64.text	03CC1E69A301FFE7B92FF8BBB50DFB64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax jaintiacus (Godwin-Austen 1871)	<div><p>Dicharax jaintiacus (Godwin-Austen, 1871)</p><p>Fig. 32</p><p>Alycaeus Jaintiacus Godwin-Austen, 1871: 92–93, pl. 5 fig. 3.</p><p>Alycaeus serratus Godwin-Austen, 1874: 148–149, pl. 3 fig. 6. Syn. nov.</p><p>Alycaeus jaintiacus var. crassus Godwin-Austen, 1914: 375, pl. 137 figs 5, 5a. Syn. nov.</p><p>Alycaeus (Alycaeus) serratus – Kobelt 1902: 351. — Ramakrishna et al. 2010: 51.</p><p>Alycaeus (Dicharax) jaintiacus – Kobelt 1902: 372. — Gude 1921: 256.</p><p>Alycaeus jaintiacus – Godwin-Austen 1914: 375, pl. 143 figs 3, 3a–b.</p><p>Alycaeus serratus – Godwin-Austen 1914: 400, pl. 144 figs 6, 6a–b. — Gude 1921: 219. Chamalycaeus (Dicharax) jaintiacus – Ramakrishna et al. 2010: 61.</p><p>Alycaeus (Dicharax) jaintiacus var. crassa – Gude 1921: 256–257.</p><p>Dicharax jaintiacus – Páll-Gergely et al. 2020: 65.</p><p>Dicharax jaintiacus crassus – Páll-Gergely et al. 2020: 65.</p><p>Dicharax (?) serratus – Páll-Gergely et al. 2020: 103.</p><p>Diagnosis</p><p>The elevated spire, rather smooth R1 (with the exception of strong, widely-spaced ribs along the suture), the long R3 swelling with straight anterior part, and the strongly fringed aperture distinguish this species from its congeners.</p><p>Type material examined</p><p>INDIA • 14 syntypes of A. jaintiacus (Fig. 32A–E); Nongjinghi, Jaintia Hills; Godwin-Austen leg.; NHMUK 1903.7.1.2686 • 4 syntypes of A. jaintiacus var. crassus in 2 vials (Fig. 32K–O); Nonjinghi, Jaintia; Godwin-Austen coll.; NHMUK 1903.7.1.2752 • 1 syntype of A. serratus (Fig. 32F–J); Laisen Valley, Jiantia Hills; NHMUK 1903.7.1.2487 .</p><p>Type localities</p><p>“in Nongjinghi, Jaintia” ( A. jaintiacus); “in Nongjinghi, Jaintia, 4563 feet ” ( A. jaintiacus var. crassus); “Laisen Trigl. station, Munipur Hills” ( A. serratus).</p><p>Differential diagnosis</p><p>The most similar is D. obscurus; for comparisons, see under that species. Species of the Dicharax species group 7 ( D. chennelli and similar species) resemble D. jaintiacus, but the distance between their two peristomes is smaller, their R3 swelling is more inflated and usually have R3 depressions. In contrast, D. jaintiacus has a less inflated R3 without depressions.</p><p>Remarks</p><p>Dicharax jaintiacus var. crassus is practically identical to the nominotypical form, therefore it is treated here as a junior synonym. Alycaeus serratus is slightly smaller and has a shallower R3 swelling, but all other shell characters are very similar, and they were described from the same geographic region; therefore, Alycaeus serratus is hereby synonymised with D. jaintiacus . Alycaeus serratus was not compared with A. jaintiacus in its original description.</p></div>	https://treatment.plazi.org/id/03CC1E69A301FFE7B92FF8BBB50DFB64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A305FFE7B92EFAE5B20EF85B.text	03CC1E69A305FFE7B92EFAE5B20EF85B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax muspratti (Godwin-Austen 1914)	<div><p>Dicharax muspratti (Godwin-Austen, 1914)</p><p>Fig. 33A–E</p><p>Alycaeus muspratti Godwin-Austen, 1914: 396, pl. 148 fig. 1.</p><p>Alycaeus (Raptomphalus) muspratti – Gude 1921: 289.</p><p>Chamalycaeus (Raptomphalus) muspratti – Ramakrishna et al. 2010: 69.</p><p>Dicharax (?) muspratti – Páll-Gergely et al. 2020: 97.</p><p>Type material examined</p><p>INDIA • 16 syntypes (Fig. 33A–E); E. Naga; R.H. Beddome coll.; NHMUK 1912.4.16.273 .</p><p>Type locality</p><p>“Eastern Naga ”.</p><p>Differential diagnosis</p><p>Dicharax dohertyi (of which no specimens were available, see under D. dohertyi) and D. rugosus are similar in sculpture and aperture formation, but they have a higher spire. Dicharax trashidentatus is very similar to D. muspratti in shell and aperture shape and shell sculpture, but it also has a higher spire and denser R1 ribs. Dicharax rugosus is also similar due to the strong R1 ribs, the large distance between the inner and outer peristomes and the fringed aperture, but it has a higher spire.</p></div>	https://treatment.plazi.org/id/03CC1E69A305FFE7B92EFAE5B20EF85B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A304FFE9B92AFEECB347FE05.text	03CC1E69A304FFE9B92AFEECB347FE05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax obscurus (Godwin-Austen 1914)	<div><p>Dicharax obscurus (Godwin-Austen, 1914)</p><p>Fig. 31F–J</p><p>Alycaeus obscurus Godwin-Austen, 1914: 378–379, pl. 154 figs 9, 9a.</p><p>Alycaeus (Dicharax) obscurus – Gude 1921: 263.</p><p>Chamalycaeus (Dicharax) obscurus – Ramakrishna et al. 2010: 64.</p><p>Dicharax (?) obscurus – Páll-Gergely et al. 2020: 100.</p><p>Type material examined</p><p>INDIA • 1 syntype (Fig. 31F–J); Cherra Poonje; Godwin-Austen leg.; NHMUK 1913.3.16.8 .</p><p>Type locality</p><p>“Cherra Poonje”.</p><p>Differential diagnosis</p><p>The most similar species in this species group is D. jaintiacus, which has a more globular shell, possesses more widely-spaced R1 ribs, and a less expanded outer peristome. Species of the Dicharax species group 7 are similar to D. obscurus in the formation of the R3 swelling, but the distance between their two peristomes is smaller and their R3 swelling is more inflated and usually have depressions and/or grooves.</p></div>	https://treatment.plazi.org/id/03CC1E69A304FFE9B92AFEECB347FE05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30BFFE9B920FE06B772F836.text	03CC1E69A30BFFE9B920FE06B772F836.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax rugosus (Godwin-Austen 1914)	<div><p>Dicharax rugosus (Godwin-Austen, 1914)</p><p>Fig. 33F–N</p><p>Alycaeus rugosus Godwin-Austen, 1914: 359, pl. 141 figs 7, 7a.</p><p>Dicharax sajodentatus Gittenberger &amp; Choki Gyeltshen in Gittenberger et al., 2024: 200, figs 19, 21. Syn. nov.</p><p>Alycaeus (Dicharax) rugosus – Gude 1921: 268–269.</p><p>Chamalycaeus (Dicharax) rugosus – Ramakrishna et al. 2010: 66.</p><p>Metalycaeus rugosus – Páll-Gergely et al. 2020: 154.</p><p>Diagnosis</p><p>The combination of the strongly ribbed R1, the rather centrally located R3 swelling, the large distance between the inner and outer peristomes and the fringed aperture distinguishes this species from its congeners.</p><p>Type material examined</p><p>BHUTAN • holotype of Dicharax sajodentatus (Fig. 33K–N) (photographs examined).</p><p>INDIA • 3 syntypes of D. rugosus (figured specimen indicated by pink wool; Fig. 33F–J); Burroi Gorge, Dafla; NHMUK 1903.7.1.2641 .</p><p>Type localities</p><p>“Burroi Gorge, Dafla Hills” ( A. rugosus); “ Samdrup Jongkhar Dzongkhag: N of Bangtar, Phuntshothang, 26°53’N 91°42’E, 375 m a.s.l.” ( D. sajodentatus).</p><p>Differential diagnosis</p><p>For differences with newly described Bhutanese species ( D. zhemdentatus, D. trashidentatus, D. umbiclausus and D. aspidentatus) see Gittenberger et al. (2024) (under the name D. sajodentatus). See also under C. asaluensis, C. crispatus, C. dohertyi and D. muspratti .</p><p>Remarks</p><p>During the examination of A. rugosus, we found traces of a spiral striation on the teleoconch, based on which we classified that species in the genus Metalycaeus (see Páll-Gergely et al. 2020). However, the re-examination revealed that those spiral lines were probably part of lower shell layers of the corroded syntypes, and moreover, Gittenberger et al. (2024) described several similar species from Bhutan that can undoubtedly be assigned to Dicharax .</p><p>The comparison of syntypes of D. rugosus and D. sajodentatus revealed that their shell shape, sculpture and formation of the peristome are identical; therefore, we treat the latter as a synonym of the former.</p></div>	https://treatment.plazi.org/id/03CC1E69A30BFFE9B920FE06B772F836	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8B9BFF95CB484F85F.text	03CC1E69A30AFFE8B9BFF95CB484F85F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 2</p><p>Remarks</p><p>This species group is characterized by a strongly elevated, short R3 swelling that is situated very close to the outer peristome, and a slightly to strongly crenulated peristome.</p><p>Dicharax bicrenatus is classified in this species group due to the small distance between the R3 swelling and the outer peristome. However, it also has a large distance between the two peristomes, which is characteristic of Dicharax species group 1.</p></div>	https://treatment.plazi.org/id/03CC1E69A30AFFE8B9BFF95CB484F85F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA00FEECB79BFCBF.text	03CC1E69A30AFFE8BA00FEECB79BFCBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax trashidentatus Gittenberger, Choki Gyeltshen & Kezang Tobgay 2024	<div><p>Dicharax trashidentatus Gittenberger, Choki Gyeltshen &amp; Kezang Tobgay, 2024</p><p>Fig. 29E–H</p><p>Dicharax trashidentatus Gittenberger, Choki Gyeltshen &amp; Kezang Tobgay in Gittenberger et al., 2024: 203, figs 21–22.</p><p>Type locality</p><p>“ Trashigang Dzongkhag: 3 km E of Trashigang, 825 m a.s.l., 27°21’’N 91°35’E ” (Bhutan).</p><p>Differential diagnosis</p><p>In shell shape and sculpture this species is practically identical with D. zhemdentatus . The only difference is that the chevrons (for the terminology see Gittenberger et al. 2024) reach the peristome edge, and do not in D. zhemdentatus . Therefore, we maintain this as a valid species until intermediate forms become available. For the differences with other Bhutanese species see Gittenberger et al. (2024). See also under Dicharax muspratti .</p></div>	https://treatment.plazi.org/id/03CC1E69A30AFFE8BA00FEECB79BFCBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA7AFCC1B244FB64.text	03CC1E69A30AFFE8BA7AFCC1B244FB64.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax umbiclausus Gittenberger, Choki Gyeltshen & Kezang Tobgay 2024	<div><p>Dicharax umbiclausus Gittenberger, Choki Gyeltshen &amp; Kezang Tobgay, 2024</p><p>Fig. 29M–P</p><p>Dicharax umbiclausus Gittenberger, Choki Gyeltshen &amp; Kezang Tobgay in Gittenberger et al., 2024: 204, figs 3, 21, 23.</p><p>Type locality</p><p>“ Samdrup Jongkhar Dzongkhag: 20 km N of Samdrup Jongkhar, 1680 m a.s.l., 26°59’N 91°31’E ” (Bhutan).</p><p>Differential diagnosis</p><p>The most similar species is D. aspidentatus, which has a slightly open umbilicus, more widely-spaced ribs, and a lower R3 swelling. For other differences, see Gittenberger et al. 2024.</p></div>	https://treatment.plazi.org/id/03CC1E69A30AFFE8BA7AFCC1B244FB64	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30AFFE8BA9FFB1BB7A0F92A.text	03CC1E69A30AFFE8BA9FFB1BB7A0F92A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax zhemdentatus Gittenberger & Choki Gyeltshen 2024	<div><p>Dicharax zhemdentatus Gittenberger &amp; Choki Gyeltshen, 2024</p><p>Fig. 30I–L</p><p>Dicharax zhemdentatus Gittenberger &amp; Choki Gyeltshen in Gittenberger et al., 2024: 204, figs 20–21.</p><p>Type locality</p><p>“ Zhemgang Dzongkhag: Ngangla - Trong (= Tong), 2 km N of Ngangla, c. 17 km NE of Panbang, 1085 m a.s.l., 26°58’N 91°03’E ” (Bhutan).</p><p>Differential diagnosis</p><p>For diagnosis and differences with other Bhutanese species see Gittenberger et al. (2024). See also under D. trashidentatus .</p></div>	https://treatment.plazi.org/id/03CC1E69A30AFFE8BA9FFB1BB7A0F92A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEBB933FEECB50EFC18.text	03CC1E69A309FFEBB933FEECB50EFC18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax bicrenatus (Godwin-Austen 1874)	<div><p>Dicharax bicrenatus (Godwin-Austen, 1874)</p><p>Fig. 34A–E</p><p>Alycaeus bicrenatus Godwin-Austen, 1874: 148, pl. 3 fig. 5.</p><p>Alycaeus (Dicharax) bicrenatus – Kobelt 1902: 365. — Gude 1921: 238–239.</p><p>Alycaeus bicrenatus – Godwin-Austen 1884: pl. 51 fig. 4; 1914: 386–387, pl. 144 figs 5, 5a–b. Chamalycaeus (Dicharax) bicrenatus – Ramakrishna et al. 2010: 56.</p><p>Dicharax (?) bicrenatus – Páll-Gergely et al. 2020: 79.</p><p>Diagnosis</p><p>The combination of the small shell size, the large distance between the inner and outer peristomes, the depressed shell, the fine ribbing, and the short R3 swelling defines this species.</p><p>Type material examined</p><p>INDIA • 7 syntypes in two vials (Fig. 34A–E); Kopamedza, Naga Hills; NHMUK 1903.7.1.2490 .</p><p>Type locality</p><p>“Kopamedza Peak, Naga Hill, 8–9,000 feet, in forest”.</p></div>	https://treatment.plazi.org/id/03CC1E69A309FFEBB933FEECB50EFC18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEBBA95FC29B7E5FA41.text	03CC1E69A309FFEBBA95FC29B7E5FA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax bjiminaensis Gittenberger & Choki Gyeltshen 2024	<div><p>Dicharax bjiminaensis Gittenberger &amp; Choki Gyeltshen, 2024</p><p>Fig. 35A–D</p><p>Dicharax bjiminaensis Gittenberger &amp; Choki Gyeltshen in Gittenberger et al., 2024: 212, figs 35–36.</p><p>Type locality</p><p>“ Thimphu Dzongkhag: road W of quarry near Bjimina (13 km SW of Thimphu), 2700 m a.s.l., 27°24’N 89°34’E ” (Bhutan).</p><p>Differential diagnosis</p><p>The combination of the densely ribbed R1, the elevated, short R3 swelling situated close to the outer peristome, the large distance between the inner and outer peristomes is true for this species and for D. lunabulbus . However, the latter species is much smaller, it has a more undulated inner peristome, a larger distance between the inner and outer peristomes, and more widely-spaced R2 ribs. Dicharax vanderbijli has a smaller distance between the inner and outer peristomes, and has more lamella-like and widely-spaced ribs.</p></div>	https://treatment.plazi.org/id/03CC1E69A309FFEBBA95FC29B7E5FA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A309FFEDB924F9C2B508FC38.text	03CC1E69A309FFEDB924F9C2B508FC38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax crenatus (Godwin-Austen 1871)	<div><p>Dicharax crenatus (Godwin-Austen, 1871)</p><p>Fig. 34F–J</p><p>Alycaeus crenatus Godwin-Austen, 1871: 90–91, pl. 3 fig. 5.</p><p>Alycaeus crenatus – Godwin-Austen 1874: 150; 1914: 388–389, pl. 143 figs 8, 8a–b. — Sowerby 1877: pl. 1, species 1, figs a–b.</p><p>Alycaeus (Dicharax) crenatus – Kobelt 1902: 366. — Gude 1921: 241.</p><p>Chamalycaeus (Dicharax) crenatus – Ramakrishna et al. 2010: 57.</p><p>Dicharax crenatus – Páll-Gergely et al. 2020: 56.</p><p>Type material examined</p><p>INDIA • 2 syntypes (Fig. 34F–J); Mokarsa, Khasi Hills; Godwin-Austen coll.; NHMUK 1903.7.1.2642 .</p><p>Type locality</p><p>“On Burrail Range, N. Cachar, at ca 5000 feet ”.</p><p>Differential diagnosis</p><p>Dicharax daflaensis has a smooth R2, while D. crenatus possesses curved R2 ribs. Moreover, the R3 swelling of D. daflaensis is more conical, i.e., less abruptly elevating. Dicharax tangmaiensis has a more strongly undulated inner peristome, and a smooth R2 surface. While in D. crenatus the R2 and R3 are of comparable length, in D. davisi and D. ochraceus the R3 is much longer than the R2.</p><p>Remarks</p><p>In the original description, Godwin-Austen (1871) gave “Burrail Range, N. Cachar” as the type locality. Later (Godwin-Austen 1914: 389) explained that he found two shells in his collection (from Mokarsa in the NW Khasi Hills, no. 2642), based on which the drawings of A. crenatus were made in the original description (Godwin-Austen 1871), but their locality was not mentioned in the original description. Based on ICZN Art. 72.4.1.1, the two shells from Mokarsa are also part of the type series. Even if Godwin-Austen referred to one of the shells as “the type shell” (Godwin-Austen 1914: 389), this does not constitute a valid lectotype designation (Art 74.5), because the condition of “unambiguousity” has not been met (we do not know which of the two shells were selected to be “the type”).</p><p>Godwin-Austen (1874) noted that “ A. crenatus was found as far east as Shiroifurar, also at Kezakenomih and Yémai”. Those samples were not examined by us.</p></div>	https://treatment.plazi.org/id/03CC1E69A309FFEDB924F9C2B508FC38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFEDB908FC49B77BF8D4.text	03CC1E69A30FFFEDB908FC49B77BF8D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax cucullatus (Theobald 1870)	<div><p>Dicharax cucullatus (Theobald, 1870)</p><p>Fig. 36A–J</p><p>Alycaeus cucullatus Theobald, 1870: 396–397, pl. 18 fig. 2.</p><p>Alycaeus davisi Godwin-Austen, 1914: 408, pl. 148 figs 9, 9a.</p><p>Alycaeus cucullatus – Sowerby 1877: pl. 2, species 12. — Godwin-Austen 1914: 407, pl. 155 fig. 5.</p><p>Alycaeus (Dicharax) cucullatus – Kobelt 1902: 367–368. — Gude 1921: 244–245.</p><p>Alycaeus (Chamalycaeus) davisi – Gude 1921: 226.</p><p>Dicharax cucullatus – Páll-Gergely et al. 2020: 57. — Jirapatrasilp et al. 2021: 4, figs 2–3, 4a–b, 5a, 6h (treated D. davisi as a junior synonym).</p><p>Dicharax davisi – Páll-Gergely et al. 2020: 58.</p><p>Type material examined</p><p>MYANMAR • 2 syntypes of D. cucullatus; Shan States; NHMUK 1888.12.4.951-952 • 4 syntypes of A. davisi; Shan States; Woodthorpe leg.; NHMUK 1903.7.1.1630 .</p><p>Type localities</p><p>“Shan States” ( A. cucullatus); “ Siam and Shan boundary” ( A. davisi).</p><p>Differential diagnosis</p><p>The most similar species is D. ochraceus, which is smaller, and has more widely-spaced R2 ribs (Jirapatrasilp et al. 2021). See also under D. digitatus .</p><p>Remarks</p><p>This is a variable species regarding the aperture shape and the height of R3 swelling (see Jirapatrasilp et al. 2021).</p></div>	https://treatment.plazi.org/id/03CC1E69A30FFFEDB908FC49B77BF8D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30FFFECB91BF875B753FBDC.text	03CC1E69A30FFFECB91BF875B753FBDC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax digitatus (H. F. Blanford 1871)	<div><p>Dicharax digitatus (H.F. Blanford, 1871)</p><p>Fig. 34K–O</p><p>Alycaeus digitatus H.F. Blanford, 1871: 41–42, pl. 2 fig. 4.</p><p>Alycaeus (Dicharax) digitatus – Kobelt 1902: 369. — Gude 1921: 248.</p><p>Alycaeus digitatus – Godwin-Austen 1914: 339–340, pl. 134 figs 5, 5a.</p><p>Chamalycaeus (Dicharax) digitatus – Ramakrishna et al. 2010: 59.</p><p>Dicharax digitatus – Páll-Gergely et al. 2020: 59.</p><p>Diagnosis</p><p>The high and short R3 swelling, which is situated close to the outer peristome, the strongly fringed inner peristome, the large distance between the inner and outer peristomes, and the protruding finger-like projections on the inner peristome distinguish this species from all congeners.</p><p>Material examined</p><p>INDIA • 1 shell (probably not syntype, but figured by Godwin-Austen 1914; Fig. 34K–O); Rechila Pk, Sikkim; W. Robert leg.; NHMUK 1903.7.1.1253 .</p><p>Type locality</p><p>“apud Darjeeling in vallo Rungno fluminis Himalayae Sikkimensis”.</p><p>Differential diagnosis</p><p>Other species of this species group may also have finger-like projections of the peristome ( D. cucullatus, D. ochraceus, D. plectocheilus, D. tangmaiensis), but in those cases the outer peristome ‘follows’ the projections, while in D. digitatus the outer peristome stops at a single, straight line. See also under D. bicrenatus .</p></div>	https://treatment.plazi.org/id/03CC1E69A30FFFECB91BF875B753FBDC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30EFFECBA72FB6CB2E8F990.text	03CC1E69A30EFFECBA72FB6CB2E8F990.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax lunabulbus Gittenberger, Choki Gyeltshen & Kezang Tobgay 2024	<div><p>Dicharax lunabulbus Gittenberger, Choki Gyeltshen &amp; Kezang Tobgay, 2024</p><p>Fig. 35E–H</p><p>Dicharax lunabulbus Gittenberger, Choki Gyeltshen &amp; Kezang Tobgay in Gittenberger et al., 2024: 214, figs 38, 43.</p><p>Type locality</p><p>“ Chhukha Dzongkhag: SE of Gedu, 26°54’N 89°33’E, 1988 m a.s.l.” (Bhutan).</p><p>Differential diagnosis</p><p>Dicharax vanderbijli has more widely-spaced R1 ribs, smaller distance between the inner and outer peristomes, and a less fringed aperture. See also under D. bicrenatus and D. bjiminaensis .</p></div>	https://treatment.plazi.org/id/03CC1E69A30EFFECBA72FB6CB2E8F990	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A30EFF91B932F9B0B5E4FEEC.text	03CC1E69A30EFF91B932F9B0B5E4FEEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax ochraceus (Godwin-Austen 1893)	<div><p>Dicharax ochraceus (Godwin-Austen, 1893)</p><p>Fig. 36K–O</p><p>Alycaeus ochraceus Godwin-Austen, 1893: 594–595 .</p><p>Alycaeus ochraceus – Godwin-Austen 1897: 3, pl. 63 figs 7, 7a–b; 1914: 411.</p><p>Alycaeus (Dicharax) ochraceus – Kobelt 1902: 374. — Gude 1921: 263.</p><p>Dicharax ochraceus – Páll-Gergely et al. 2020: 69.</p><p>Type material examined</p><p>MYANMAR • 2 syntypes (Fig. 36K–O); Ruby mine Disr., Up. Burma; Doherty leg.; NHMUK 1903.7.1.2684 .</p><p>Type locality</p><p>“Ruby Mines District, Upper Burmah”.</p><p>Differential diagnosis</p><p>See under D. cucullatus (the most similar species) and D. digitatus .</p></div>	https://treatment.plazi.org/id/03CC1E69A30EFF91B932F9B0B5E4FEEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF91B90DFE9CB54CFB57.text	03CC1E69A373FF91B90DFE9CB54CFB57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax plectocheilus (Benson 1859)	<div><p>Dicharax plectocheilus (Benson, 1859)</p><p>Fig. 37A–D</p><p>Alycaeus plectocheilus Benson, 1859: 180 .</p><p>Alycaeus plectocheilus – Sowerby 1877: pl. 2, species 14. — Godwin-Austen 1914: 342–343, pl. 134</p><p>figs 4, 4a–c. Alycaeus (Dicharax) plectochilus [sic] – Kobelt 1902: 375. — Gude 1921: 264–265. Alycaeus plectocheilus, large var. – Godwin-Austen 1914: 342–343, pl. 133 figs 3, 3a–c. Chamalycaeus (Dicharax) plectochilus – Ramakrishna et al. 2010: 65. Dicharax plectocheilus – Páll-Gergely et al. 2017: 54. — Preece et al. 2022: 80, fig. 38e–f. —</p><p>Gittenberger et al. 2024: 214, figs 39–40, 43. Dicharax (?) plectocheilus – Páll-Gergely et al. 2020: 101.</p><p>Type material examined</p><p>INDIA • 2 syntypes (Fig. 37A–D); Darjiling, Rungun Valley; Blanford coll.; NHMUK 1906.4.4.184 .</p><p>Type locality</p><p>“in valle Rungun”.</p><p>Differential diagnosis</p><p>There are two other species in this species group with smooth R2 surface: D. subdigitatus and D. tangmaiensis . The former has a larger shell, has a larger distance between the R3 swelling and the peristome, and less protruding peristomal projections; the latter is much larger than D. plectocheilus and has a less elevated R3 swelling. See also under D. digitatus .</p></div>	https://treatment.plazi.org/id/03CC1E69A373FF91B90DFE9CB54CFB57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A373FF93B939FAEBB36FFE26.text	03CC1E69A373FF93B939FAEBB36FFE26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax subdigitatus (Godwin-Austen 1876)	<div><p>Dicharax subdigitatus (Godwin-Austen, 1876)</p><p>Fig. 37E–I</p><p>Alycaeus sub-digitatus Godwin-Austen, 1876: 177 .</p><p>Alycaeus (Dicharax) daflaensis var. subdigitata – Kobelt 1902: 368.</p><p>Alycaeus (Dicharax) daflaensis var. subdigitatus – Gude 1921: 246.</p><p>Dicharax (?) daflaensis subdigitatus – Páll-Gergely et al. 2020: 84.</p><p>Type material examined</p><p>INDIA • 3 syntypes; Shengorh Peak, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7.1.2498 .</p><p>Type locality</p><p>“Shengorh Peak” and “Tánir ridge at 4000 feet ”.</p><p>Differential diagnosis</p><p>Dicharax tangmaiensis is larger, and has a shorter distance between the R3 swelling and the peristome. See also under D. plectocheilus .</p><p>Remarks</p><p>The shell regions and the sculpture are similar to D. digitatus, but the undulation of the peristome started anterior to the R3 swelling. Moreover, the R3 swelling is situated at a greater distance from the peristome edge. Therefore, D. subdigitatus is a species in its own right, instead of a subspecies of D. daflaensis .</p></div>	https://treatment.plazi.org/id/03CC1E69A373FF93B939FAEBB36FFE26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93B9BFF9EDB5B5F8A5.text	03CC1E69A371FF93B9BFF9EDB5B5F8A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 3</p><p>Remarks</p><p>This species group is characterized by the relatively large (around 5 mm), usually finely ribbed shell, two R3 swellings, and a rounded (not fringed) aperture.</p><p>One species is known from the Himalayan region ( D. mutatus), while the other three are known from central Myanmar ( D. bifrons, D. spatiosus, D. woodthorpei).</p></div>	https://treatment.plazi.org/id/03CC1E69A371FF93B9BFF9EDB5B5F8A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93B939FE5AB296FB86.text	03CC1E69A371FF93B939FE5AB296FB86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax tangmaiensis (Chen & Zhang 2001)	<div><p>Dicharax tangmaiensis (Chen &amp; Zhang, 2001)</p><p>Fig. 37J–N</p><p>Chamalycaeus tangmaiensis Chen &amp; Zhang, 2001: 184–185, 188–189, figs 1–4.</p><p>Dicharax (?) tangmaiensis – Páll-Gergely et al. 2017: 107–108.</p><p>Dicharax tangmaiensis – Páll-Gergely et al. 2020: 73, fig. 17.</p><p>Diagnosis</p><p>A large species with smooth R2. See under D. digitatus and D. plectocheilus and D. subdigitatus .</p><p>Type material examined</p><p>CHINA • holotype (Fig. 37J–N); Tong-Mai Town, Bo-Mi County, Tibet Autonomous Region; 20 Jun. 1980; Chen De-Niu leg.; CASIZ TM 0010054 • 1 paratype; same data as for holotype; CASIZ TM 0010056 .</p><p>Type locality</p><p>“Tongmai Town, (30°01’N, 95°E), Bomi County, Tibet Autonomous Region, China ”.</p></div>	https://treatment.plazi.org/id/03CC1E69A371FF93B939FE5AB296FB86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF93BA89FBBBB7A8FA5C.text	03CC1E69A371FF93BA89FBBBB7A8FA5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax vanderbijli Gittenberger & Choki Gyeltshen 2024	<div><p>Dicharax vanderbijli Gittenberger &amp; Choki Gyeltshen, 2024</p><p>Fig. 35I–L</p><p>Dicharax vanderbijli Gittenberger &amp; Choki Gyeltshen in Gittenberger et al., 2024: 212, figs 35, 37.</p><p>Type locality</p><p>“ Paro Dzongkhag: 12 km SW of Paro, 3680 m a.s.l., 27°22’N 89°19’E ” (Bhutan).</p><p>Differential diagnosis</p><p>Most similar to D. lunabulbus and D. bjiminaensis . For comparisons with those species, see the respective species.</p></div>	https://treatment.plazi.org/id/03CC1E69A371FF93BA89FBBBB7A8FA5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A371FF95B975F8A6B376FE7D.text	03CC1E69A371FF95B975F8A6B376FE7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax bifrons (Theobald 1870)	<div><p>Dicharax bifrons (Theobald, 1870)</p><p>Fig. 38A–E</p><p>Alycaeus bifrons Theobald, 1870: 396, pl. 18 fig. 1. Alycaeus bifrons – Sowerby 1877: pl. 6, species 48. — Godwin-Austen 1914: 407, pl. 139 figs 3, 3a. Alycaeus (Dicharax) bifrons – Kobelt 1902: 365–366. — Gude 1921: 239.</p><p>Dicharax (?) bifrons – Páll-Gergely et al. 2020: 79.</p><p>Type material examined</p><p>MYANMAR • 3 syntypes (Fig. 38A–E); Shan States; NHMUK 1888.12.4.956-958 .</p><p>Type locality</p><p>“ Shan States”.</p><p>Differential diagnosis</p><p>The most similar species is D. mutatus, which has a longer R2 and more pronounced R3 swellings.</p></div>	https://treatment.plazi.org/id/03CC1E69A371FF95B975F8A6B376FE7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A377FF95B921FE0EB7A8FB68.text	03CC1E69A377FF95B921FE0EB7A8FB68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax mutatus (Godwin-Austen 1876)	<div><p>Dicharax mutatus (Godwin-Austen, 1876)</p><p>Fig. 38F–J</p><p>Alycaeus mutatus Godwin-Austen, 1876: 177–178, pl. 7 figs 11, 11a.</p><p>Alycaeus (Dicharax) mutatus – Kobelt 1902: 373–374. — Gude 1921: 260–261.</p><p>Alycaeus mutatus – Godwin-Austen 1914: 357, pl. 145 figs 9, 9a.</p><p>Chamalycaeus (Dicharax) mutatus – Ramakrishna et al. 2010: 63.</p><p>Type material examined</p><p>INDIA • 9 syntypes (Fig. 38F–J); Toruputu Peak, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7.1.2495 .</p><p>Type locality</p><p>“On Torúpútú, Tánir, and Shengorh Peaks, at 6–7000 feet elevation, in the dead leaves and moss about the roots of the forest”.</p><p>Differential diagnosis</p><p>Dicharax spatiosus is larger, has a longer R2, a more elevated main R3 swelling, and a clearer distinction between the inner and outer peristomes. For differences with the most similar species, D. bifrons, see under that species.</p></div>	https://treatment.plazi.org/id/03CC1E69A377FF95B921FE0EB7A8FB68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A377FF95BAD0FB19B426F83D.text	03CC1E69A377FF95BAD0FB19B426F83D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax spatiosus Pall-Gergely & Hunyadi 2022	<div><p>Dicharax spatiosus Páll-Gergely &amp; Hunyadi, 2022</p><p>Fig. 39A–E</p><p>Dicharax spatiosus Páll-Gergely &amp; Hunyadi, 2022: 97, fig. 5.</p><p>Diagnosis</p><p>The large shell, long R2, two elevated swellings on R3, and the circular (not fringed) aperture distinguishes this species from all congeners. See also under D. mutatus and also the original description of D. spatiosus for further comparisons.</p><p>Type material examined</p><p>MYANMAR • holotype (Fig. 39A–E); Shan State, 7.5 km N of Pinlaung, Tar Kge, near “Big Bang Cave”; 20°810.2730′ N, 96°847.4420′ E; 1540 m a.s.l.; A. Hunyadi, K. Okubo and J.U. Otani leg.; coll. HA (to be deposited in the HNHM) .</p><p>Type locality</p><p>“ Myanmar, Shan State, Pinlaung centre N 7.5 km, Tar Kge, near „Big Bang Cave”, 20810.2730N, 96847.4420E, 1540 m a.s.l.”.</p></div>	https://treatment.plazi.org/id/03CC1E69A377FF95BAD0FB19B426F83D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A376FF97B934FEECB787FC5E.text	03CC1E69A376FF97B934FEECB787FC5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax woodthorpi (Godwin-Austen 1914)	<div><p>Dicharax woodthorpi (Godwin-Austen, 1914)</p><p>Fig. 39F–J</p><p>Alycaeus woodthorpi Godwin-Austen, 1914: 414, pl. 155 fig. 14.</p><p>Alycaeus (Dicharax) woodthorpei [sic] – Gude 1921: 275.</p><p>Dicharax (?) woodthorpei – Páll-Gergely et al. 2020: 107; 2021: 37, fig. 20b.</p><p>Diagnosis</p><p>This species differs from the other three species of the same species group by the smooth R2.</p><p>Type material examined</p><p>MYANMAR • 22 syntypes in two vials (Fig. 39F–J); Fort Stedman, Burma; Woodthorpe leg.; NHMUK 1903.7.1.3064 .</p><p>Additional material examined</p><p>MYANMAR • 2 shells; Burma; E.R. Sykes coll.; acc. no. 1825; NHMUK 20150126.</p><p>Type locality</p><p>“Fort Stedman, Burma ”.</p><p>Differential diagnosis</p><p>There are several species with smooth R2 surface (mostly in Dicharax species group 8), but D. woodthorpei can be distinguished from them by the relatively large shell, the double R3 swelling and the rounded aperture.</p></div>	https://treatment.plazi.org/id/03CC1E69A376FF97B934FEECB787FC5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A375FF97B937FAA4B578F863.text	03CC1E69A375FF97B937FAA4B578F863.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax duoculmen (Godwin-Austen 1914)	<div><p>Dicharax duoculmen (Godwin-Austen, 1914)</p><p>Fig. 40A–E</p><p>Alycaeus duoculmen Godwin-Austen, 1914: 365, pl. 157 figs 2, 2a.</p><p>Alycaeus (Raptomphalus) duoculmen – Gude 1921: 286–287.</p><p>Chamalycaeus (Raptomphalus) duoculmen – Ramakrishna et al. 2010: 68.</p><p>Dicharax (?) duoculmen – Páll-Gergely et al. 2020: 86.</p><p>Type material examined</p><p>INDIA • holotype (single shell mentioned in the original description: Fig. 40A–E); Tsanspu Valley; Oakes leg.; NHMUK 1903.7.1.3582 .</p><p>Type locality</p><p>“Tsanspu Valley”.</p><p>Differential diagnosis</p><p>The most similar species is D. magnificus, which has a stronger R1 sculpture, a more strongly fringed aperture and a sharper keel on the umbilical side of R2.</p></div>	https://treatment.plazi.org/id/03CC1E69A375FF97B937FAA4B578F863	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A375FF97B9BFFBE3B523FAA7.text	03CC1E69A375FF97B9BFFBE3B523FAA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 4</p><p>Remarks</p><p>This species group is characterized by the relatively large (around 5 mm), finely ribbed R1, smooth R2, two R3 swellings, and the fringed aperture. Species of the Dicharax species group 3 also have double R3 swellings, but have rounded apertures.</p><p>Both species are known from the Indian Himalaya.</p></div>	https://treatment.plazi.org/id/03CC1E69A375FF97B9BFFBE3B523FAA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF99B9BFFA97B283F939.text	03CC1E69A37BFF99B9BFFA97B283F939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 5</p><p>Remarks</p><p>This species group is characterized by shells with an elevated spire and relatively low R3 swelling. Other traits are variable. It is likely that D. globosus and D. vestitus are relatives, because other than the shell height they share the strong ribs, which are peculiarly oblique. Dicharax burchi, and D. verrucosus are also similar in the R3 swelling that is situated close to the aperture. Dicharax conicus may not be related to the other high-spired species, and can be recognized by its peculiar outer peristome.</p></div>	https://treatment.plazi.org/id/03CC1E69A37BFF99B9BFFA97B283F939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF99B937FEECB46CFAF3.text	03CC1E69A37BFF99B937FEECB46CFAF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax magnificus (Godwin-Austen 1914)	<div><p>Dicharax magnificus (Godwin-Austen, 1914)</p><p>Fig. 40F–P</p><p>Alycaeus (Raptomphalus) magnificus Godwin-Austen, 1914: 366, pl. 156 figs 1, 1a–b.</p><p>Alycaeus (Raptomphalus) magnificus – Gude 1921: 288.</p><p>Chamalycaeus (Raptomphalus) magnificus – Ramakrishna et al. 2010: 69.</p><p>Metalycaeus (?) magnificus – Páll-Gergely et al. 2020: 146, fig. 31b.</p><p>Type material examined</p><p>INDIA • 1 syntype (Fig. 40F–J); Yamney Valley, Abor Hills; NHMUK 1903.7.1.3115 .</p><p>Additional material examined</p><p>INDIA • 1 shell (Fig. 40K–P); Arunachal Pradesh, 2 km from Kalek towards <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=95.172966&amp;materialsCitation.latitude=28.101116" title="Search Plazi for locations around (long 95.172966/lat 28.101116)">Rottung</a>, base of the dry bamboo clumps, in semi-evergreen forest; 28°6.067′ N, 95°10.378′ E; 784 m a.s.l.; 24 April 2022; Aravind N.A. and Surya Narayanan leg.; NZSI LM1914 • 7 shells; same data as for preceding; ATREE .</p><p>Type locality</p><p>“Yamne valley, Abor Hills”.</p><p>Differential diagnosis</p><p>See under Dicharax duoculmen .</p><p>Remarks</p><p>Based on the spiral striae visible on the protoconch of the syntype, this species has been classified in Metalycaeus (see Páll-Gergely et al. 2020). Re-examination of that syntype revealed that those spiral striae are parts of lower shell layers; therefore, those are not homologous with those of species of Metalycaeus, which are raised striae. Furthermore, additional, fresh shells collected at Kalek-Rottung Road clearly lack spiral striation, showing that this is a species of Dicharax . Dicharax magnificus is the type species of Raptomphalus Godwin-Austen, 1914, and as a consequence, that taxon is a synonym of Dicharax (instead of Metalycaeus).</p></div>	https://treatment.plazi.org/id/03CC1E69A37BFF99B937FEECB46CFAF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37BFF9BBAD4F949B2EDFDD2.text	03CC1E69A37BFF9BBAD4F949B2EDFDD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax burchi Jirapatrasilp & Pall-Gergely 2021	<div><p>Dicharax burchi Jirapatrasilp &amp; Páll-Gergely, 2021</p><p>Fig. 41A–E</p><p>Dicharax burchi Jirapatrasilp &amp; Páll-Gergely in Jirapatrasilp et al., 2021: 9, figs 6f, 10, 11a.</p><p>Type material examined</p><p>THAILAND • holotype; Khon Kaen Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.956635&amp;materialsCitation.latitude=16.80911" title="Search Plazi for locations around (long 101.956635/lat 16.80911)">Tham Phaya Nakharat</a>; 16°48′32.8″ N, 101°57′23.9″ E; 21 Jul. 2020; P. Tongkerd and A. Pholyotha leg.; CUMZ 7428. For other specimens, see Jirapatrasilp et al. (2021) .</p><p>Additional material examined</p><p>MYANMAR • 1 shell (Fig. 41A–E); Shan State, ca 6 km E of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.3004&amp;materialsCitation.latitude=20.133368" title="Search Plazi for locations around (long 97.3004/lat 20.133368)">Hsihseng</a> centre, right side of rd + 500 m on unpaved rd, limestone hill; 20°8.002′ N, 97°18.024′ E; 1000 m a.s.l.; 7 Oct. 2018; A. Hunyadi, K. Okubo and J.U. Otani leg.; coll. HA (locality code: 2018/41) .</p><p>Type locality</p><p>“Tham Phaya Nakharat, Khon Kaen Province, Thailand, 16°48’32.8”N, 101°57’23.9”E ”.</p><p>Differential diagnosis</p><p>Differs from the most similar D. verrucosus by the more elevated spire and the longer R3.</p></div>	https://treatment.plazi.org/id/03CC1E69A37BFF9BBAD4F949B2EDFDD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9BB922FD75B346F8FB.text	03CC1E69A379FF9BB922FD75B346F8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax conicus (Godwin-Austen 1871)	<div><p>Dicharax conicus (Godwin-Austen, 1871)</p><p>Fig. 42</p><p>Alycaeus conicus Godwin-Austen, 1871: 87–88, pl. 3 fig. 1.</p><p>Alycaeus conicus var. nanus Godwin-Austen, 1914: 388, pl. 138 figs 6, 6a–b. Syn. nov.</p><p>Dicharax conicus jatingaensis Páll-Gergely in Páll-Gergely et al., 2020: 56 (replacement name for Alycaeus conicus var. nanus Godwin-Austen, 1914, non Alycaeus nanus Möllendorff, 1886)</p><p>Alycaeus conicus – Sowerby 1877: pl. 1, species 9. — Godwin-Austen 1914: 387–388, pl. 143 figs 4, 4a–b. — Gude 1921: 208.</p><p>Alycaeus (Alycaeus) conicus – Kobelt 1902: 342. — Ramakrishna et al. 2010: 47.</p><p>Alycaeus conicus var. nana – Gude 1921: 208.</p><p>Dicharax conicus – Páll-Gergely et al. 2020: 45.</p><p>Diagnosis</p><p>Besides the elevated spire, this species can be distinguished from its congeners based on the nearly smooth R1, and the peculiar outer peristome, which is strongly expanded and turns anteriorly.</p><p>Type material examined</p><p>INDIA • 12 syntypes of A. conicus (Fig. 42A–E); Samiamri, E of the Kopili R.; Godwin-Austen leg.; NHMUK 1903.7.1.2674 • 12 syntypes of A. conicus var. nanus (Fig. 42F–J); Jatinga Valley, N. Cachar; NHMUK 1903.7.1.2675 .</p><p>Additional material examined</p><p>INDIA • 1 shell (included with type lot, but not mentioned in the original description and not considered as part of the type series); Khasi Hills; NHMUK 1903.7.1.2565 .</p><p>Type localities</p><p>“Was abundant on the Limestone Hill east of Kopili river, North Cachar District, and was occasionally also found in other places, but rare” ( A. conicus); “Jatinga Valley, North Cachar Hills” ( A. conicus var. nanus).</p><p>Remarks</p><p>Besides a slight difference in shell size, we found no important conchological traits that differ between A. conicus and A. conicus var. nanus; therefore, we see no reason to maintain the latter as a subspecies.</p></div>	https://treatment.plazi.org/id/03CC1E69A379FF9BB922FD75B346F8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A379FF9DB924F88EB79FFC90.text	03CC1E69A379FF9DB924F88EB79FFC90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax globulus (Godwin-Austen 1874)	<div><p>Dicharax globulus (Godwin-Austen, 1874)</p><p>Fig. 43A–E</p><p>Alycaeus globulus Godwin-Austen, 1874: 147–148, pl. 3 fig. 4.</p><p>Alycaeus (Dicharax) globulus – Kobelt 1902: 371. — Gude 1921: 254.</p><p>Alycaeus globulus – Godwin-Austen 1914: 392, pl. 144 figs 4, 4a–b.</p><p>Chamalycaeus (Dicharax) globulus – Ramakrishna et al. 2010: 60.</p><p>Dicharax (?) globulus – Páll-Gergely et al. 2020: 63.</p><p>Diagnosis</p><p>Differs from all congeners by the deep longitudinal grooves of the R3.</p><p>Type material examined</p><p>INDIA • 13 syntypes (Fig. 43A–E); Phunggum, Lahupa Naga Hills, Munipur; NHMUK 1903.7.1.2486 .</p><p>Type locality</p><p>“Phunggum, a Naga village at head of the Lanier valley, at 5,000 feet ”.</p><p>Remarks</p><p>It appears that the outer peristome of this species is situated close to the R3 swelling. Such extended space between the inner and outer peristomes is visible in other species as well, and most extreme in the cases of D. bicrenatus, D. digitatus, D. rugosus and D. umbiclausus . However, in D. globulus the relatively large area between the two peristomes bears deep longitudinal grooves between the aperture and the R3 swelling.</p></div>	https://treatment.plazi.org/id/03CC1E69A379FF9DB924F88EB79FFC90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37FFF9DBADEFCB1B527FA33.text	03CC1E69A37FFF9DBADEFCB1B527FA33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax verrucosus Pall-Gergely & Hunyadi 2021	<div><p>Dicharax verrucosus Páll-Gergely &amp; Hunyadi, 2021</p><p>Fig. 41F–J</p><p>Dicharax verrucosus Páll-Gergely &amp; Hunyadi in Páll-Gergely et al., 2021: 44, fig. 34.</p><p>Type material examined</p><p>MYANMAR • holotype (Fig. 41F–J); Shan State, Taunggyi, hill above Aye Say Tee Pagoda, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.0506&amp;materialsCitation.latitude=20.791483" title="Search Plazi for locations around (long 97.0506/lat 20.791483)">Dragon Cave</a>; 20°47.489′ N, 97°03.036′ E; 1380 m a.s.l.; A. Hunyadi, K. Okubo and J.U. Otani leg.; HNHM 104862. See other specimens in Páll-Gergely et al. (2021) .</p><p>Type locality</p><p>“ Myanmar, Shan State, Taunggyi, hill above Aye Say Tee Pagoda, Dragon Cave, 20°47.489’N, 97°03.036’E, 1380 m a.s.l.”</p><p>Differential diagnosis</p><p>See under D. burchi and in the original description.</p></div>	https://treatment.plazi.org/id/03CC1E69A37FFF9DBADEFCB1B527FA33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37FFF9FBAC7FA50B586FDC6.text	03CC1E69A37FFF9FBAC7FA50B586FDC6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax vestitus subsp. vestitus (W. T. Blanford 1862)	<div><p>Dicharax vestitus vestitus (W.T. Blanford, 1862)</p><p>Fig. 43F–J</p><p>Alycaeus vestitus W.T. Blanford, 1862: 138–139 .</p><p>Alycaeus vestitus var. minor W.T. Blanford, 1862: 138 .</p><p>Alycaeus vestitus – Sowerby 1877: pl. 1, species 3. — Godwin-Austen 1914: 424–425, pl. 139 figs 2, 2a. — Gude 1921: 220–221.</p><p>Alycaeus (Alycaeus) vestitus – Kobelt 1902: 352.</p><p>Dicharax vestitus – Páll-Gergely et al. 2017: 71; 2020: 75.</p><p>Type material examined</p><p>MYANMAR • holotype (single shell mentioned in the original description: Fig. 43F–J); Moditoung; NHMUK 1906.4.4.53 .</p><p>Additional material examined</p><p>MYANMAR • 2 additional non-type specimens in the same lot from Alori Khyoung and Mamya Khyoung; NHMUK 1906.4.4.53.</p><p>Type locality</p><p>“in montibus Arakanensibus”.</p><p>Differential diagnosis</p><p>Probably the most closely related species is D. globulus due to the peculiar oblique R1 ribs. That species, however, has longitudinal grooves on R3, and a fringed aperture.</p></div>	https://treatment.plazi.org/id/03CC1E69A37FFF9FBAC7FA50B586FDC6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF9FB9BFFADBB774F9AE.text	03CC1E69A37DFF9FB9BFFADBB774F9AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 6</p><p>Remarks</p><p>This species group is characterized by large, depressed shells and rounded (not fringed) apertures. Species of this species group are distributed from Myanmar (Ayeyarwady and Yangon Regions) to the Naga Hills.</p></div>	https://treatment.plazi.org/id/03CC1E69A37DFF9FB9BFFADBB774F9AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF9FBAE6FD7AB478FAA6.text	03CC1E69A37DFF9FBAE6FD7AB478FAA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax vestitus subsp. akyabensis (Godwin-Austen 1914)	<div><p>Dicharax vestitus akyabensis (Godwin-Austen, 1914)</p><p>Fig. 43K–O</p><p>Alycaeus vestitus var. akyabensis Godwin-Austen, 1914: 425–426, pl. 155 fig. 7.</p><p>Alycaeus vestitus var. akyabensis – Gude 1921: 221.</p><p>Dicharax vestitus akyabensis – Páll-Gergely et al. 2020: 75.</p><p>Type material examined</p><p>MYANMAR • 2 syntypes (Fig. 43K–O); Baumi, Akyab; NHMUK 1888.12.4.251-252 .</p><p>Type locality</p><p>“Baumi, Akyab”.</p><p>Remarks</p><p>Dicharax vestitus akyabensis is smaller and has a thinner peristome than the nominotypical subspecies, which probably indicates intraspecific variability. Nevertheless, we maintain the two subspecies until more material becomes available.</p></div>	https://treatment.plazi.org/id/03CC1E69A37DFF9FBAE6FD7AB478FAA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A37DFF85B90CF9D3B230FD0D.text	03CC1E69A37DFF85B90CF9D3B230FD0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax glaber (W. T. Blanford 1865)	<div><p>Dicharax glaber (W.T. Blanford, 1865)</p><p>Figs 44–49</p><p>Alycaeus glaber W.T. Blanford, 1865: 84 .</p><p>Alycaeus edei Godwin-Austen, 1914: 391–392, pl. 149 figs 2, 2a. Syn. nov.</p><p>Alycaeus levis Godwin-Austen, 1914: 394, pl. 138 figs 3, 3a. Syn. nov.</p><p>Alycaeus glaber – Sowerby 1877: pl. 4, species 31. — Godwin-Austen 1914: 418–419, pl. 151 fig. 1. Alycaeus (Dicharax) glaber – Kobelt 1902: 371. — Gude 1921: 253.</p><p>Alycaeus levis – Gude 1921: 209–210.</p><p>Alycaeus (Chamalycaeus) edei – Gude 1921: 227.</p><p>Alycaeus (Alycaeus) levis – Ramakrishna et al. 2010: 48.</p><p>Chamalycaeus (Chamalycaeus) edei – Ramakrishna et al. 2010: 53.</p><p>Dicharax (?) edei – Páll-Gergely et al. 2020: 86.</p><p>Dicharax (?) glaber – Páll-Gergely et al. 2020: 89.</p><p>Dicharax (?) levis – Páll-Gergely et al. 2020: 92.</p><p>Type material examined</p><p>MYANMAR • possible syntypes of A. glaber (Fig. 44A–E); Akyab; W.T. Blanford coll.; NHMUK 1906.4.4.181 • 2 shells of ( A. glaber); Akyab, Arakan; NHMUK 20191066 • 8 syntypes of A. edei in 2 vials (Fig. 44F–J); Naraindhur, Cachar; F. Ede leg.; NHMUK 1903.7.1.1665 • holotype of A. levis (single shell mentioned in the original description: Fig. 44K–O); Munipur; Godwin-Austen coll.; NHMUK 1903.7.1.2631 .</p><p>Additional material examined</p><p>INDIA – Mizoram • 1 empty shell; Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 29; NZSI LM1730 • 1 empty shell; same data as for preceding; Ar 39; NZSI LM1731 • 1 empty shell; same data as for preceding; Ar 52; NZSI LM1732 • 1 empty shell; same data as for preceding; Ar 30; NZSI LM1733 • 1 empty shell; same data as for preceding; Ar64 (Fig. 48); NZSI LM1734 • 2 empty shells; same data as for preceding; Ar54 (Fig. 46); NZSI LM1735 • 2 empty shells; same data as for preceding; Ar 20 (Fig. 47B–D [operculum]); NZSI LM1736 • 1 empty shell (Fig. 45); same data as for preceding; Ar 19; NZSI LM1737 • 1 empty shell; same data as for preceding; Ar 65; NZSI LM1738 • 1 empty shell; same data as for preceding; Ar41 (Fig. 47A); ZSI LM1739 • 1 empty shell; same data as for preceding; Ar 28; ZSI LM1905 .</p><p>Type localities</p><p>“Akyab, Arakan; the hills south of the harbour” ( A. glaber); “Naraindhur, Cachar, No. 1665 B.M.” [probably Narainpur Village, Rajabazar Tehsil, Cachar District: 24°30.4′ N, 92°52.2′ E] ( A. edei); “Gaziphimi, Lahupa Naga Hills, Munipur” [probably Gaziphema, Naga Hills, Manipur: 25°30′ N, 94° 23.3′ E] ( A. levis).</p><p>Differential diagnosis</p><p>There are few species of Dicharax of this size. For comparisons with other species, see under D. magnus and D. ingrami .</p><p>Additional information on the morphology</p><p>A cross sectional view of R2 was examined in a specimen (Fig. 48): anterior crust forms a significantly elevated rib, which has an anterior projection (i.e., the R2 ribs are curved towards the aperture); the anterior crust folds over the much weaker posterior crust; microtunnel narrow compared to the ribs, its cross sectional view subcircular.</p><p>A D. glaber shell from the Blue Mountain apparently has short chevrons on the inner surface of its aperture (Fig. 47A). The tendency towards a fringed aperture appears in several species (see other examples in Páll-Gergely et al. 2017, 2021).</p><p>Operculum (Fig. 47B–D): colour chocolate brown, outer surface multispiral, without elevated lamina, inner side with a strongly elevated, knob-like nipple.</p><p>Distribution</p><p>This species is known from northeastern Myanmar and the neighbouring Indian areas (Fig. 49).</p><p>Remarks</p><p>The type specimens of A. glaber, A. edei and A. levis are very similar, and the notable differences (i.e., the more rounded shape of A. glaber and the strongly reflected outer peristome of A. edei) are considered to be intraspecific variability. The shells from the Blue Mountain are identical to the type specimens of A. edei with the exception of the strongly reflected outer peristome (similar cases were reported for D. notus (Godwin-Austen, 1914), see Páll-Gergely et al. 2021: fig. 26).</p></div>	https://treatment.plazi.org/id/03CC1E69A37DFF85B90CF9D3B230FD0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF87B91AFEECB321F977.text	03CC1E69A365FF87B91AFEECB321F977.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax ingrami (W. T. Blanford 1862)	<div><p>Dicharax ingrami (W.T. Blanford, 1862)</p><p>Fig. 50</p><p>Alycaeus ingrami W.T. Blanford, 1862: 135–136 .</p><p>Alycaeus ingrami var. nagaensis Godwin-Austen, 1871: 92, pl. 5 fig. 2. Syn. nov.</p><p>Alycaeus anonymus Godwin-Austen, 1914: 405–406, pl. 139 figs 1, 1a. Syn. nov.</p><p>Alycaeus ingrami – Sowerby 1877: pl. 6, species 54. — Godwin-Austen 1886: 193–194, pl. 44 figs 1,</p><p>1a–c; 1914: 421. Alycaeus nagaensis – Godwin-Austen 1884: pl. 51 figs 3, 7; 1886: 195, pl. 44 figs 3, 3a–c; 1914: 396–</p><p>397, pl. 143 figs 2, 2a– b. Alycaeus (Chamalycaeus) ingrami – Kobelt 1902: 357. — Gude 1921: 228–229. Alycaeus (Chamalycaeus) nagaensis – Kobelt 1902: 359. — Gude 1921: 230. Alycaeus anonymus – Gude 1921: 205. Chamalycaeus (Chamalycaeus) nagaensis – Ramakrishna et al. 2010: 54. Dicharax anonymus – Páll-Gergely et al. 2020: 53. Dicharax (?) ingrami – Páll-Gergely et al. 2020: 90. Dicharax (?) nagaensis – Páll-Gergely et al. 2020: 98.</p><p>Type material examined</p><p>MYANMAR • 7 shells, probable syntypes of A. ingrami (figured by Godwin-Austen 1886 and Fig. 50A– E); Tongoop, Arakan; Blanford coll.; NHMUK 1906.4.4.68 • 1 shell of A. ingrami; Arakan, “authentic”, “type var.”, 1944, No. 450; NHMUK • 2 syntypes of A. anonymus (Fig. 50K–O); Akouktoung, Pegu; NHMUK 1906.4.4.67a • 7 syntypes of A. nagaensis in 2 vials (Fig. 50F–J); Asalu, N. Cachar; NHMUK 1903.7.1.2615 .</p><p>Type localities</p><p>“prope Tongoop in Arakan ” ( A. ingrami); “Akouk-toung, Pegu: Type; also Thoudaung and Yenandoung, Pegu ” ( A. anonymus); “Neighbourhood of Asálú, rather local in its distribution, but abundant” ( A. nagaensis).</p><p>Differential diagnosis</p><p>Dicharax glaber has glossy initial whorls and R3, while those are regularly ribbed in D. ingrami (although the R3 is glossy in the type specimens of A. nagaensis). No other differences have been found between D. glaber and D. ingrami, and the difference regarding the R3 sculpture is rather minor, therefore, they may be considered synonyms. However, since we know of no specimens showing intermediate character states, we maintain both species as valid. See also under D. magnus .</p><p>Remarks</p><p>The type specimens of A. anonymus and A. ingrami are very similar, the only differences are that the former has a slightly keeled body whorl and a stronger sculpture. Nevertheless, these differences are considered intraspecific variability. Godwin-Austen (1914), when describing A. anonymus, did not compare it with any other species. The type specimens of A. nagaensis also do not differ from those of A. anonymus and A. ingrami except for their larger size, which is not sufficient for species level distinction. Therefore, both A. nagaensis and A. anonymus are junior synonyms of D. ingrami .</p></div>	https://treatment.plazi.org/id/03CC1E69A365FF87B91AFEECB321F977	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A365FF89B920F908B7C3FDC9.text	03CC1E69A365FF89B920F908B7C3FDC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax magnus (Godwin-Austen 1893)	<div><p>Dicharax magnus (Godwin-Austen, 1893)</p><p>Fig. 51A–E</p><p>Alycaeus magnus Godwin-Austen, 1893: 594 .</p><p>Alycaeus (Alycaeus) magnus – Kobelt 1902: 346–347. — Ramakrishna et al. 2010: 49.</p><p>Alycaeus magnus – Godwin-Austen 1914: 395, pl. 138 figs 1, 1a. — Gude 1921: 210–211, fig. 33. Dicharax (?) magnus – Páll-Gergely et al. 2020: 95.</p><p>Type material examined</p><p>INDIA • 1 syntype (Fig. 51A–E); Naga Hills; Beddome fr.; NHMUK 1903.7.1.1480 .</p><p>Type locality</p><p>“ Naga Hills, 150 miles eastward of Kohima”.</p><p>Differential diagnosis</p><p>This species is much larger than D. glaber, has ribbed initial whorls, and the outer peristome partly covers the umbilicus. The same is true when comparing A. magnus with D. ingrami, but both species have ribbed initial whorls.</p></div>	https://treatment.plazi.org/id/03CC1E69A365FF89B920F908B7C3FDC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36BFF89B9BFF8DDB52DF859.text	03CC1E69A36BFF89B9BFF8DDB52DF859.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 7</p><p>Remarks</p><p>This species group is characterized by a strongly swollen R3, which has its more elevated posterior end (“tip”) close to the constriction between R2 and R3.</p></div>	https://treatment.plazi.org/id/03CC1E69A36BFF89B9BFF8DDB52DF859	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36BFF89B902FD7AB2AFF8AC.text	03CC1E69A36BFF89B902FD7AB2AFF8AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax umbonalis (Benson 1856)	<div><p>Dicharax umbonalis (Benson, 1856)</p><p>Fig. 51F–N</p><p>Alycaeus umbonalis Benson, 1856: 225–226 .</p><p>Alycaeus sandowayensis Godwin-Austen, 1914: 423–424, pl. 139 figs 4, 4a. Syn. nov.</p><p>Alycaeus umbonalis – Sowerby 1877: pl. 4, species 36. — Godwin-Austen 1886: 194–195, pl. 44 figs 2, 2a–c; 1914: 413, 424.</p><p>Alycaeus (Chamalycaeus) umbonalis – Kobelt 1902: 364. — Gude 1921: 235–236.</p><p>Alycaeus (Chamalycaeus) sandowayensis – Gude 1921: 232.</p><p>Dicharax (?) sandowayensis – Páll-Gergely et al. 2020: 103.</p><p>Dicharax (?) umbonalis – Páll-Gergely et al. 2020: 106.</p><p>Diagnosis</p><p>This species differs from all other congeners of this species group by the wide umbilicus and less inflated R2.</p><p>Type material examined</p><p>MYANMAR • holotype of A. sandowayensis (single shell mentioned in the original description: Fig. 51F–J); Mai-i, Sandoway Dist., Arakan; Stoliczka leg.; NHMUK 1903.7.1.2558 • 3 syntypes of A. umbonalis (Fig. 51K–N); Burmah; UMZC I.103005 .</p><p>Additional material examined</p><p>MYANMAR • 5 shells of A. umbonalis; Akoutong, Burma; Blanford coll.; NHMUK.</p><p>Type localities</p><p>“ad Akaouktong, prope ripas fluvii Irawadi ” ( A. umbonalis); “Mai-i, Sandoway District, Arakan ” ( A. sandowayensis).</p><p>Remarks</p><p>The holotype of A. sandowayensis is a smaller shell than typical A. umbonalis, but since all shell characters are identical, we treat it as a synonym of the latter species. In the original description of A. sandowayensis, Godwin-Austen (1914) did not compare it with any other species.</p></div>	https://treatment.plazi.org/id/03CC1E69A36BFF89B902FD7AB2AFF8AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88B928FEECB5A7FC62.text	03CC1E69A36AFF88B928FEECB5A7FC62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax chennelli (Godwin-Austen 1886)	<div><p>Dicharax chennelli (Godwin-Austen, 1886)</p><p>Fig. 52A–E</p><p>Alycaeus chennelli Godwin-Austen, 1886: 192–193, pl. 48 fig. 2.</p><p>Alycaeus chennelli – Godwin-Austen 1914: 387.</p><p>Alycaeus chennelli var. – Godwin-Austen 1914: 387.</p><p>Alycaeus (Dicharax) chennelli – Kobelt 1902: 366. — Gude 1921: 240–241.</p><p>Chamalycaeus (Dicharax) chennelli – Ramakrishna et al. 2010: 57.</p><p>Dicharax chennelli – Páll-Gergely et al. 2020: 55.</p><p>Type material examined</p><p>INDIA • 8 syntypes (Fig. 52A–E); Piknui, Naga Hills; A. Chennell leg.; NHMUK 1903.7.1.2612 • 10 shells of “ chennelli var.”; Lhota Naga Hills; A. Chennell leg.; NHMUK 1903.7.1.2613 .</p><p>Type localities</p><p>“Piknúi, Naga Hills”; “Lhota Naga Hills” ( chennelli var.).</p><p>Differential diagnosis</p><p>The only stable character which distinguishes D. chennelli from D. diagonius is the presence of a lower apertural incision in the former, whilst it is absent in the latter.</p></div>	https://treatment.plazi.org/id/03CC1E69A36AFF88B928FEECB5A7FC62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF88BAC5FBE6B7E3F8FD.text	03CC1E69A36AFF88BAC5FBE6B7E3F8FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax damsangensis (Godwin-Austen 1886)	<div><p>Dicharax damsangensis (Godwin-Austen, 1886)</p><p>Fig. 53A–E</p><p>Alycaeus damsangensis Godwin-Austen, 1886: 192, pl. 43 figs 3, 3a–c.</p><p>Alycaeus (Dicharax) damsangensis – Kobelt 1902: 368. — Gude 1921: 246–247.</p><p>Alycaeus (Charax) damsangensis – Godwin-Austen 1914: 339.</p><p>Dicharax damsangensis – Páll-Gergely et al. 2020: 55. — Gittenberger et al. 2024: 207, figs 27–29. (probably not this species).</p><p>Type material examined</p><p>BHUTAN • 12 syntypes (Fig. 53A–E); Damsang, W. Bhutan; Robert leg.; NHMUK 1903.7.1.2677 .</p><p>Type locality</p><p>“Damsang Peak, Western Bhutan Hills”.</p><p>Differential diagnosis</p><p>The most similar species is D. notatus, which has a more inflated R3 and denser R1 ribs. Dicharax nowgongensis (Godwin-Austen, 1914) is also similar, but its basal lip is rounded.</p><p>Remarks</p><p>The specimens figured by Gittenberger et al. (2024) from Bhutan differ from the type specimens in the denser R1 ribbing and the ribbed R3 (vs smooth in D. damsangensis). Therefore, they likely represent an undescribed species.</p></div>	https://treatment.plazi.org/id/03CC1E69A36AFF88BAC5FBE6B7E3F8FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36AFF8DB92FF88DB739FCE2.text	03CC1E69A36AFF8DB92FF88DB739FCE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax diagonius (Godwin-Austen 1871)	<div><p>Dicharax diagonius (Godwin-Austen, 1871)</p><p>Fig. 52F–J</p><p>Alycaeus diagonius Godwin-Austen, 1871: 88–89, pl. 3 fig. 2.</p><p>Alycaeus diagonus [sic] – Sowerby 1877: pl. 1, species 2.</p><p>Alycaeus (Dicharax) diagonius – Kobelt 1902: 368–369. — Gude 1921: 247–248.</p><p>Alycaeus diagonius – Godwin-Austen 1914: 389–390, pl. 143 figs 5, 5a–b.</p><p>Chamalycaeus (Dicharax) diagonius – Ramakrishna et al. 2010: 58.</p><p>Dicharax damsangensis – Páll-Gergely et al. 2020: 59.</p><p>Type material examined</p><p>INDIA • 10 syntypes (Fig. 52F–J); Diyung Valley, N of Asalu, N. Cachar; Godwin-Austen coll.; NHMUK 1903.7.1.2678 .</p><p>Type locality</p><p>“The Diyung valley, north of Asálú, in Cachar District”.</p><p>Differential diagnosis</p><p>This species has a longer and more pointed R3 swelling and more widely-spaced R1 than Dicharax subculmen (Godwin-Austen, 1893) . See also under D. chennelli and D. habiangensis (Godwin-Austen, 1914) .</p></div>	https://treatment.plazi.org/id/03CC1E69A36AFF8DB92FF88DB739FCE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36FFF8DBAC0FC65B2B9F86A.text	03CC1E69A36FFF8DBAC0FC65B2B9F86A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax habiangensis (Godwin-Austen 1914)	<div><p>Dicharax habiangensis (Godwin-Austen, 1914)</p><p>Fig. 54</p><p>Alycaeus habiangensis Godwin-Austen, 1914: 374, pl. 138 figs 2, 2a–b.</p><p>Alycaeus generosus Godwin-Austen, 1914: 374, pl. 138 figs 8, 8a–b. Syn. nov.</p><p>Alycaeus (Dicharax) habiangensis – Gude 1921: 254.</p><p>Alycaeus (Cycloryx) generosus – Gude 1921: 279.</p><p>Chamalycaeus (Dicharax) habiangensis – Ramakrishna et al. 2010: 61.</p><p>Cycloryx generosus – Ramakrishna et al. 2010: 71.</p><p>Dicharax generosus – Páll-Gergely et al. 2020: 63.</p><p>Dicharax habiangensis – Páll-Gergely et al. 2020: 63.</p><p>Type material examined</p><p>INDIA • holotype (single shell mentioned in the original description: Fig. 54A–E) of A. habiangensis; Habiang Garo, W. Khasi; Godwin-Austen leg.; NHMUK 1903.7.1.2649 • 2 syntypes of A. generosus (Fig. 54F–J); Khasi Hills, Godwin-Austen coll.; NHMUK 1903.7.1.2566 .</p><p>Type localities</p><p>“Habiang Garo, on the West Khasi border” ( A. habiangensis); “Khasi Hills” ( A. generosus).</p><p>Differential diagnosis</p><p>Both D. subculmen and D. diagonius have a longer R2, a ribbed R1, and a less bulging R3. See also under D. nitidus .</p><p>Remarks</p><p>The only difference between the two syntypes of A. generosus and the single, subadult syntype of A. habiangensis is the very slightly shallower R3 dent of the latter, suggesting that they represent the same species. The original description of A. habiangensis immediately followed that of A. generosus, and in the face of the nearly identical shells, they were not compared with each other; except for the umbilical view, they were not illustrated from the same views. We selected A. habiangensis to be the accepted name because it is known from a more precise locality than A. generosus .</p></div>	https://treatment.plazi.org/id/03CC1E69A36FFF8DBAC0FC65B2B9F86A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFF8FB960FEECB74DFB78.text	03CC1E69A36DFF8FB960FEECB74DFB78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax hebes (Benson 1857)	<div><p>Dicharax hebes (Benson, 1857)</p><p>Fig. 53F–J</p><p>Alycaeus hebes Benson, 1857: 204–205 .</p><p>Alycaeus hebes – Sowerby 1877: pl. 6, species 52. — Godwin-Austen 1886: 191, pl. 43 figs 1, 1a–c;</p><p>1914: 374–375, pl. 145 figs 5, 5a–b. Alycaeus (Dicharax) hebes – Kobelt 1902: 371. — Gude 1921: 255. Chamalycaeus (Dicharax) hebes – Ramakrishna et al. 2010: 61. Dicharax hebes – Páll-Gergely et al. 2017: 10, fig. 5; 2020: 64, fig. 13a; 2024, fig. 12d. — Preece et al.</p><p>2022: 80, fig. 31c–d.</p><p>Diagnosis</p><p>The relatively large shell, the nearly smooth R1, the formation of the R3 swelling and the rounded aperture distinguishes this species from its congeners.</p><p>Type material examined</p><p>INDIA • neotype (designated by Preece et al. 2022: Fig. 53F–J); Teria Ghat is in the Khasi Hills; Theobald leg.; NHMUK 1888.12.4.909 .</p><p>Additional material examined</p><p>INDIA • 3 shells; Khasi Berge; Möllendorff coll.; SMF 109244 • 1 shell; Nongphrian, Khasi Hills; Dr Alfred Oberwimmer coll.; NHMW (mixed sample with D. theobaldi) • 1 shell; same data as for preceding; NHMW 14715 (mixed sample with D. theobaldi) .</p><p>Type locality</p><p>“ad Teria Ghát”.</p></div>	https://treatment.plazi.org/id/03CC1E69A36DFF8FB960FEECB74DFB78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A36DFFB1B913FB09B7AEFD12.text	03CC1E69A36DFFB1B913FB09B7AEFD12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax nitidus (W. T. Blanford 1862)	<div><p>Dicharax nitidus (W.T. Blanford, 1862)</p><p>Fig. 55</p><p>Alycaeus nitidus W.T. Blanford, 1862: 141 .</p><p>Alycaeus kurzianus Theobald &amp; Stoliczka, 1872: 330, pl. 11 fig. 2. Syn. nov.</p><p>Alycaeus kurzianus – Sowerby 1877: pl. 3, species 22. — Godwin-Austen 1914: 409–410, pl. 151 figs 7, 7a.</p><p>Alycaeus nitidus – Sowerby 1877: pl. 3, species 25. — Godwin-Austen 1914: 421–422, pl. 151 figs 4, 4a.</p><p>Alycaeus (Chamalycaeus) nitidus – Kobelt 1902: 360. — Gude 1921: 230–231.</p><p>Alycaeus (Dicharax) kurzianus – Kobelt 1902: 373. — Gude 1921: 258–259.</p><p>Chamalycaeus (Chamalycaeus) nitidus – Ramakrishna et al. 2010: 54.</p><p>Dicharax nitidus – Páll-Gergely et al. 2020: 68.</p><p>Dicharax (?) kurzianus – Páll-Gergely et al. 2020: 90, fig. 91.</p><p>Type material examined</p><p>MYANMAR • 3 possible syntypes of A. nitidus (Fig. 55A–E); Manya Khyoung, Arakan; Blanford coll.; NHMUK 1906.4.4.54 • 3 probable syntypes of A. kurzianus (labelled as “ kurtzianus ”; the locality does not match with the original description, but the specimens are identical to the figured one) (Fig. 55F–J); Nr. Prome, Pegu; F. Stoliczka leg.; NHMUK 1903.7.1.2700 • 1 syntype of A. kurzianus; Nattoung, Prome; W. Theobald coll.; NZSI M.24974 • 6 syntypes of A. kurzianus; same data as for preceding; NZSI M.25021 .</p><p>Type localities</p><p>“prope Tongoop in Arakan ” ( A. nitidus); “Nattoung in provincia Barmana, Prome dicta” ( A. kurzianus).</p><p>Differential diagnosis</p><p>The most similar species due to the tiny, glossy shell and the short R2 is D. habiangensis, which has a larger R3 swelling and a basally rounded aperture. See also under D. semivivus sp. nov.</p><p>Remarks</p><p>Dicharax nitidus and D. kurzianus agree in all important traits (shell and aperture shape, shell size, length of shell regions, sculpture), and no notable difference was found between them. Therefore, D. kurzianus is a junior synonym of D. nitidus . The two species were not compared in the original description of D. kurzianus .</p><p>Dicharax nitidus was described from Tongoop (Taungup Pass at 18°40.1′ N, 94°44.5′ E), whereas D. kurzianus from Nattoung, which could not be located on the map. According to the additional information about Nattung, it is in “Mendon District, Pegu ”, and close to Prome (= Pyay). A mountain with the name “Nattaung” is located at 18°48.6′ N, 97°2.2′ E, ca 190 km east from Pyay, therefore it is probably not the same as Nattung. Nevertheless, the Taungup Pass and Pyay are situated geographically close to each other.</p></div>	https://treatment.plazi.org/id/03CC1E69A36DFFB1B913FB09B7AEFD12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A353FFB1B91DFD37B4D2FA09.text	03CC1E69A353FFB1B91DFD37B4D2FA09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax notatus (Godwin-Austen 1876)	<div><p>Dicharax notatus (Godwin-Austen, 1876)</p><p>Fig. 53K–O</p><p>Alycaeus notatus Godwin-Austen, 1876: 176, pl. 7 figs 9, 9a, b.</p><p>Alycaeus notatus – Godwin-Austen 1886: 191–192, pl. 43 figs 2, 2a–c; 1914: 358–359, pl. 145 figs 8, 8a. Alycaeus (Dicharax) notatus – Kobelt 1902: 374. — Gude 1921: 262.</p><p>Chamalycaeus (Dicharax) notatus – Ramakrishna et al. 2010: 64.</p><p>Dicharax notatus – Páll-Gergely et al. 2020: 68.</p><p>Diagnosis</p><p>The elevated spire, relatively dense R1 ribs and the strongly inflated R3 swelling distinguishes this species from its congeners. See under D. damsangensis .</p><p>Type material examined</p><p>INDIA • 4 syntypes (Fig. 53K–O); Toruputu Peak, Dafla Hills; elev. 3000 ft; NHMUK 1903.7.1.2672 • 2 syntypes; Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7.1.2544. Both samples are in the same box but in different vials .</p><p>Type locality</p><p>“On the slopes of Torúpútú Peak at 3000 feet ”.</p></div>	https://treatment.plazi.org/id/03CC1E69A353FFB1B91DFD37B4D2FA09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A353FFB0BAC5FA3AB5A7FAE5.text	03CC1E69A353FFB0BAC5FA3AB5A7FAE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax nowgongensis (Godwin-Austen 1914)	<div><p>Dicharax nowgongensis (Godwin-Austen, 1914)</p><p>Fig. 56</p><p>Alycaeus nowgongensis Godwin-Austen, 1914: 397, pl. 137, figs 4, 4a–b.</p><p>Alycaeus nowgongensis – Gude 1921: 213.</p><p>Alycaeus (Alycaeus) nowgongensis – Ramakrishna et al. 2010: 49.</p><p>Dicharax nowgongensis – Páll-Gergely et al. 2020: 69.</p><p>Type material examined</p><p>INDIA • holotype (single shell mentioned in the original description: Fig. 56A–G); Koliaghur nr Tezpur, Assam; NHMUK 1903.7.1.2682 .</p><p>Type locality</p><p>“Koliaghur or Koliahur, Nowgoug District, Assam ”.</p><p>Differential diagnosis</p><p>Similar to D. chennelli, D. damsangensis, D. diagonius and D. vitreus, which have similar R1 sculptures. Unfortunately, the holotype specimen is broken at its R3, making it difficult to examine some key characters. Nevertheless, D. nowgongensis has a rounded aperture and a relatively low-spired shell, which differs from most congeners of the same species group.</p></div>	https://treatment.plazi.org/id/03CC1E69A353FFB0BAC5FA3AB5A7FAE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A352FFB2BACFFA65B799F9D9.text	03CC1E69A352FFB2BACFFA65B799F9D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax praeda Pall-Gergely & Aravind 2025	<div><p>Dicharax praeda Páll-Gergely &amp; Aravind sp. nov.</p><p>urn:lsid:zoobank.org:act: B752AA96-1639-4C84-8102-4798657CCCD9</p><p>Figs 12B–C, 57</p><p>Diagnosis</p><p>A small species of Dicharax with glossy R1 and R3, a combined length of R2+R3 reaching nearly half a whorl, and a strongly inflated R3 bearing a single dent near the suture on the upper part of the R3 swelling.</p><p>Etymology</p><p>The specific epithet ‘ praeda ’ (Latin for ‘prey’) refers to the dent on R3 resembling a bite trace. To be used as a noun in apposition.</p><p>Type material examined</p><p>Holotype</p><p>INDIA – Mizoram • empty shell (D: 3 mm, H: 2.4 mm, Figs 12B–C, 57); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 67; NZSI LM1740. Paratype</p><p>INDIA – Mizoram • 1 empty shell; Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 67; NZSI LM1741 .</p><p>Description</p><p>Shell small, yellowish, glossy, semitranslucent; shell outline rather rounded in dorsal view except for the conspicuously inflated R3; spire elevated, conical; body whorl rounded; protoconch low, glossy, consisting of ca 1.5 whorls; R1 of 1.75–2 whorls, glossy, smooth, with some inconspicuous, widely-spaced, incomplete (i.e., present only near the suture) ribs on the last ca 0.75 whorl of R1; spiral striation absent; boundary between R1 and R2 conspicuous due to the sudden change to a regularly ribbed surface; R2 with ca 22–26 straight, sharp, lamella-like ribs; R2 ribs slightly curved towards aperture, the spaces between the ribs are wider than the width of a single rib; R2+R3 nearly half whorl; R3 slightly longer than R2; boundary between R2 and R3 clearly visible due to a deep constriction; entire R3 strongly swollen, forming a blunt tip towards constriction; a shallow, inconspicuous, elongated dent is situated near the upper end of the swelling, nearly touching the suture; aperture strongly oblique to shell axis, rounded with a slight upper (parieto-palatal) incision; boundary between inner and outer peristomes conspicuous; inner peristome slightly thickened and protruding; outer peristome more conspicuous, moderately sharp, strongly expanded, especially on the upper palatal and basal side; umbilicus very narrow, slit-like.</p><p>MEASUREMENTS. D: 2.9–3 mm, H: 2.4 mm.</p><p>Distribution</p><p>Known only from the Blue Mountain region.</p><p>Differential diagnosis</p><p>The most similar species is Dicharax diagonius, which has complete R1 ribs on the entire region (absent on the first 1.5 whorl, and on the last ca 1 whorl present only near the suture in the new species), more curved R2 ribs, and a more pointed tip of the R3 swelling.</p><p>Dicharax chennelli differs from the new species in the following characteristics: the ribs near the suture of R1 are stronger in D. chennelli, and the R2 ribs are more curved towards the aperture, forming an almost continuous flat area, while the R2 ribs of the new species are rather straight, and therefore they do not form a flat surface. The dorsal side of the R3 swelling of D. chennelli bears two dents, one situated on the upper part, and the lower at its middle, both being the same distance from the suture. In contrast, the upper dent nearly touches the suture in the new species, and the lower dent is not visible. Furthermore, the tip of the swelling (i.e., the part situated closest to the constriction) is sharper in D. chennelli, and blunter in the new species. Lastly, D. chennelli has a basal incision, while the aperture of the new species is perfectly rounded.</p><p>Dicharax habiangensis has a shorter R2 and R3, the R2 ribs are more curved, the dent on the R3 is situated at some distance from the suture (touching the suture in the new species), and the entire R3 is separated from the penultimate whorl by the deeper suture.</p><p>Dicharax subculmen has a lower spire, denser ribbing on R1, and a deeper dent on the R3 swelling. Dicharax nowgongensis has stronger and more complete R1 ribs, but the R3 of the holotype of that species was broken.</p></div>	https://treatment.plazi.org/id/03CC1E69A352FFB2BACFFA65B799F9D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A350FFB5B936F969B76DFDEA.text	03CC1E69A350FFB5B936F969B76DFDEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax subculmen (Godwin-Austen 1893)	<div><p>Dicharax subculmen (Godwin-Austen, 1893)</p><p>Fig. 52K–O</p><p>Alycaeus subculmen Godwin-Austen, 1893: 593 .</p><p>Alycaeus subculmen – Godwin-Austen 1897: 4, pl. 63 figs 4, 4a; 1914: 398.</p><p>Alycaeus (Dicharax) subculmen – Kobelt 1902: 377. — Gude 1921: 270–271.</p><p>Chamalycaeus (Dicharax) subculmen – Ramakrishna et al. 2010: 67.</p><p>Dicharax (?) subculmen – Páll-Gergely et al. 2020: 105.</p><p>Diagnosis</p><p>The small shell, the rather weak R1 ribs, and the relatively less swollen R3 swelling distinguish this species from its congeners. See also under D. habiangensis and Dicharax praeda sp. nov.</p><p>Type material examined</p><p>INDIA • 3 syntypes (Fig. 52K–O); Naga Hills; Doherty leg.; NHMUK 1903.7.1.2687 .</p><p>Type locality</p><p>“ Naga Hills”.</p></div>	https://treatment.plazi.org/id/03CC1E69A350FFB5B936F969B76DFDEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A357FFB5BAE9FD9FB5A7FBFA.text	03CC1E69A357FFB5BAE9FD9FB5A7FBFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax vitreus Gittenberger & Kezang Tobgay 2024	<div><p>Dicharax vitreus Gittenberger &amp; Kezang Tobgay, 2024</p><p>Fig. 58</p><p>Dicharax vitreus Gittenberger &amp; Kezang Tobgay in Gittenberger et al., 2024: 205, figs 25–27.</p><p>Type locality</p><p>“ Dagana Dzongkhag: [302] Lhamoizingkha, Pubji near Quarry, ca 250 m a.s.l., 26º45’N 89º48’E ” (Bhutan).</p><p>Differential diagnosis</p><p>Similar to D. chennelli and D. diagonius, but the R2 ribs are seemingly more widely-spaced (although the photos in the original description of D. vitreus are of low quality). Moreover, D. vitreus lacks the basal incision in the aperture, which is present in D. chennelli .</p></div>	https://treatment.plazi.org/id/03CC1E69A357FFB5BAE9FD9FB5A7FBFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A356FFB4B92CFE39B43EF80F.text	03CC1E69A356FFB4B92CFE39B43EF80F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax akhaensis (Godwin-Austen 1914)	<div><p>Dicharax akhaensis (Godwin-Austen, 1914)</p><p>Fig. 59</p><p>Alycaeus akhaensis Godwin-Austen, 1914: 352, pl. 141 figs 1, 1a–b.</p><p>Alycaeus burroiensis Godwin-Austen, 1914: 354, pl. 141 figs 6, 6a. Syn. nov.</p><p>Alycaeus gemma Godwin-Austen, 1914: 355–356, pl. 149 figs 6, 6a. Syn. nov.</p><p>Alycaeus (Dicharax) gemma – Gude 1921: 252.</p><p>Alycaeus (Cycloryx) burroiensis – Gude 1921: 277.</p><p>Alycaeus (Raptomphalus) akhaensis – Gude 1921: 286.</p><p>Chamalycaeus (Dicharax) gemma – Ramakrishna et al. 2010: 60.</p><p>Chamalycaeus (Raptomphalus) akhaensis – Ramakrishna et al. 2010: 68.</p><p>Dicharax (?) akhaensis – Páll-Gergely et al. 2020: 76.</p><p>Dicharax (?) burroiensis – Páll-Gergely et al. 2020: 80.</p><p>Dicharax (?) gemma – Páll-Gergely et al. 2020: 88.</p><p>Diagnosis</p><p>This species is characterized by strong, widely-spaced R1 ribs, a short R2+R3, a smooth R2, two low R3 swellings and a slightly fringed peristome.</p><p>Type material examined</p><p>INDIA • holotype of A. akhaensis (single shell mentioned in the original description: Fig. 59A–E); Akha Hills, Barowli River; Godwin-Austen coll.; NHMUK 1903.7.1.2683 • 1 syntype of A. burroiensis (Fig. 59F–J); Burroi Rr., Dafla; NHMUK 1903.7.1.2653 • 3 corroded syntypes of A. gemma in two vials (Fig. 59K–O); No. 7. camp, Dikrang Valley, Dafla; NHMUK 1903.7.1.2601 .</p><p>Type localities</p><p>“Barowli Gorge, Durrang District, Assam, foot of the Akha Hills” ( A. akhaensis); “Burroi Gorge, Dafla Hills” ( A. burroiensis); “No. 7 Camp, Dikrang Valley, Dafla Hills” ( A. gemma).</p><p>Differential diagnosis</p><p>Dicharax lahupaensis (Godwin-Austen, 1914) has a ribbed R2, and its aperture is less fringed.</p><p>Remarks</p><p>Both A. akhaensis and A. burroiensis are represented in the NHM by single, corroded specimens. Acknowledging that the important characters (mostly the fine sculpture and the formation of R3) could not be very clearly seen on the shells, we could not find any conchological differences that would justify their distinction. Therefore, we treat A. burroiensis as a junior synonym of Dicharax akhaensis . Alycaeus gemma also agrees in all important shell characters (ratios of shell regions, R3 swellings, fringed aperture), but its shell has a rather oval outline in dorsal view. Nevertheless, this is also considered part of intraspecific variability, and therefore, A. gemma is also treated as a junior synonym of A. akhaensis . See also under D. khasiacus .</p></div>	https://treatment.plazi.org/id/03CC1E69A356FFB4B92CFE39B43EF80F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A356FFB4B9BFFEECB495FE08.text	03CC1E69A356FFB4B9BFFEECB495FE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 8</p><p>Diagnosis</p><p>This species group is characterized by mostly small (smaller than 3 mm), mostly glossy species. The aperture is not fringed or slightly fringed.</p></div>	https://treatment.plazi.org/id/03CC1E69A356FFB4B9BFFEECB495FE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A354FFB9B92FFEECB474FA87.text	03CC1E69A354FFB9B92FFEECB474FA87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax birugosus (Godwin-Austen 1893)	<div><p>Dicharax birugosus (Godwin-Austen, 1893)</p><p>Fig. 60</p><p>Alycaeus bi-rugosus Godwin-Austen, 1893: 593.</p><p>Alycaeus birugosus var. minor Godwin-Austen, 1914: 370, pl. 155 figs 9, 9a.</p><p>Alycaeus bi-rugosus – Godwin-Austen 1897: 387, pl. 63 figs 5, 5a.</p><p>Alycaeus (Dicharax) birugosus – Kobelt 1902: 366.</p><p>Alycaeus birugosus – Godwin-Austen 1914: 370, 387.</p><p>Alycaeus canaliculus Godwin-Austen, 1914: 371, pl. 154 fig. 11.</p><p>Alycaeus birugosus var. – Godwin-Austen 1914: 370, pl. 154 figs 7, 7a.</p><p>Alycaeus (Chamalycaeus) canaliculus – Gude 1921: 225.</p><p>Alycaeus (Dicharax) birugosus – Gude 1921: 239–240.</p><p>Alycaeus (Dicharax) birugosus var. minor – Gude 1921: 240.</p><p>Chamalycaeus (Chamalycaeus) canaliculus – Ramakrishna et al. 2010: 53.</p><p>Chamalycaeus (Dicharax) birugosus – Ramakrishna et al. 2010: 57.</p><p>Dicharax (?) birugosus – Páll-Gergely et al. 2020: 79, fig. 19 (treated A. canaliculus as a junior synonym).</p><p>Diagnosis</p><p>The characteristic, oblique main R3 swelling, and the small secondary swelling distinguish this species from its congeners. See under Dicharax umashaankeri sp. nov.</p><p>Type material examined</p><p>INDIA • 2 syntypes of A. birugosus (Fig. 60A–E); Khasi Hills; Godwin-Austen leg.; NHMUK 1903.7.1.2628 • 1 syntype of Alycaeus birugosus var. minor (labelled as duorugosus var. minor); Garo Hills; NHMUK 1903.7.1.2755 • 1 syntype of A. canaliculus (Fig. 60F–J); Teria Ghat, Khasi; Godwin-Austen leg.; NHMUK 1903.7.1.2764.</p><p>Additional material examined</p><p>INDIA • 7 shells (labelled as “ birugosus var.”); Jawai, Jiantia Hills, 282a; Godwin-Austen leg., NHMUK 1903.7.1.2571.</p><p>Type localities</p><p>“Khasi Hills and Manipur ” ( A. birugosus); “Garo Hills” ( A. birugosus var. minor); “Teria Ghat, foot of the Khasi Hills” ( A. canaliculus).</p></div>	https://treatment.plazi.org/id/03CC1E69A354FFB9B92FFEECB474FA87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35BFFBDB92CFA84B4B4FE33.text	03CC1E69A35BFFBDB92CFA84B4B4FE33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax daflaensis (Godwin-Austen 1876)	<div><p>Dicharax daflaensis (Godwin-Austen, 1876)</p><p>Fig. 61A–E</p><p>Alycaeus daflaensis Godwin-Austen, 1876: 176–177, pl. 7 figs 12, 12a–b.</p><p>Alycaeus (Dicharax) daflaensis – Kobelt 1902: 368. — Gude 1921: 245–246.</p><p>Alycaeus daflaensis – Godwin-Austen 1914: 354–355, pl. 145 figs 11, 11a–b.</p><p>Chamalycaeus (Dicharax) daflaensis – Ramakrishna et al. 2010: 58.</p><p>Dicharax (?) daflaensis – Páll-Gergely et al. 2020: 84.</p><p>Type material examined</p><p>INDIA • lectotype (designated by Páll-Gergely et al. 2020) (Fig. 61A–E); Toruputu Peak, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7.1.2497.1 • 6 paralectotypes; same data as for preceding; NHMUK 1903.7.1.2497.2-7 .</p><p>Type locality</p><p>“Torúpútú Peak, 7000 feet ”.</p><p>Differential diagnosis</p><p>Within this species group, D. gemmula is the most similar due to the similarly shaped R3 swelling, the smooth R2 and the overall smooth shell surface. Nevertheless, the aperture of D. gemmula is not fringed. Dicharax lahupaensis is also similar, but it has elevated R2 ribs (smooth in D. daflaensis), and its R3 swelling is situated at a greater distance from the aperture. Dicharax pachitaensis (Godwin-Austen, 1886) has a less elevated R3 swelling, stronger R1 ribs, more rounded aperture, and a keeled umbilical part of R2. See also under D. crenatus .</p></div>	https://treatment.plazi.org/id/03CC1E69A35BFFBDB92CFA84B4B4FE33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFBDB934FE57B739FB05.text	03CC1E69A35FFFBDB934FE57B739FB05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax dalingensis (Godwin-Austen 1914)	<div><p>Dicharax dalingensis (Godwin-Austen, 1914)</p><p>Fig. 62A–E</p><p>Alycaeus dalingensis Godwin-Austen, 1914: 338–339, pl. 134 figs 3, 3a–c.</p><p>Alycaeus (Dicharax) dalingensis – Gude 1921: 246.</p><p>Dicharax (?) dalingensis – Páll-Gergely et al. 2020: 85.</p><p>Type material examined</p><p>INDIA • 7 syntypes (Fig. 62A–E); Rechila Pk, Sikkim; W. Robert leg.; NHMUK 1903.7.1.1251 .</p><p>Type locality</p><p>“Rechila Peak, Daling District, on Sikkim and Bhutan Boundary (10,300 ft.)”.</p><p>Differential diagnosis</p><p>The most similar species are D. lenticulus (Godwin-Austen, 1874), D. nongtungensis (Godwin-Austen, 1914) and D. politus (W.T. Blanford, 1865), due to the smooth shell surface and the long, inflated R3 swelling. Dicharax lenticulus is larger and has a more depressed shell. Dicharax nongtungensis is smaller, more depressed, and has a more strongly expanded outer peristome. Lastly, D. politus is much more depressed, its R2 ribs are separated (R2 surface is smooth in D. dalingensis), and has a more inflated R3 swelling. See also under D. ganeshaiahi sp. nov. and D. nattoungensis (Godwin-Austen, 1914) .</p></div>	https://treatment.plazi.org/id/03CC1E69A35FFFBDB934FE57B739FB05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A35FFFA1BAEFFB05B4FFFA29.text	03CC1E69A35FFFA1BAEFFB05B4FFFA29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax ganeshaiahi Pall-Gergely & Aravind 2025	<div><p>Dicharax ganeshaiahi Páll-Gergely &amp; Aravind sp. nov.</p><p>urn:lsid:zoobank.org:act: ADABA8B7-0D4C-4330-83F7-4CBE3CC2604A</p><p>Figs 64, 65A–C, 66A–B</p><p>Diagnosis</p><p>A small species of Dicharax with a glossy shell, smooth, short R2 (combined length of R2+R3 ca quarter whorl, R3 ca twice as long as R2), an R3 with a central, blunt swelling, a rounded aperture with shallow upper and lower incisions, and a strongly expanded outer peristome.</p><p>Etymology</p><p>This species is named after Prof. K.N. Ganeshaiah, Founder, Trustee of Ashoka Trust for Research in Ecology and the Environment (ATREE), Bangalore, India. He has contributed immensely to the understanding of the ecology, evolution, and conservation of Indian flora and fauna. He is also a popular science communicator in the vernacular language and has published over 32 books in the Kannada language.</p><p>Type material examined</p><p>Holotype</p><p>INDIA – Mizoram • empty shell (D: 3.2 mm, H: 2.3 mm, Fig. 64); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 62; NZSI LM1755.</p><p>Paratypes</p><p>INDIA – Mizoram • 8 empty shells; Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 62; NZSI LM1756 • 2 empty shells; same data as for preceding; Ar 47; NZSI LM1757 • 1 empty shell (Fig. 65A–C, 66A–B); same data as for preceding; Ar 27; NZSI LM1758 • 6 empty shells; same data as for preceding; Ar 72; NZSI LM1759 .</p><p>Additional material examined</p><p>INDIA – Mizoram • 1 corroded empty shell; same data as for holotype; Ar 47; NZSI LM1760 • 6 corroded empty shells; same data as for holotype; Ar 72; NZSI LM1761 .</p><p>Description</p><p>Shell small, yellowish or pinkish when fresh, semitranslucent; entire shell surface glossy; shell outline rounded in dorsal view, R2 and end of R3 conspicuously swollen, although not in all specimens; spire elevated, conical; body whorl rounded; protoconch low, smooth, consisting of 1.25–1.5 whorls; R1 of ca 1.5–1.75 whorls, smooth, glossy, last ca quarter whorl of some specimens with weak, widely-spaced ribs, while in other specimens entire R3 remains smooth; some weak ribs of the last ca quarter whorl visible inside umbilicus; boundary between R1 and R2 not conspicuous, in the case of specimens with ribbed end of R1 the change between the two regions is more clearly visible due to the large increase in rib density; R2 without elevated ribs, but very densely arranged lighter and darker alternating stripes (more than 40 in number); cross sectional view of R2 was examined in a specimen (Fig. 66C–D): anterior crust forms no rib, it is entirely flat; instead of folding over the posterior crust, the posterior crust covers the larger part of the microtunnels, and the anterior crust only covers the edge of the posterior crust; microtunnel wide compared to anterior and posterior crusts, its cross sectional view teardrop-shaped; R2+R3 ca 90 degrees; R3 ca twice as long as R2; boundary between R2 and R3 clearly visible due to a shallow constriction; R3 with a low, central, triangular swelling; aperture strongly oblique to shell axis, rounded with a shallow upper (parieto-palatal), and a shallow lower (basal) incision; inner and outer peristomes clearly separated (mostly at the parietal, palatal and columellar part and less so at the basal part); inner peristome slightly protruding; outer peristome strongly expanded and reflected; umbilicus ca one fourth of shell width, almost rounded, although its part facing R2 is nearly straight, forming a slight keel on the umbilical side of R2.</p><p>OPERCULUM. Inner side smooth, without central nipple; outer side smooth, without any sculpture (Ar27).</p><p>MEASUREMENTS. D: 2.8–3.4 mm, H: 2.1–2.4 mm.</p><p>Differential diagnosis</p><p>Dicharax dalingensis (Fig. 62A–E) is similar in having a smooth R2, but has longer R2 and R3, and a higher spire.</p><p>Dicharax lenticulus (Fig. 63) also has a smooth R2, but is much larger, and has a longer R2 and R3. Moreover, both D. dalingensis and D. lenticulus have an incision between the penultimate whorl and peristome, while the peristome is attached to the penultimate whorl in the new species.</p><p>Dicharax pachitaensis (Fig. 61K–O), which is probably the most similar species, also has a smooth R2, but the combined length of R2 and R3 is usually (not always) longer than that of the new species. Moreover, D. pachitaensis has prominent ribs near the suture on the dorsal side, the R3 swelling is situated a greater distance from the peristome, and its aperture has deeper upper and lower incisions.</p><p>Remarks</p><p>The cross-sectional view of R2 of two species of Alycaeinae with smooth R2 is known: Dicharax moellendorffi (Kobelt &amp; Möllendorff, 1886) (see Páll-Gergely et al. 2017) and Metalycaeus minatoi Páll-Gergely, 2017 (see Páll-Gergely &amp; Asami 2017). Even though the R2 surfaces are similar, the fine structure of the R2 (i.e., how the anterior and posterior crusts fold over each other to make the microtunnels) is very different. In the case of D. moellendorffi, the anterior crust has a strong posterior projection that covers the anterior crust. In the case of Metalycaeus minatoi, the anterior and posterior crusts meet just above the microtunnel, and in the new species the posterior crust has an elongated anterior projection covering the microtunnel.</p></div>	https://treatment.plazi.org/id/03CC1E69A35FFFA1BAEFFB05B4FFFA29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A343FFA3B97AFA59B4A3FC5A.text	03CC1E69A343FFA3B97AFA59B4A3FC5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax gemmula (Benson 1859)	<div><p>Dicharax gemmula (Benson, 1859)</p><p>Fig. 67</p><p>Alycaeus gemmula Benson, 1859: 179–180 .</p><p>Alycaeus (Charax) subhumilis Möllendorff, 1897a: 41 . Syn. nov.</p><p>Alycaeus gemmula – Sowerby 1877: pl. 5, species 37. — Godwin-Austen 1886: 190, pl. 48 figs 4, 4a–c. Alycaeus (Dicharax) gemmula – Kobelt 1902: 370–371. — Gude 1921: 252–253.</p><p>Alycaeus (Dicharax) subhumilis – Kobelt 1902: 377. — Gude 1921: 271, pl. 1 figs 1–2.</p><p>Alycaeus (Charax) gemmula – Godwin-Austen 1914: 340.</p><p>Alycaeus (Charax) subhumilis – Godwin-Austen 1914: 344–345, pl. 133 figs 2–2c.</p><p>Chamalycaeus (Dicharax) subhumilis – Zilch 1957: 146, pl. 6 fig. 24. — Ramakrishna et al. 2010: 67. Chamalycaeus (Dicharax) gemmula – Ramakrishna et al. 2010: 60.</p><p>Dicharax (?) gemmula – Páll-Gergely et al. 2020: 89.</p><p>Dicharax (?) subhumilis – Páll-Gergely et al. 2020: 105, fig. 25.</p><p>Diagnosis</p><p>This species is characterized by a tiny, glossy shell, smooth R2 and a single, sharp R3 swelling, R2+R3 only slightly longer than a quarter whorl.</p><p>Type material examined</p><p>INDIA • neotype (designated by Preece et al. 2022: Fig. 67A–E); Darjiling; Blanford coll.; NHMUK 1906.4.4.55 • lectotype of A. subhumilis (designated by Zilch 1957: Fig. 67F–I); Vorderindien: Darjiling; Möllendorff coll.; SMF 109224 • 1 paralectotype of A. subhumilis; same data as for preceding; SMF 109225 .</p><p>Type localities</p><p>“in valle Rungun” ( A. gemmula); “in montibus Darjiling Indiae” ( A. subhumilis).</p><p>Differential diagnosis</p><p>Dicharax lahupaensis is similar in general shell and aperture shape, but it has two R3 swellings, and its R2 has separate ribs (R3 surface is smooth in D. gemmula). See also under D. daflaensis .</p><p>Remarks</p><p>No notable differences were found between the types of A. gemmula and A. subhumilis, therefore the latter is a junior synonym of the former.</p></div>	https://treatment.plazi.org/id/03CC1E69A343FFA3B97AFA59B4A3FC5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A341FFA5B92CFBEFB5E1FDB9.text	03CC1E69A341FFA5B92CFBEFB5E1FDB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax khasiacus (Godwin-Austen 1871)	<div><p>Dicharax khasiacus (Godwin-Austen, 1871)</p><p>Fig. 68</p><p>Alycaeus Khasiacus Godwin-Austen, 1871: 90, pl. 3 fig. 4.</p><p>Alycaeus duorugosus Godwin-Austen, 1914: 391 . Syn. nov.</p><p>Alycaeus khasiacus – Sowerby 1877: pl. 1, species 8. — Godwin-Austen 1914: 356, 376–377, 393, pl. 143 figs 7, 7a–b.</p><p>Alycaeus (Dicharax) khasiacus – Kobelt 1902: 372–373. — Gude 1921: 257–258.</p><p>Alycaeus khasiacus var. – Godwin-Austen 1914: 356.</p><p>Alycaeus (Dicharax) duorugosus – Gude 1921: 249.</p><p>Chamalycaeus (Dicharax) duorugosus – Ramakrishna et al. 2010: 59.</p><p>Chamalycaeus (Dicharax) khasiacus – Ramakrishna et al. 2010: 62.</p><p>Dicharax (?) khasiacus – Páll-Gergely et al. 2020: 90.</p><p>Dicharax (?) duorugosus – Páll-Gergely et al. 2020: 86.</p><p>Diagnosis</p><p>Dicharax khasiacus is characterized by the rounded outline, the short, smooth R2, and the two, anteriorly situated R3 swellings.</p><p>Type material examined</p><p>INDIA • 29 syntypes of A. khasiacus (Fig. 68A–E); Lailangkote, Khasi Hills; NHMUK 1903.7.1.2650 • 1 syntype of A. duorugosus (Fig. 68F–J); Burrail; Godwin-Austen coll.; NHMUK 1903.7.1.2771 .</p><p>Additional material examined</p><p>INDIA • 4 shells; Jawai, Assam; Oberwimmer coll.; NHMW • 2 shells; same data as for preceding; NHMW 14711 • 4 shells; Jawai, India; Dr. Leo P. Rušnov coll., ex coll. Dr W. Blume ex coll. W.F. Webb; NHMW 15168 .</p><p>Type localities</p><p>“On the highest parts of the Khasi and Jiantia Hills” ( A. khasiacus); “Burrail Range, Naga ” “Also Angaoluo Trigonometrical Station, No. 2572; South Barak, No. 2629, and Munipur, No. 2654 B.M.” ( A. duorugosus).</p><p>Differential diagnosis</p><p>Dicharax lahupaensis has a ribbed R2 (smooth in D. khasiacus) and a thicker peristome. Dicharax akhaensis is larger, has a shorter R3 swelling, and a less fringed aperture.</p><p>Remarks</p><p>The single examined syntype of A. duorugosus fits the morphological variability of the 29 syntypes of A. khasiacus; therefore the former is a junior synonym of the latter.</p></div>	https://treatment.plazi.org/id/03CC1E69A341FFA5B92CFBEFB5E1FDB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A347FFA5B938FDCAB48CFB16.text	03CC1E69A347FFA5B938FDCAB48CFB16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax lahupaensis (Godwin-Austen 1914)	<div><p>Dicharax lahupaensis (Godwin-Austen, 1914)</p><p>Fig. 61F–J</p><p>Alycaeus lahupaensis Godwin-Austen, 1914: 394, pl. 141 figs 3, 3a.</p><p>Alycaeus (Raptomphalus) lahupaensis – Gude 1921: 287–288.</p><p>Chamalycaeus (Raptomphalus) lahupaensis – Ramakrishna et al. 2010: 68.</p><p>Dicharax lahupaensis – Páll-Gergely et al. 2020: 66.</p><p>Diagnosis</p><p>This species is characterized by a ribbed R2, two low R3 swellings, a fringed aperture and a strongly expanded outer peristome. See also under D. akhaensis, D. daflaensis, D. gemmula and D. khasiacus .</p><p>Type material examined</p><p>INDIA • 10 syntypes (Fig. 61F–J); Gaziphimih, N.E. Munipur; NHMUK 1903.7.1.2655 .</p><p>Type locality</p><p>“Gaziphimi, Lahupa Naga Hills, Munipur”.</p></div>	https://treatment.plazi.org/id/03CC1E69A347FFA5B938FDCAB48CFB16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A347FFA4B92FFB2BB2B0FE05.text	03CC1E69A347FFA4B92FFB2BB2B0FE05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax lenticulus (Godwin-Austen 1874)	<div><p>Dicharax lenticulus (Godwin-Austen, 1874)</p><p>Fig. 63</p><p>Alycaeus lenticulus Godwin-Austen, 1874: 147 .</p><p>Alycaeus lenticulus – Godwin-Austen 1914: 340–341, pl. 136 figs 2, 2a.</p><p>Alycaeus (Dicharax) lenticulus – Gude 1921: 259–260.</p><p>Chamalycaeus (Dicharax) lenticulus – Ramakrishna et al. 2010: 63</p><p>Dicharax (?) lahupaensis – Páll-Gergely et al. 2020: 92.</p><p>Type material examined</p><p>INDIA • holotype (single shell mentioned in the original description: Fig 63F–J); Darjeeling; Godwin-Austen coll.; NZSI M.8075 • several shells (Fig. 63A–E); Rissom Pk. (W. Bhutan); Godwin-Austen Coll.; NHMUK 1903.07.01.2552 .</p><p>Type locality</p><p>“Darjeeling”.</p><p>Differential diagnosis</p><p>Dicharax nongtungensis is smaller, has a thicker peristome and a narrower umbilicus. Dicharax politus has a more depressed shell and a more strongly inflated R3 swelling. See also under D. dalingensis, D. ganeshaiahi sp. nov., and D. nattoungensis .</p><p>Remarks</p><p>This species was described based on a strongly corroded shell (Godwin-Austen 1874). Later, Godwin-Austen (1914) provided a more complete description and drawings based on multiple additional specimens. We received photos of the holotype (Fig. 63F–J) from the Zoological Survey of India, and could confirm that the figured shell (Fig. 63A–E) is conspecific with the holotype.</p></div>	https://treatment.plazi.org/id/03CC1E69A347FFA4B92FFB2BB2B0FE05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A346FFA4BAC0FE06B254FA88.text	03CC1E69A346FFA4BAC0FE06B254FA88.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax multirugosus (Godwin-Austen 1874)	<div><p>Dicharax multirugosus (Godwin-Austen, 1874)</p><p>Fig. 69A–E</p><p>Alycaeus multirugosus Godwin-Austen 1874: 149, pl. 3 fig. 7.</p><p>Alycaeus (Dicharax) multirugosus – Kobelt 1902: 373. — Gude 1921: 260.</p><p>Alycaeus multirugosus – Godwin-Austen 1914: 395–396, pl. 144 figs 7, 7a.</p><p>Chamalycaeus (Dicharax) multirugosus – Ramakrishna et al. 2010: 63.</p><p>Dicharax (?) multirugosus – Páll-Gergely et al. 2020: 97.</p><p>Type material examined</p><p>INDIA • 2 syntypes (Fig. 69A–E); head of Lanier River NE of Munipur; Godwin-Austen coll.; NHMUK 1903.7.1.2485 .</p><p>Type locality</p><p>“Hills at head of the Lanier River, Naga Hills, ca. 5–6,000 feet ”.</p><p>Remarks</p><p>An easily recognisable species based on the three R3 swellings.</p><p>Differential diagnosis</p><p>Due to the complicated R3 swelling, this species is similar to some species of the Dicharax species group 7, but differs from most of those species by the more depressed shell, the rounded aperture, and the R3 swelling separated into 3 parts. See also under D. umashaankeri sp. nov.</p></div>	https://treatment.plazi.org/id/03CC1E69A346FFA4BAC0FE06B254FA88	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A346FFA6BAC5FAB8B58EFEF8.text	03CC1E69A346FFA6BAC5FAB8B58EFEF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax nattoungensis (Godwin-Austen 1914)	<div><p>Dicharax nattoungensis (Godwin-Austen, 1914)</p><p>Fig. 70A–E</p><p>Alycaeus nattoungensis Godwin-Austen, 1914: 410–411, pl. 155 figs 15, 15a.</p><p>Alycaeus nattoungensis – Gude 1921: 212–213.</p><p>Dicharax (?) nattoungensis – Páll-Gergely et al. 2020: 98, fig. 24.</p><p>Type material examined</p><p>MYANMAR • holotype (single shell mentioned in the original description: Fig. 70A–E); Nattoung Hills, Mendon District, Pegu, Burma; Theobald coll.; NZSI M.8036 .</p><p>Type locality</p><p>“Nattoung Hills”.</p><p>Differential diagnosis</p><p>Dicharax dalingensis, D. lenticulus and D. politus have a more inflated R3 swelling preceded by a deeper constriction between R2 and R3. Dicharax nongtungensis has a more oval outline in dorsal view and a more expanded outer peristome (thicker peristome). Dicharax pusillus is similar in general shell shape, but is much smaller and has a more strongly expanded peristome.</p></div>	https://treatment.plazi.org/id/03CC1E69A346FFA6BAC5FAB8B58EFEF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34BFFA9BAC9FEECB762FB51.text	03CC1E69A34BFFA9BAC9FEECB762FB51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax nongtungensis (Godwin-Austen 1914)	<div><p>Dicharax nongtungensis (Godwin-Austen, 1914)</p><p>Fig. 70F–J</p><p>Alycaeus nongtungensis Godwin-Austen, 1914: 378, pl. 138 figs 5, 5a.</p><p>Alycaeus (Dicharax) nongtungensis – Gude 1921: 261.</p><p>Chamalycaeus (Dicharax) nongtungensis – Ramakrishna et al. 2010: 64.</p><p>Dicharax (?) nongtungensis – Páll-Gergely et al. 2020: 98. — Gittenberger et al. 2024: 210, figs 33, 35. (identification questionable).</p><p>Type material examined</p><p>INDIA • 21 syntypes (Fig. 70F–J); Nongtung; Godwin-Austen coll.; NHMUK 1903.7.1.2692 .</p><p>Additional material examined</p><p>INDIA • 2 shells; Nontung; Dr Leo P. Rušnov coll., ex coll. Dr. W. Blume, ex coll. W.F. Webb; NHMW 15144 • 4 shells; Nongtung, Burmah; NHMW 14890.</p><p>Type locality</p><p>“Nongtung, Jaintia Hills”.</p><p>Differential diagnosis</p><p>Dicharax politus is more depressed, has a deeper constriction and a more inflated R3 swelling. Dicharax pusillus is smaller, has a shorter R2 and a less expanded outer peristome. See also under D. dalingensis, D. lenticulus and D. nattoungensis .</p><p>Remarks</p><p>The specimen form Bhutan is indeed similar to the syntypes of D. nongtungensis, but has a less inflated R3 swelling.</p></div>	https://treatment.plazi.org/id/03CC1E69A34BFFA9BAC9FEECB762FB51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34BFFA9B911FAF2B7ECF869.text	03CC1E69A34BFFA9B911FAF2B7ECF869.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax notus (Godwin-Austen 1914)	<div><p>Dicharax notus (Godwin-Austen, 1914)</p><p>Fig. 71A–E</p><p>Alycaeus notus Godwin-Austen, 1914: 411, pl. 155 fig. 12.</p><p>Alycaeus (Dicharax) notus – Gude 1921: 262.</p><p>Dicharax notus – Páll-Gergely et al. 2020: 69; 2021: 32, figs 24–27. — Jirapatrasilp et al. 2021: 7, figs 4c–d, 5b, 6e, 7a–b.</p><p>Diagnosis</p><p>Dicharax notus and D. omissus are relatively widespread and variable species in northern Thailand and neighbouring central Myanmar. Regarding their conchological variability and identification, see Páll-Gergely et al. (2021).</p><p>Type material examined</p><p>MYANMAR • 15 syntypes (Fig. 71A–E); Fort Stedman, Burma; Woodthorpe coll.; NHMUK 1903.7.1.3065. See also Páll-Gergely et al. (2021) .</p><p>Type locality</p><p>“Fort Stedman, Burma ”.</p></div>	https://treatment.plazi.org/id/03CC1E69A34BFFA9B911FAF2B7ECF869	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34AFFABB920FEECB435FDEF.text	03CC1E69A34AFFABB920FEECB435FDEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax omissus (Godwin-Austen 1914)	<div><p>Dicharax omissus (Godwin-Austen, 1914)</p><p>Fig. 71F–J</p><p>Alycaeus omissus Godwin-Austen, 1914: 411, pl. 155 fig. 13.</p><p>Alycaeus (Chamalycaeus) omissus – Gude 1921: 231.</p><p>Dicharax (?) omissus – Páll-Gergely et al. 2020: 100; 2021: 38, figs 29–30. — Jirapatrasilp et al. 2021: 7, figs 6a, 8a–b.</p><p>Diagnosis</p><p>See under D. notus and Páll-Gergely et al. (2021).</p><p>Type material examined</p><p>THAILAND / MYANMAR • 2 syntypes (Fig. 71F–J); Siam and Shan boundary; Woodthorpe coll.; NHMUK 1903.7.1.1228. See also Páll-Gergely et al. (2021) .</p><p>Type locality</p><p>“ Siam and Shan boundary”.</p></div>	https://treatment.plazi.org/id/03CC1E69A34AFFABB920FEECB435FDEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A349FFABB93FFD93B2E3FAED.text	03CC1E69A349FFABB93FFD93B2E3FAED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax pachitaensis (Godwin-Austen 1886)	<div><p>Dicharax pachitaensis (Godwin-Austen, 1886)</p><p>Fig. 61K–O</p><p>Alycaeus pachitaensis Godwin-Austen, 1886: 190–191, pl. 48 figs 5, 5a–c.</p><p>Alycaeus (Dicharax) pachitaensis – Kobelt 1902: 374. — Gude 1921: 264.</p><p>Alycaeus pachitaensis – Godwin-Austen 1914: 359.</p><p>Chamalycaeus (Dicharax) pachitaensis – Ramakrishna et al. 2010: 65.</p><p>Dicharax (?) pachitaensis – Páll-Gergely et al. 2020: 100.</p><p>Diagnosis</p><p>This species is characterized by a smooth shell (with the exception of widely-spaced R1 ribs along the suture), a smooth R2, two low R3 swellings, a rounded aperture with upper and lower incisions, and a keeled umbilical part of R2. See also under D. daflaensis and D. ganeshaiahi sp. nov.</p><p>Type material examined</p><p>INDIA • 3 syntypes (Fig. 61K–O); Pachita village, Dafla Hills; Godwin-Austen coll.; NHMUK 1903.7.1.2614 .</p><p>Type locality</p><p>“Pachita village (Camp no. 7 of the Expeditionary Force, 1874–75), Dafla Hills, Assam ”.</p></div>	https://treatment.plazi.org/id/03CC1E69A349FFABB93FFD93B2E3FAED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A349FFAAB912FA9DB498FEEF.text	03CC1E69A349FFAAB912FA9DB498FEEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax politus (W. T. Blanford 1865)	<div><p>Dicharax politus (W.T. Blanford, 1865)</p><p>Fig. 62K–O</p><p>Alycaeus politus W.T. Blanford, 1865: 83–84 .</p><p>Alycaeus politus – Sowerby 1877: pl. 5, species 39. — Godwin-Austen 1914: 422, pl. 139 figs 5, 5a. — Gude 1921: 214–215.</p><p>Alycaeus (Alycaeus) politus – Kobelt 1902: 348.</p><p>Dicharax politus – Páll-Gergely et al. 2020: 71.</p><p>Diagnosis</p><p>This species is characterized by a glossy, depressed shell, a finely ribbed R2, a strongly inflated, long R3 swelling, and a deep constriction between R2 and R3. See also under D. dalingensis, D. lenticulus, D. nattoungensis and D. nongtungensis .</p><p>Type material examined</p><p>MYANMAR • 3 probable syntypes (Fig. 62K–O); Phungdo, Arakan; Blanford coll.; NHMUK 1906.4.4.178 .</p><p>Type locality</p><p>“Phuong do, near Cape Negrais, Arakan ”.</p></div>	https://treatment.plazi.org/id/03CC1E69A349FFAAB912FA9DB498FEEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFAAB923FE9CB2DAFBAB.text	03CC1E69A348FFAAB923FE9CB2DAFBAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax pusillus (Godwin-Austen 1871)	<div><p>Dicharax pusillus (Godwin-Austen, 1871)</p><p>Fig. 70K–O</p><p>Alycaeus pusillus Godwin-Austen, 1871: 89–90, pl. 3 fig. 3.</p><p>Alycaeus pusillus – Sowerby 1877: pl. 1, species 7. — Godwin-Austen 1914: 379–380, 398, pl. 143 figs 6, 6a–b. — Gude 1921: 215–216.</p><p>Alycaeus (Alycaeus) pusillus – Kobelt 1902: 348. — Ramakrishna et al. 2010: 50.</p><p>Dicharax (?) pusillus – Páll-Gergely et al. 2020: 102.</p><p>Diagnosis</p><p>This species is characterized by a small shell, a smooth R2, and the nearly smooth transition between R2 and R3 (no constriction between the two). See also under D. nattoungensis and D. nongtungensis .</p><p>Type material examined</p><p>INDIA • 5 syntypes (Fig. 70K–O); Jawai, Jaintia; NHMUK 1903.7.1.2688 .</p><p>Type locality</p><p>“near Jawai”, “on the banks of the Kopili river on the road from Jawai to Asálú, viâ Súfai”.</p></div>	https://treatment.plazi.org/id/03CC1E69A348FFAAB923FE9CB2DAFBAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A348FFACBADCFBD8B2DBFD35.text	03CC1E69A348FFACBADCFBD8B2DBFD35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax semivivus Pall-Gergely & Aravind 2025	<div><p>Dicharax semivivus Páll-Gergely &amp; Aravind sp. nov.</p><p>urn:lsid:zoobank.org:act: FDBFC556-730F-4891-9B87-ABDBD8B2EBA4</p><p>Fig. 72</p><p>Diagnosis</p><p>A small species of Dicharax with smooth R1, having some widely-spaced ribs on the last whorl of R1, a combined R2+R3 length of ca 150 degrees, an R3 with a single, low, central swelling, and a moderately fringed peristome.</p><p>Etymology</p><p>The specific epithet ‘ semivivus ’ (‘half-alive, almost dead’) refers to the condition of the holotype, which was corroded, but was probably alive when found.</p><p>Type material examined</p><p>Holotype</p><p>INDIA – Mizoram • empty shell (D: 2.4 mm, H: 1.8 mm, Fig. 72); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 45; NZSI LM1754.</p><p>Description</p><p>Shell small, off-white, semitranslucent; shell surface corroded but the holotype (single available specimen) was probably alive judging from the in situ presence of the operculum; shell outline slightly oval in dorsal view; spire elevated, conical; body whorl rounded; protoconch low, smooth, its boundary not visible due to corrosion consisting of 1.5 whorls; R1 of ca 1.75 whorls, corroded, nearly smooth, last whorl with widely-spaced, low ribs near suture and inside umbilicus, no ribs reach mid part of whorls; boundary between R1 and R2 conspicuous due to the sudden increase in rib density; R2 with ca 26 low ribs, spaces between the ribs are approximately the width of a single rib or narrower; R2+R3 ca 150 degrees; R3 slightly longer than R2; boundary between R2 and R3 clearly visible due to a deep constriction; R3 with a low central swelling, anterior and posterior slopes are declivous; aperture strongly oblique to shell axis, rounded with a shallow upper (parieto-palatal), a deep lower (basal) incision and a fringed palatal part; together with the upper and lower incisions, there are 5 incisions on the peristome; peristome thin overall, boundary between inner and outer peristomes conspicuous; inner peristome slightly protruding; outer peristome only slightly expanded, not reflected, undulated following the fringed inner peristome; umbilicus ca one fourth of shell width, nearly rounded.</p><p>OPERCULUM. The outer side is smooth, inner side could not be examined.</p><p>MEASUREMENTS. D: 2.4 mm, H: 1.8 mm.</p><p>Distribution</p><p>Known only from the Blue Mountain region.</p><p>Differential diagnosis</p><p>The most similar species is Dicharax nitidus (Fig. 55), but it is larger, and has a longer R3, with a more posteriorly-situated swelling. Dicharax jaintiacus (Fig. 32) is larger, has a higher spire and a narrower umbilicus, a more strongly expanded outer peristome and a more strongly fringed aperture.</p></div>	https://treatment.plazi.org/id/03CC1E69A348FFACBADCFBD8B2DBFD35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34EFFAFB922FD56B7D8FD23.text	03CC1E69A34EFFAFB922FD56B7D8FD23.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax theobaldi (W. T. Blanford 1862)	<div><p>Dicharax theobaldi (W.T. Blanford, 1862)</p><p>Fig. 69F–J</p><p>Alycaeus Theobaldi W.T. Blanford, 1862: 142–143 .</p><p>Alycaeus theobaldi var. diyungensis Godwin-Austen, 1914: 401–402, pl. 138 fig. 4. Syn. nov. Alycaeus maosmaiensis Godwin-Austen, 1922: 365, text figs. Syn. nov.</p><p>Alycaeus theobaldi – Sowerby 1877: pl. 5, species 44. — Godwin-Austen 1914: 359–360, pl. 149 figs 3, 3a–b; 1914: 382–383, pl. 145 figs 4, 4a.</p><p>Alycaeus (Dicharax) theobaldi – Kobelt 1902: 377–378. — Gude 1921: 272–273.</p><p>Chamalycaeus (Dicharax) theobaldi – Ramakrishna et al. 2010: 67.</p><p>Dicharax maosmaiensis – Páll-Gergely et al. 2020: 67.</p><p>Dicharax theobaldi – Páll-Gergely et al. 2020: 73.</p><p>Dicharax theobaldi diyungensis – Páll-Gergely et al. 2020: 73.</p><p>Diagnosis</p><p>The combination of the strong, widely-spaced R1 ribs, the relatively widely-spaced R2 ribs and the blunt, central R3 swelling distinguish this species from its congeners.</p><p>Type material examined</p><p>INDIA • 1 syntype of A. theobaldi (Fig. 69F–J); Khasi Hills, NHMUK 1906.4.4.60 • 12 syntypes of Alycaeus theobaldi var. diyungensis (Fig. 69F–J); Diyung Valley, N of Asalu; NHMUK 1903.7.1.2546 • 1 syntype of A. maosmaiensis (Fig. 69K–O) separated in a vial with pink wool + 4 additional syntypes; Maosmai, nr Cherrapoonjee, Khasi; NHMUK 20191067 .</p><p>Additional material examined</p><p>INDIA • 3 shells; Nongphrian, Khasi Hills; Dr Alfred Oberwimmer coll.; NHMW (mixed sample with D. hebes) • 1 shell; same data as for preceding; NHMW 14715 (mixed sample with D. hebes) .</p><p>Type localities</p><p>“in montibus Khasi” ( A. theobaldi); “Diyung Valley, north of Asalu, N. Cachar” ( A. theobaldi var. diyungensis); “Khasi Hills, near Cherrapunji, at the mouth of the Maosmai cave” ( A. maosmaiensis).</p><p>Remarks</p><p>According to the original description of Alycaeus theobaldi var. diyungensis, that variety differs from the typical A. theobaldi in the following traits: “This Diyung Valley species is a departure from A. theobaldi of the Khasi Hills in having the apex less high and conical, the suture more impressed, the shell more openly umbilicated; costulation next suture stronger” (Godwin-Austen 1914: 402). However, we do not see any of these differences comparing the type specimens of the two taxa. Therefore, we treat D. theobaldi diyungensis as a junior synonym of D. theobaldi .</p><p>The type specimens of D. theobaldi, D. theobaldi diyungensis and D. maosmaiensis are similar in shell size and shell shape, while the sculpture of R1 shows differences. Namely, D. theobaldi and D. theobaldi diyungensis have strong, widely-spaced ribs that are more or less regular at the beginning of R1, and become irregular and more widely-spaced towards its end, while D. maosmaiensis has a smooth beginning of R1, and at the end of R1 the ribs are relatively weak, and slightly denser than in the other two nominal taxa. Nevertheless, these differences are considered part of the intraspecific variability. A similar case (intraspecific variability in R1 rib density) was shown in the case of Dicharax notus (see Páll-Gergely et al. 2021).</p></div>	https://treatment.plazi.org/id/03CC1E69A34EFFAFB922FD56B7D8FD23	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A34DFF51BAF8FD27B2F5F982.text	03CC1E69A34DFF51BAF8FD27B2F5F982.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax umashaankeri Pall-Gergely & Aravind 2025	<div><p>Dicharax umashaankeri Páll-Gergely &amp; Aravind sp. nov.</p><p>urn:lsid:zoobank.org:act: DB661D38-F5BD-4993-96D6-BA0C40F17753</p><p>Figs 65D–F, 66C–D, 73</p><p>Diagnosis</p><p>A small species of Dicharax with glossy R1 and R3, a combined R2+R3 length of ca 150 degrees, an R3 with a clearly separated main swelling and a secondary swelling, and a rounded aperture with a slight upper incision.</p><p>Etymology</p><p>This species is named after Prof. R. Uma Shaanker, Founder Trustee of Ashoka Trust for Research in Ecology and the Environment (ATREE), Bangalore, India. He has contributed immensely to the understanding of ecology, evolution, and conservation of flora and fauna of the Western Ghats, India.</p><p>Type material examined</p><p>Holotype</p><p>INDIA – Mizoram • empty shell (D: 2.7 mm, H: 2 mm, Fig. 73); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 73; NZSI LM1742.</p><p>Paratypes</p><p>INDIA – Mizoram • 28 empty shells; same data as for holotype; NZSI LM1743 • 2 empty shells; same data as for holotype; Ar 58; NZSI LM1744 • 1 empty shell; same data as for holotype; Ar 75; NZSI LM1907 • 3 empty shells; same data as for holotype; Ar 40; NZSI LM1745 • 1 empty shell; same data as for holotype; Ar 50; NZSI LM1746 • 3 empty shells; same data as for holotype; Ar 63; NZSI LM1747 • 1 empty shell; same data as for holotype; Ar 25; NZSI LM1748 • 1 empty shell; same data as for holotype; Ar 24; NZSI LM1749 • 1 empty shell; same data as for holotype; Ar 15; NZSI LM1750 • 1 empty shell; same data as for holotype; Ar 15; NZSI LM1908 .</p><p>Additional material examined</p><p>INDIA – Mizoram • 2 juvenile empty shell (Fig. 65D–F, 66C–D); same data as for holotype; NZSI LM1751 • 1 broken shell; same data as for holotype; Ar 21, NZSI LM1752 • 3 corroded/broken shells; same data as for holotype; Ar 36, NZSI LM1753 .</p><p>Description</p><p>Shell small, light yellowish to orange or ochre, in some specimens with reddish apex; glossy, semitranslucent; shell outline slightly oval in dorsal view; spire slightly elevated, low conical; body whorl rounded; protoconch low, glossy, consisting of 1.25–1.5 whorls; R1 of ca 1.5 whorls, glossy, nearly smooth, with some inconspicuous, dense, low riblets, in some specimens ribs increase in strength on the last ca quarter whorl; spiral striation absent; boundary between R1 and R2 conspicuous due to the sudden change to a regularly, densely ribbed surface; R2 with ca 18–22 straight, sharp, lamella-like ribs; R2 ribs low, blunt, curved towards aperture, spaces between the ribs are approximately the width of a single rib; cross sectional view of R2 was examined in a specimen (Fig. 66C–D): anterior crust forms a significantly elevated rib, which has an anterior projection (i.e., the R2 ribs are curved towards the aperture); anterior crust folds over the much weaker posterior crust; microtunnel wide compared to ribs, its cross sectional view teardrop-shaped; R2+R3 ca 150 degrees; R3 slightly longer than R2; boundary between R2 and R3 clearly visible due to a deep constriction; R3 forms a blunt tip towards constriction and possesses two swellings; the upper swelling is visible only on the dorsal side, stops at the centre of the whorl, while the lower swelling (i.e., the one situated closer to the aperture) reaches the ventral side of shell; the dent separating the two swellings is elongated, beginning at the suture; aperture strongly oblique to shell axis, rounded with a very slight upper (parieto-palatal) incision; boundary between inner and outer peristomes conspicuous; inner peristome slightly thickened, protruding; outer peristome more conspicuous, sharp, strongly expanded, and reflected towards umbilicus; umbilicus very narrow, ca 1/5 –1/6 of shell width, elongated, oval.</p><p>OPERCULUM. Outer surface multispiral, without elevated lamina, inner side with an elevated, flat-topped nipple.</p><p>MEASUREMENTS. D: 2.6–3.1 mm, H: 2–2.2 mm.</p><p>Distribution</p><p>Known only from the Blue Mountain region.</p><p>Differential diagnosis</p><p>Dicharax akhaensis is similar in having two swellings and being glossy, but differs from the new species by having more widely-spaced R1 ribs, denser R2 ribs (and possibly lower, although the holotype is corroded), a fringed aperture, and a wider umbilicus.</p><p>Dicharax birugosus is similar in colour, size, sculpture and length of R1 and R2, but its R3 is shorter than that of the new species, the aperture is slightly fringed and has a deep basal incision. Moreover, the two swellings are arranged differently: the main swelling is oblique to the constriction in Dicharax birugosus, and rather parallel in the new species, while the area on which the secondary swelling is situated is much wider in the new species.</p><p>Dicharax subculmen belongs to the “Elephant Dicharax ” species group but is somewhat similar to this new species, at least in dorsal view. Nevertheless, the new species has a more depressed shell and a wider umbilicus, a deeper ditch between the primary and secondary R3 swellings, a longer R3 with a larger distance between the peristome and the secondary swelling, and a thinner peristome.</p><p>Dicharax kurzianus has a fringed aperture.</p><p>Dicharax multirugosus has a more depressed shell, and has three separate R3 swellings.</p></div>	https://treatment.plazi.org/id/03CC1E69A34DFF51BAF8FD27B2F5F982	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B3FF51B9BFF986B574F8B0.text	03CC1E69A3B3FF51B9BFF986B574F8B0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 9</p><p>Remarks</p><p>This species group is characterized by a low and blunt R3 swelling that is situated close to the outer peristome, and a slightly fringed to rounded peristome.</p></div>	https://treatment.plazi.org/id/03CC1E69A3B3FF51B9BFF986B574F8B0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B3FF50B977F8D1B7F7FD4E.text	03CC1E69A3B3FF50B977F8D1B7F7FD4E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax montanus (Nevill 1881)	<div><p>Dicharax montanus (Nevill, 1881)</p><p>Fig. 74A–E</p><p>Alycaeus montanus Nevill, 1881: 149, pl. 6 fig. 5.</p><p>Alycaeus (Chamalycaeus) montanus – Kobelt 1902: 359. — Gude 1921: 229.</p><p>Alycaeus montanus – Godwin-Austen 1914: 341–342, pl. 136 figs 3, 3a.</p><p>Chamalycaeus (Chamalycaeus) montanus – Ramakrishna et al. 2010: 54.</p><p>Dicharax (?) montanus – Páll-Gergely et al. 2020: 97, fig. 23.</p><p>Diagnosis</p><p>This species differs from the other species of this species group by the absence of an R3 swelling.</p><p>Type material examined</p><p>INDIA • 1 syntype (labelled as holotype) (Fig. 74A–E); Sikkim; elev. 11000 ft; Dr Stoliczka coll.; NZSI M.8082 .</p><p>Type locality</p><p>“ Sikkim, at 11,000 ft ”.</p></div>	https://treatment.plazi.org/id/03CC1E69A3B3FF50B977F8D1B7F7FD4E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B2FF50B938FCF1B5F5FAC2.text	03CC1E69A3B2FF50B938FCF1B5F5FAC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax rechilaensis (Godwin-Austen 1914)	<div><p>Dicharax rechilaensis (Godwin-Austen, 1914)</p><p>Fig. 74F–J</p><p>Alycaeus rechilaensis Godwin-Austen, 1914: 343–344, pl. 134 figs 2, 2a.</p><p>Alycaeus (Dicharax) rechilaensis – Gude 1921: 267.</p><p>Dicharax (?) rechilaensis – Páll-Gergely et al. 2020: 102.</p><p>Type material examined</p><p>INDIA • holotype (single shell mentioned in the original description: Fig. 74F–J); Rechila Pk, Sikkim; N. Robert leg.; NHMUK 1903.7.1.1252 .</p><p>Type locality</p><p>“Rechila Peak, Daling District, on Sikhim-Bhutan Frontier, 10,300 feet ”.</p><p>Differential diagnosis</p><p>Dicharax strangulatus has a shallower constriction between R2 and R3, and the R3 swelling of that species is less steep. See also under D. montanus and D. strigatus .</p></div>	https://treatment.plazi.org/id/03CC1E69A3B2FF50B938FCF1B5F5FAC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B2FF53B91EFA45B7C0FEEC.text	03CC1E69A3B2FF53B91EFA45B7C0FEEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax strangulatus (L. Pfeiffer 1846)	<div><p>Dicharax strangulatus (L. Pfeiffer, 1846)</p><p>Fig. 74K–O</p><p>Cyclostoma strangulatum L. Pfeiffer, 1846: 86 .</p><p>Alycaeus strangulatus – Sowerby 1877: pl. 6, species 47. — Godwin-Austen 1914: 337, pl. 136 figs 1, 1a. Alycaeus (Dicharax) strangulatus – Kobelt 1902: 376. — Gude 1921: 269.</p><p>Chamalycaeus (Dicharax) strangulatus – Ramakrishna et al. 2010: 66.</p><p>Dicharax strangulatus – Sajan et al. 2020: 523, figs 1a–b, 2a–l. — Páll-Gergely et al. 2020: 104.</p><p>Type material examined</p><p>INDIA • 1 shell (“possible syntype ”) (Fig. 74K–O); NHMUK 1856.9.15.18 • at least 5 shells; Mussoorie, N. W Himalaya,“figd”; NHMUK 1903.7.1.2501 • several shells; NHMUK 1928.7.28.85-104 (from general collection). See also newly collected specimens examined in Sajan et al. (2020).</p><p>Type locality</p><p>“Bengalia”.</p><p>Differential diagnosis</p><p>See under D. rechilaensis .</p></div>	https://treatment.plazi.org/id/03CC1E69A3B2FF53B91EFA45B7C0FEEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52BA8CFAB5B7D6F86E.text	03CC1E69A3B0FF52BA8CFAB5B7D6F86E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax bhucarinatus Gittenberger & Sherub Sherub 2024	<div><p>Dicharax bhucarinatus Gittenberger &amp; Sherub Sherub, 2024</p><p>Fig. 75A–D</p><p>Dicharax bhucarinatus Gittenberger &amp; Sherub Sherub in Gittenberger et al., 2024: 209, figs 27, 32.</p><p>Diagnosis</p><p>Differs from its congeners by the densely ribbed shell, and the keeled body whorl.</p><p>Type locality</p><p>“ Samdrup Jongkhar Dzongkhag: 7 km NNW of Daifam (= Jomotshangkha), Toka Phu, 940 m a.s.l., 26º57’N 92º04’E ” (Bhutan).</p><p>Differential diagnosis</p><p>As mentioned in the original description (Gittenberger et al. 2024), the only species of Alycaeinae in the region with a keeled body whorl is Chamalycaeus sculptilis, which has a beaded peristome. Nevertheless, D. bhucarinatus was described based on the corroded holotype and therefore, the sculpture could not be examined. Thus, it may belong to Chamalycaeus (more likely than Metalycaeus, as mentioned by Gittenberger et al. 2024).</p></div>	https://treatment.plazi.org/id/03CC1E69A3B0FF52BA8CFAB5B7D6F86E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52B947FB44B33BFA95.text	03CC1E69A3B0FF52B947FB44B33BFA95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 10</p><p>Remarks</p><p>This species group is characterized by dense ribbing on the entire shell, and rounded aperture.</p></div>	https://treatment.plazi.org/id/03CC1E69A3B0FF52B947FB44B33BFA95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B0FF52B924FEECB282FBC7.text	03CC1E69A3B0FF52B924FEECB282FBC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax strigatus (Godwin-Austen 1874)	<div><p>Dicharax strigatus (Godwin-Austen, 1874)</p><p>Fig. 74P–R</p><p>Alycaeus strigatus Godwin-Austen, 1874: 146–147, pl. 3 fig. 2.</p><p>Alycaeus (Chamalycaeus) strigatus – Kobelt 1902: 363. — Gude 1921: 234–235.</p><p>Alycaeus strigatus – Godwin-Austen 1914, 381, 401, pl. 144 figs 2, 2a–b.</p><p>Chamalycaeus (Chamalycaeus) strigatus – Ramakrishna et al. 2010: 56.</p><p>Dicharax (?) strigatus – Páll-Gergely et al. 2020: 105.</p><p>Material examined</p><p>The syntypes should be in the Indian Museum (see Nevill 1878), but they have not been located there; therefore, they are considered lost (Páll-Gergely et al. 2020).</p><p>Type locality</p><p>“ Assam ”.</p><p>Remarks</p><p>This species may be conspecific with D. rechilaensis, but in the absence of available specimens of D. strigatus, we cannot compare the two species properly. According to the original description of D. strigatus, the shell is 0.15 inch (ca 3.8 mm), while D. rechilaensis is 5 mm (Godwin-Austen 1914). The drawing of D. strigatus in lateral view (Fig. 74Q) agrees clearly with the photo of D. rechilaensis in the same view (Fig. 74G). However, the apical view (Fig. 74R) shows a shallower constriction than in D. rechilaensis (Fig. 74I). Because of the mismatch of the measurements and the anomalies of the figures, we maintain this as a valid species, until more information becomes available.</p></div>	https://treatment.plazi.org/id/03CC1E69A3B0FF52B924FEECB282FBC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B7FF55BA7CFEECB2DCFC84.text	03CC1E69A3B7FF55BA7CFEECB2DCFC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax bhudepressus Gittenberger, Choki Gyeltshen & Kezang Tobgay 2024	<div><p>Dicharax bhudepressus Gittenberger, Choki Gyeltshen &amp; Kezang Tobgay, 2024</p><p>Fig. 75E–H</p><p>Dicharax bhudepressus Gittenberger, Choki Gyeltshen &amp; Kezang Tobgay in Gittenberger et al., 2024: 211, figs 34–35.</p><p>Diagnosis</p><p>Not similar to any other species of Dicharax . Although no spiral striation was mentioned in the original description, this species rather resembles some species of Metalycaeus, such as M. heudei (Bavay &amp; Dautzenberg, 1900) (see Páll-Gergely et al. 2017). However, the photos in the original description are not of sufficient resolution to see the fine sculpture. We keep this species in Dicharax until we will have the opportunity to examine the holotype or additional material. Nevertheless, this species is not similar to any species of Metalycaeus from the Himalayas.</p><p>Type locality</p><p>“ Zhemgang Dzongkhag: 19.5 km SE of Tingtibi, 780 m a.s.l., 27°01’N 90°49’E ” (Bhutan).</p></div>	https://treatment.plazi.org/id/03CC1E69A3B7FF55BA7CFEECB2DCFC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B7FF54BA90FC85B292FE5A.text	03CC1E69A3B7FF54BA90FC85B292FE5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax crassipalatum Gittenberger & Sherub Sherub 2024	<div><p>Dicharax crassipalatum Gittenberger &amp; Sherub Sherub, 2024</p><p>Fig. 75I–L</p><p>Dicharax crassipalatum Gittenberger &amp; Sherub Sherub in Gittenberger et al., 2024: 209, figs 27, 30–31.</p><p>Diagnosis</p><p>The dense ribbing, simple peristome and the morphology of R3 swelling distinguish this species from all other Alycaeinae from the region.</p><p>Type locality</p><p>“ Mongar Dzongkhag: 3 km N of Mongar, 1070 m a.s.l., 27º19’N 91º13’E ” (Bhutan).</p></div>	https://treatment.plazi.org/id/03CC1E69A3B7FF54BA90FC85B292FE5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF54B901FD04B44EFAE4.text	03CC1E69A3B6FF54B901FD04B44EFAE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax avae (W. T. Blanford 1863)	<div><p>Dicharax avae (W.T. Blanford, 1863)</p><p>Fig. 76</p><p>Alycaeus avae W.T. Blanford, 1863: 323–324 .</p><p>Alycaeus avae – Sowerby 1877: pl. 3, species 20. — Godwin-Austen 1914: 406–407, pl. 151 fig. 6. Alycaeus (Dicharax) avae – Kobelt 1902: 365. — Gude 1921: 238.</p><p>Dicharax avae – Páll-Gergely et al. 2020: 64.</p><p>Diagnosis</p><p>This species can be recognized based on the relatively large, densely ribbed shell, and the strongly oblique aperture.</p><p>Type material examined</p><p>MYANMAR • 6 syntypes (Fig. 76); Shan Hills, E of Ava, Burma; Blanford coll.; NHMUK 1906.4.4.61 .</p><p>Type locality</p><p>“The hills east of Mandalay and Ava”.</p></div>	https://treatment.plazi.org/id/03CC1E69A3B6FF54B901FD04B44EFAE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF54B946FDEFB7F8FD07.text	03CC1E69A3B6FF54B946FDEFB7F8FD07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 11</p><p>Remarks</p><p>This species group is characterized by a relatively large, ribbed shell, and a slightly fringed aperture (both from Myanmar).</p></div>	https://treatment.plazi.org/id/03CC1E69A3B6FF54B946FDEFB7F8FD07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B6FF58B921FA65B7B1FDAA.text	03CC1E69A3B6FF58B921FA65B7B1FDAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax succineus (W. T. Blanford 1862)	<div><p>Dicharax succineus (W.T. Blanford, 1862)</p><p>Figs 77–80</p><p>Alycaeus succineus W.T. Blanford, 1862: 139–140 .</p><p>Alycaeus vulcani W.T. Blanford, 1863: 323 . Syn. nov.</p><p>Alycaeus blanfordi Godwin-Austen, 1914: 418, pl. 148 fig. 3. Syn. nov.</p><p>Alycaeus succineus – Sowerby 1877: pl. 2, species 16. — Godwin-Austen 1914: 424, pl. 151 fig. 2.</p><p>Alycaeus vulcani – Sowerby 1877: pl. 2, species 17. — Godwin-Austen 1914: 413–414, pl. 151 figs 5, 5a. — Gude 1921: 221–222.</p><p>Alycaeus (Dicharax) succineus – Kobelt 1902: 377. — Gude 1921: 271–272.</p><p>Alycaeus (Alycaeus) vulcani – Kobelt 1902: 352.</p><p>Alycaeus blanfordi – Gude 1921: 206.</p><p>Chamalycaeus vulcani – Páll-Gergely et al. 2020: 48.</p><p>Dicharax (?) succineus – Páll-Gergely et al. 2020: 106.</p><p>Dicharax (?) blanfordi – Páll-Gergely et al. 2020: 106.</p><p>Diagnosis</p><p>The large shell, widely-spaced ribs, the long R2, relatively low R3 swelling and the fringed aperture distinguish this species from its congeners. Based on the lamella-like ribs, this species rather resembles species of Chamalycaeus and Metalycaeus (e.g., M. brahma, see Páll-Gergely et al. 2020), but lacks the spiral striation on its entire shell.</p><p>Type material examined</p><p>MYANMAR • 1 syntype of A. succineus (Fig. 77A–E); Tangoop Pass, Arakan Hills; NHMUK 1906.4.4.52 • holotype of A. blanfordi (single specimen mentioned in the original description: Fig. 77F– J); Chwegalé, Arakan Hills; NHMUK 1906.4.4.177 • 1 shell, labelled as syntype of A. vulcani; Ava, Burma; MCZ 135705 • 4 shells, possible syntypes of A. vulcani; Puppadoung; ex coll. Theobald; NHMUK 1888.12.4.939-942 • 2 shells (possible syntypes of A. vulcani); Puppa Hill, Ava; Blanford leg.; Crosse coll. 1899, Sykes coll. 1954; NHMUK • possible syntypes of A. vulcani (Fig. 78A–E); Pupa Hills, Burma, extinct volcano; W.T. Blanford coll.; NHMUK 1906.4.4.49 • possible syntypes of A. vulcani (Fig. 78F–J); Puppa Ava, Burma; H.F. Blanford coll., ex coll. auctoris; NHMUK 20160325 .</p><p>Additional material examined</p><p>INDIA – Mizoram • 1 empty shell (Fig. 79); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 70; NZSI LM1727 • 2 empty shells; same data as for preceding; Ar26; NZSI LM1728 • 1 empty shell; same data as for preceding; Ar22; NZSI LM1729 .</p><p>Type localities</p><p>“in montibus Arakanensibus” ( A. succineus); “on the upper portion of the isolated peak of Puppa, an extinct volcano lying ca 40 miles E. S. E. of the town of Pu-gán in the territories of the king of Ava” ( A. vulcani); “Chwegali, Arakan Hills” [probably Chaingalain Village, Rati Taung: 20°23.8’N 92°39.9’E] ( A. blanfordi).</p><p>Distribution</p><p>This species is known from the western part of Myanmar and the Indian state of Mizoram (Fig. 80).</p><p>Remarks</p><p>The type specimens of Alycaeus blanfordi, A. succineus and A. vulcani are nearly identical. There is no notable difference in shell size, shape, and sculpture. Therefore, they are treated as synonyms, and the oldest name, D. succineus, is valid. Blanford (1863), when describing D. vulcani, compared that species with D. succineus, and wrote that it was more globose than A. succineus . The opposite is true; the syntype of A. succineus is slightly more globular than that of A. vulcani, although the difference represents intraspecific variability. Godwin-Austen (1914) did not compare A. blanfordi with any other species and only noted that the species had very distinctive characters and was worthy of being named.</p><p>The specimens from the Blue Mountain differ from the ones from the Arakan Mountains (types of A. blanfordi, A. succineus and A. vulcani) by having a glossy R3, while that of the specimens in the Arakan Mountains are matte with some growth lines. We consider this difference insufficient for species-level distinction.</p></div>	https://treatment.plazi.org/id/03CC1E69A3B6FF58B921FA65B7B1FDAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B9FF5BB947F9FAB445F972.text	03CC1E69A3B9FF5BB947F9FAB445F972.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax species group 12</p><p>Remarks</p><p>This species group is not based on morphology, but all species from Myanmar which could not be classified elsewhere are lumped here.</p></div>	https://treatment.plazi.org/id/03CC1E69A3B9FF5BB947F9FAB445F972	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B9FF5AB93AF917B71FFCEF.text	03CC1E69A3B9FF5AB93AF917B71FFCEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax ataranensis (Godwin-Austen 1914)	<div><p>Dicharax ataranensis (Godwin-Austen, 1914)</p><p>Fig. 81A–J</p><p>Alycaeus ataranensis Godwin-Austen, 1914: 426, pl. 148 figs 4, 4a–b.</p><p>Alycaeus (Dicharax) ataranensis – Gude 1921: 237.</p><p>Dicharax ataranensis – Páll-Gergely et al. 2020: 77, fig. 18; 2021: 25, fig. 18.</p><p>Diagnosis</p><p>R1 nearly smooth (with only some strong, widely-spaced ribs near the suture), R3 swelling conical, upper edge of peristome strongly protruding. Based on these traits, this species can be easily distinguished from its congeners.</p><p>Type material examined</p><p>MYANMAR • holotype (single shell mentioned in the original description: Fig. 81A–E); Ataran, Burma; ex Dr F. Stoliczka coll.; NZSI M.8073 .</p><p>Additional material examined</p><p>MYANMAR • 1 shell (Fig. 81F–J); Kayin State, ca 51 km ESE of Mudon Centre, NE of Htimahto, NE point of <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=97.97&amp;materialsCitation.latitude=16.039934" title="Search Plazi for locations around (long 97.97/lat 16.039934)">Kwooprai Hill</a>; 16°2.396′ N, 97°58.200′ E; 30 m a.s.l.; 10 Oct. 2018; A. Hunyadi, K. Okubo and J.U. Otani leg.; Coll. HA .</p><p>Type locality</p><p>“Ataran”.</p></div>	https://treatment.plazi.org/id/03CC1E69A3B9FF5AB93AF917B71FFCEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5AB926FC9CB281FA4D.text	03CC1E69A3B8FF5AB926FC9CB281FA4D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax dohertyi (Godwin-Austen 1893)	<div><p>Dicharax dohertyi (Godwin-Austen, 1893)</p><p>Fig. 81K–L</p><p>Alycaeus dohertyi Godwin-Austen, 1893: 595 .</p><p>Alycaeus dohertyi – Godwin-Austen 1897: 3, pl. 63 figs 3, 3a. — Godwin-Austen 1914: 408.</p><p>Alycaeus (Dicharax) dohertyi – Kobelt 1902: 369. — Gude 1921: 248–249.</p><p>Dicharax (?) dohertyi – Páll-Gergely et al. 2020: 85.</p><p>Material examined</p><p>The type specimens of this species are probably lost (Páll-Gergely et al. 2020).</p><p>Type locality</p><p>“Momeit, Burmah”.</p><p>Differential diagnosis</p><p>According to the drawings (Fig. 81K–L), the most similar species is probably D. rugosus, but in that species only the inner surface of the peristome is fringed. See also under D. muspratti .</p></div>	https://treatment.plazi.org/id/03CC1E69A3B8FF5AB926FC9CB281FA4D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3B8FF5CB925F9FDB7FAFBD2.text	03CC1E69A3B8FF5CB925F9FDB7FAFBD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax stoliczkii (Godwin-Austen 1874)	<div><p>Dicharax stoliczkii (Godwin-Austen, 1874)</p><p>Fig. 82</p><p>Alycaeus Stoliczkii Godwin-Austen, 1874: 147, pl. 3 fig. 3.</p><p>Alycaeus (Charax) peilei Preston, 1914: 22–23, figure on page 23. Syn. nov.</p><p>Alycaeus stoliczkii – Sowerby 1877: pl. 6, species 53. — Godwin-Austen 1914: 399–400, pl. 144 figs 3, 3a–b.</p><p>Alycaeus (Chamalycaeus) stoliczkai [sic] – Kobelt 1902: 363. — Gude 1921: 233–234.</p><p>Alycaeus peilei – Godwin-Austen 1914: 397–398.</p><p>Alycaeus (Dicharax) peilei – Gude 1921: 264.</p><p>Chamalycaeus (Chamalycaeus) stoliczkai [sic] – Ramakrishna et al. 2010: 55.</p><p>Chamalycaeus (Dicharax) peilei – Ramakrishna et al. 2010: 65.</p><p>Dicharax (?) peilei – Páll-Gergely et al. 2020: 101.</p><p>Dicharax (?) stoliczkii – Páll-Gergely et al. 2020: 104.</p><p>Diagnosis</p><p>The large shell size, the long R2+R3, and the aperture shape distinguish this species from all other Alycaeinae .</p><p>Type material examined</p><p>INDIA • 3 syntypes of A. stoliczkii (Fig. 82A–E); Naga Hills; NHMUK 1903.7.1.2622 • 1 syntype of A. peilei (Fig. 82F–J); Naga Hills; Preston coll.; NHMUK 1915.1.4.1281 .</p><p>Type localities</p><p>“Angaoluo Peak, Nágá Hills at 7,000 feet ”; “further to the east at Kezakenomih, and at the head of the Lanier River at ca. 5,000 feet where the specimens are much larger” ( A. stoliczkii); “ Naga Hills” ( A. peilei).</p><p>Differential diagnosis</p><p>The most similar species is D. glaber, but D. stoliczkii has a more oval shell (rather rounded in D. glaber), a fringed aperture and a secondary swelling touching the penultimate whorl near the peristome.</p><p>Remarks</p><p>Alycaeus (Charax) peilei is identical with Alycaeus stoliczkii and therefore it is treated as a junior synonym. In the original description of the first species, Preston (1914) did not compare his new species with any other species.</p></div>	https://treatment.plazi.org/id/03CC1E69A3B8FF5CB925F9FDB7FAFBD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3BEFF5CB969FB70B760F9B4.text	03CC1E69A3BEFF5CB969FB70B760F9B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metalycaeus Pilsbry 1900	<div><p>Genus Metalycaeus Pilsbry, 1900</p><p>Alycaeus (Metalycaeus) Pilsbry, 1900: 382 (section of Alycaeus).</p><p>Metalycaeus – Páll-Gergely &amp; Asami 2017: 4. — Páll-Gergely et al. 2017: 73; 2020: 137 (treated Raptomphalus Godwin-Austen, 1914 as a junior synonym).</p><p>Type species</p><p>Alycaeus (Metalycaeus) melanopoma Pilsbry, 1900 (Páll-Gergely et al. 2017; fig. 31a) (junior synonym of Metalycaeus nipponensis (Reinhardt, 1877), see Minato 1988), subsequent designation by Thiele (1929: 108).</p></div>	https://treatment.plazi.org/id/03CC1E69A3BEFF5CB969FB70B760F9B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
03CC1E69A3BEFF44BAC6F9D5B22DFB82.text	03CC1E69A3BEFF44BAC6F9D5B22DFB82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metalycaeus polygonoma (W. T. Blanford 1862)	<div><p>Metalycaeus polygonoma (W.T. Blanford, 1862)</p><p>Figs 83–88</p><p>Alycaeus polygonoma W.T. Blanford, 1862: 140–141 .</p><p>Alycaeus distinctus Godwin-Austen, 1893: 592 . Syn. nov.</p><p>Alycaeus rotundatus Godwin-Austen, 1914: 359, pl. 154 fig. 6. Syn. nov.</p><p>Alycaeus kamakiaensis Godwin-Austen, 1914: 375–376, pl. 141 fig. 8. Syn. nov.</p><p>Metalycaeus posteriobulbus Gittenberger &amp; Sherub Sherub in Gittenberger et al., 2024: 193, figs 1, 6–7. Syn. nov.</p><p>Alycaeus ingrami var. – Godwin-Austen 1871: 91–92, pl. 4 fig. 3.</p><p>Alycaeus polygonus [sic] – Sowerby 1877: pl. 2, species 11.</p><p>Alycaeus (Dicharax) polygonoma – Kobelt 1902: 375. — Gude 1921: 265–266. Alycaeus polygonoma – Godwin-Austen 1914: 423, pl. 141 fig. 5.</p><p>Alycaeus distinctus – Godwin-Austen 1914: 363; 1914: 390, pl. 145 figs 3, 3a–b.</p><p>Alycaeus distinctus var. – Godwin-Austen 1914: 363, pl. 137 figs 2, 2a–b, 391, pl. 149 fig. 4 (2 varieties). Alycaeus kamakiaensis – Gude 1921: 209.</p><p>Alycaeus rotundatus – Gude 1921: 217.</p><p>Alycaeus (Chamalycaeus) distinctus – Gude 1921: 226–227.</p><p>Alycaeus (Alycaeus) kamakiaensis – Ramakrishna et al. 2010: 48.</p><p>Alycaeus (Alycaeus) rotundatus – Ramakrishna et al. 2010: 51.</p><p>Chamalycaeus (Chamalycaeus) distinctus – Ramakrishna et al. 2010: 53.</p><p>Metalycaeus distinctus – Páll-Gergely et al. 2020: 142.</p><p>Metalycaeus kamakiaensis – Páll-Gergely et al. 2020: 145.</p><p>Metalycaeus polygonoma – Páll-Gergely et al. 2020: 151.</p><p>Metalycaeus rotundatus – Páll-Gergely et al. 2020: 153.</p><p>Diagnosis</p><p>This species can be recognised based on the relatively large aperture, finely undulated peristome, the elongated basal peristome edge and the blunt centrally located R3 swelling. We indicate that the</p><p>Himalayan species of Metalycaeus are not revised in this study, and a comprehensive revision should identify the species-specific traits.</p><p>Type material examined</p><p>INDIA • 10 syntypes of A. distinctus (Fig. 83F–J); Naga Hills; NHMUK 1903.7.1.2619 • 1 syntype of A. kamakiaensis (Fig. 84A–E); Kamakia Hill, Gowhatty, Assam; NHMUK 1903.7.1.2705 • holotype (single shell mentioned in the original description: Fig. 83F–J) of A. rotundatus; Dafla Hills (?); NHMUK 1903.7.1.2543 .</p><p>MYANMAR • 2 syntypes of A. polygonoma in different vials (Fig. 83A–E); Tongoop Pass, Arakan Hills; NHMUK 1906.4.4.51 .</p><p>Additional material examined</p><p>BANGLADESH • 2 shells (Fig. 88); Chittagong, natural mango garden on a river bank; 22°31.559′ N, 92°7.652′ E; 27 Sep. 2024; T. Saito and M.S. Shariar leg.; Coll. PGB .</p><p>INDIA • 2 shells of “ Alycaeus distinctus var.”; Sadia, Assam; NHMUK 1903.7.1.2620 • 19 + 6 shells of “ Alycaeus distinctus var.”; Jatinga Valley, N. Cachar; Godwin-Austen coll.; NHMUK 1903.7.1.2576 • 1 empty shell (Figs 12D, 85); Blue Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.135&amp;materialsCitation.latitude=22.727" title="Search Plazi for locations around (long 93.135/lat 22.727)">Lawngtlai district</a>; 22.727° N, 93.135° E; 1400 m a.s.l.; 27 Jan. 2019; N.A. Aravind leg.; Ar 48; NZSI LM1762 • 2 empty shells (Figs 86–87 [operculum]); same data as for preceding; Ar 71; NZSI LM1763 • 1 empty shell; same data as for preceding; Ar 43; NZSI</p><p>LM1909 • 5 empty shells; same data as for preceding; Ar46; NZSI LM1910 • 2 empty shells; same data as for preceding; Ar51; NZSI LM1911.</p><p>Type localities</p><p>“in montibus Arakanensibus” ( A. polygonoma); “Neighbourhood of Asálú, N. Cachar Hills” ( A. distinctus); “Kamakia Temple Hill near Gowhatty, Assam ” ( A. kamakiaensis); “Dafla hills” ( A. rotundatus); “ Zhemgang Dzongkhag: between Duenmang Tsachu and Gongphu Zero Point, 24 km SE of Zhemgang, 335 m a.s.l., 27°02’N 90°48’E, scree in warm broadleaf forest” ( M. posteriobulbus).</p><p>Distribution</p><p>The new information reveals that M. polygonoma has a larger distribution than previously thought; its area spans from Bhutan and the Dafla Hills (India) to the Arakan Mountains in Myanmar (Fig. 89). Furthermore, this species is reported from Bangladesh for the first time. With these localities, M. polygonoma is one of the few species of Alycaeinae with a large distributional area besides Metalycaeus heudei (Páll-Gergely et al. 2017, 2021), Dicharax fimbriatus and D. cristatus (Möllendorff, 1886) (see Páll-Gergely et al. 2017).</p><p>Remarks</p><p>Type specimens of A. polygonoma, A. distinctus, A. kamakiaensis, A. rotundatus and M. posteriobulbus do not differ in any important conchological characters; therefore, the latter four are junior synonyms of the former. The types of those five species and the newly collected samples from the Blue Mountain show slight variability in shell size, length of R2, width of umbilicus, and the degree of undulation of the peristome, but all differences are attributed to intraspecific variability.</p></div>	https://treatment.plazi.org/id/03CC1E69A3BEFF44BAC6F9D5B22DFB82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Aravind, Neelavar Ananthram	Páll-Gergely, Barna, Gojšina, Vukašin, Aravind, Neelavar Ananthram (2025): A review of Chamalycaeus, Cycloryx and Dicharax species of the Himalaya and Myanmar and seven new species of Alycaeinae from the Blue Mountain, Mizoram, India (Gastropoda, Caenogastropoda, Cyclophoridae). European Journal of Taxonomy 1029: 1-158, DOI: 10.5852/ejt.2025.1029.3131, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3131/13935
