taxonID	type	description	language	source
03CF1E339367FF93FF73CD665BBE9BE3.taxon	description	(Fig. 3 C, 4 C) Valve elongated in lateral view, not trapezoidal, with rounded tip; there are sclerotized teeth on the dorsal part of the valve near the tip. Tegumen massive. Uncus not massive. Subuncii broad and not pointed; they have approximately the same thickness in the entire length. Juxta small, not strongly sclerotized. Aedeagus long. Vesica entirely different from Davidina and Oeneis (Protoeneis) with two tooth-like, sharply terminated cornuti. Neominois ridingsii (type species of Neominois) (Fig. 3 D, 4 D) Valve with a specific hump-like convexity on costa; without sclerotized teeth. Tegumen massive. Uncus not massive. Subuncii thin and pointed, they gradually thin to the top. Aedeagus long. Vesica simple, membranous, without sclerotized teeth.	en	Lukhtanov, Vladimir A, Dubatolov, Vladimir V (2020): Phylogenetic position and taxonomic rearrangement of Davidina (Lepidoptera: Nymphalidae), an enigmatic butterfly genus new for Europe and America. Zoological Journal of the Linnean Society 190 (3): 1036-1053, DOI: 10.1093/zoolinnean/zlaa104, URL: https://academic.oup.com/zoolinnean/article/190/3/1036/5907879
03CF1E339367FF9DFCCBCC8F5E5C995B.taxon	description	Karanasa kirgisorum Avinov & Sweadner (Figs 3 F, 4 F) Valve elongated in lateral view, not trapezoidal, with rounded tip; there are sclerotized teeth on the dorsal part of valve near the tip. Tegumen massive. Subuncii have approximately the same thickness in the entire length. Juxta massive and strongly sclerotized. Aedeagus relatively short, curved and armed with spikes. Vesica with two cornuti, each possessing a single sclerotized tooth. Generally, the male genitalia in Karanasa are similar to those in Oeneis (Oeneis), except for the massive and strongly sclerotized juxta and the aedeagus, which is curved and armed with spikes. Thus, the following characters of the male genitalia are unique for one or two of the studied genera and subgenera and represent potential (syn) apomorphies. (1) Trapezoidal, nearly triangular shape of valve in lateral view (character 1) and vesica with two large, oval sclerotized cornuti, each cornutus with several claw-like teeth forming a strongly sclerotized ridge (character 2) (unique characters for Davidina and Oeneis (Protoeneis). (2) Valve distinctly narrowed in distal portion, not trapezoidal, with rounded tip (character 3) [unique character for Oeneis (Oeneis) and Karanasa]. (3) Specific shape of valve with a hump-like convexity on costa (character 4). This character is found in Neominois ridingsii (de Lesse, 1951; Warren et al., 2008; this study) and seems to be unique for Neominois. However, it is not genus-specific, since it is not found in the second species of Neominois, N. carmen Warren, Austin et al., 2008 (Warren et al., 2008). (3) Bilobate valve (character 5), found by us for Paroeneis and previously reported for Aulocera (de Lesse, 1951; Chou, 1998; Sharma & Rose, 2014). (4) Massive and strongly sclerotized juxta (character 6) and aedeagus armed with spikes (character 7) (unique character for Karanasa). PHYLOGENETIC ANALYSIS OF THE MORPHOLOGICAL CHARACTERS 8. Valve with the distal process (JakŠić, 1998: fig. 61, 4 – 9, fig. 62, 1 – 5; Nekrutenko, 1985: fig, 59, 64) (1); valve without the distal process (0). 9. The distal part of the valve pointed and curved up (JakŠić, 1998: fig. 61, 1; figs 68 – 70; Nekrutenko, 1985: fig, 58, 65, 66) (1); the distal part of the valve not pointed and not curved up (0). 10. The distal process of the valve long (JakŠić, 1998: fig. 61, 4 – 9; Nekrutenko, 1985: fig, 59) (1); the distal process of the valve short and massive (JakŠić, 1998: fig. 62, 1 – 5; Nekrutenko, 1985: fig. 64) (0). 11. Valve broad, with distal extension (JakŠić, 1998: figs 63 – 66) (1); valve without distal extension (0). 12. Jullien organ present (JakŠić, 1998: figs 63 – 66) (1); Jullien organ absent (0). Maximum parsimony phylogenetic analysis (MP) of the morphological matrix reveals six clades that appear in all nine MP trees. The 100 % consensus of the nine MP trees is shown in Figure 5. The same topology is revealed by BI analysis of the morphological matrix. Three clades, (Davidina + Protoeneis), (Minois + Arethusana) and [Brintesia + (Minois + Arethusana)] were highly supported in the BI analysis and had a Based on the original data on characters 1 – 7 and published data on characters 4, 8 – 12 (de Lesse, 1951; Nekrutenko, 1985, JakŠić, 1998; Warren et al., 2008), we created the following matrix of the binary characters (Table 1): 1. Valve trapezoidal, nearly triangular shape in lateral view (1); valve elongated, not trapezoidal or triangular in lateral view (0). 2. Vesica with two large oval sclerotized cornuti, each cornutus with several claw-like teeth forming a strongly sclerotized ridge (1); vesica simple, membranous, without sclerotized teeth or with two single teeth (0). 3. Valve with rounded tip covered by numerous sclerotized teeth (1); tip of the valve not rounded (0). 4. Valve with a large hump-like convexity on costa (1); valve without a large hump-like convexity on costa (0). 5. Valve bilobate (1); valve not bilobate (0). 6. Juxta large, massive and strongly sclerotized (1); juxta small, not strongly sclerotized (0). 7. Aedeagus armed with spikes (1); aedeagus not armed with spikes (0). medium support in our MP analysis. Three clades, (Aulocera + Paroeneis), (Oeneis + Karanasa) and (Satyrus + Pseudochazara + Chazara), had low support. WING VENATION According to Kuznezov (1930), there is a clear distinction between Davidina (based on D. armandi, the type species of the genus) and Oeneis (based on O. norna, the type species of the genus) in the forewing venation. First, in Davidina all radial branches, except R 1, form a common and rather long stalk R 2 + 3 + 4 + 5; in Oeneis r 2 is free, i. e. R 2 and R 3 + 4 + 5 start independently from the same point of the discal cell. Second, in Davidina M 1 rises at a distance from this stalk, the vein r-m 1 being, thus, well developed; in Oeneis M 1 rises from one point with R 3 + 4 + 5, and r-m 1 is, thus, absent. Kuznezov’s observation was based on two specimens only. We analysed 18 specimens of D. armandi and discovered that four types of venation are present in Davidina. 1. All radial branches, except R 1, form a common and rather long stalk R 2 + 3 + 4 + 5; M 1 rises at a distance from this stalk, the vein r-m 1 being, thus, well developed [Davidina type according to Kuznezov (1930), found in 11 samples, Fig. 6 A]. 2. Veins R 2, R 3 + 4 + 5 and M 1 rise from the same point on the discal cell; the vein r-m 1 is absent [Oeneis type according to Kuznezov (1930), found in four samples, Fig. 6 B]. 3. Veins R 2 and R 3 + 4 + 5 rise from the same point on discal cell; M 1 rises at a distance from this point, the vein r-m 1 being, thus, well developed (found in two samples, Fig. 6 C); 4. All radial branches, except R 1, form a common and rather long stalk R 2 + 3 + 4 + 5; this stalk and M 1 rise from the same point on the discal cell; the vein r-m 1 is absent (found in a single sample, not shown). Thus, the wing venation is not stable in D. armandi. It is not genus-specific, but variable, even on population level. The supposed differences between Oeneis and Davidina in wing venation (Kuznezov, 1930) is not confirmed.	en	Lukhtanov, Vladimir A, Dubatolov, Vladimir V (2020): Phylogenetic position and taxonomic rearrangement of Davidina (Lepidoptera: Nymphalidae), an enigmatic butterfly genus new for Europe and America. Zoological Journal of the Linnean Society 190 (3): 1036-1053, DOI: 10.1093/zoolinnean/zlaa104, URL: https://academic.oup.com/zoolinnean/article/190/3/1036/5907879
03CF1E33936EFF85FEEECD0E5B139D35.taxon	description	= Leechia Röber, 1907. In Seitz, Grossschmetterlinge der Erde 1: 43, t. 19 b. Type-species: Leechia alticola Röber, 1907 (In Seitz, Grossschmetterlinge der Erde 1: 43, t. 19 b). = Sinosatyrus Lee Chuan-Lung, 1988. Chinese Science Bull. 33 (4): 320. Type-species: Sinosatyrus beyingensis Lee, 1988. = Protoeneis P. Gorbunov, 2001. In Gorbunov, The butterflies of Russia: classification, genitalia, keys for identification (Lepidoptera: Hesperioidea and Papilionoidea): 228. Type-species: Chionobas nanna Ménétriés, 1858 (Bull. Class. phys. - math. Acad. imp. Sci. St. Petersb. 17 (12 / 14): 216). 1. Davidina armandi Oberthür, 1879. Et. Entom. 4: 19, t. 2, fig. 1. TL: Pe-hoa-tschan, China. Holotype in BMNH (Fig. 1). = alticola (Röber, 1907) (Leechia alticola). In Seitz, Grossschmetterlinge der Erde 1: 43, t. 19 b. TL: ‘ Central-China (Tschang-yang) ’. TM in BMNH. = beyingensis (Lee Chuan-Lung, 1988) (Sinosatyrus beyingensis). Chinese Science Bull. 33 (4): 320. Distribution: N. E. China. Note. Lee (1988) spelled his new species of Sinosatyrus in two different ways: beÿingensis and beÿiengensis (both on p. 320, note the diaeresis mark placed over the ‘ y’). Both spellings appear to be misprints of the letters ‘ ij ’, thus possibly Lee’s original intention was to name the species beijingensis but that was not what was printed. Here, acting as First Reviser, we select beyingensis as the correct original spelling, simultaneously deleting the diacritic mark, following Arts. 32.2.1 and 32.5.2.1 of the ICZN. 2. Davidina diluta (Lukhtanov, 1994) (Oeneis diluta). In Lukhtanov & Lukhtanov, Die Tagfalter Nordwestasiens, Herbipoliana 3: 138, t. 25, fig. 6. TL: Susch, Ujukski-Gebirge, Tuva, S. Siberia, Russia. HT in ZISP. Distribution: Russia: C. Tuva. 3. Davidina lederi (Alpheraky, 1897) (Oeneis Tarpeia Pall. var. Lederi). Mém. Lép. Rom. 9: 196. TL: ‘ Urga’ [Ulanbator], Mongolia (probably in the Khangai Mts). LT in ZISP, designated by U. Eitschberger & V. A. Lukhtanov (1994), Atalanta 25: 163. = sapozhnikovi Korshunov, 1982 (Oeneis sapozhnikovi). Gel’minty, kleshchi i nasekomye: 86, figs 1 – 2. TL – 10 km NW Mungen-Mort, Central Aimak, Mongolia. HT in SZM. Distribution: N. and C. Mongolia. Russia: Irkut River valley in S. Siberia, S. Tuva. 4. Davidina mongolica (Oberthür, 1876) 4 a. Davidina mongolica mongolica (Oberthür, 1876) (Chionobas mongolica). Et. Entom. 2: 31, t. 4, fig. 6. TL: ‘ Mongolie orientale. Elle vole en été dans les montagnes, à une altitude moyenne de 500 mètres’ TM in BMNH. = tsingtaua Austaut, 1911 (Oeneis mongolica tsingtaua). Ent. Zeitschr. 24: 244. TL: Tsingtau, Shantung, China or. TM -?. = tsingtauana auct. (misspelling). = mandschurica O. Bang-Haas, 1939 (Oeneis mongolica mandschurica). Iris 53: 49, t. 1, fig. 6. TL: Kintschou, Prov. Fengtien, Manchuria mer. or., China. TM in ZMHU. f. extincta O. Bang-Haas, 1939. Iris 53: 49. TL: Kintschou, Prov. Fengtien, Manchuria mer. or., China. TM in ZMHU. Distribution: N. E. China: Inner Mongolia, Liaoning. 4 b. Davidina mongolica hoenei (Gross, 1970) (Oeneis mongolica hoenei). Mitt. Münch. Ent. Ges. 58: 22. TL: ‘ Mienshan, Shansi, Nordchina’. HT in MAKB. Distribution: N. China: Shansi [Shanxi]. 4 c. Davidina mongolica coreana (Matsumura, 1927) (Oeneis nanna coreana). Ins. Mats. 1: 163, t. 5, fig. 9. TL: ‘ Genzan’, Korea. TM in HUS (?). = okamotonis Matsumura, 1927 (‘ Oeneis nanna coreana okamotonis n. ab. ’, p. 164; ‘ Oeneis nanna okamotonis ’, p. 160). Ins. Mats. 1: 164, t. 5, f. 3. TL: ‘ Suigun’, Korea. TM in HUS (?). f. fumosa O. Bang-Haas, 1939. Iris 53: 49, t. 1, f. 6. TL: ‘ Corea int., bei Gensan’. TM in ZMHU. Distribution: N. Korea. 4 d. Davidina mongolica walkyria (Fixsen, 1887) (Oeneis walkyria). Mém. Lép. Rom. 3: 310. t. 14, fig. 4. TL: ‘ in der Umgebung Pung-Tungs’, Korea. TM in ZISP. = shonis Matsumura, 1927 (‘ Oeneis nanna walkyria shonis n. ab. ’, p. 163; ‘ Oeneis nanna shonis ’, p. 150). Ins. Mats. 1: 163, t. 5, fig. 2. TL: ‘ Mt. Daitoku’, Korea. TM in HUS (?). f. hakuba Doi, 1934 (Oeneis nanna walkyria f. hakuba). In Mori, Doi & Cho, Coloured butterflies from Korea (2): 16, t. 12, fig. 4. TL: Korea. TM -?. f. soibona Doi, 1934 (Oeneis nanna walkyria f. soibona). In Mori, Doi & Cho, Coloured butterflies from Korea (2): 16, t. 12, fig. 7. TL: Korea. TM -?.	en	Lukhtanov, Vladimir A, Dubatolov, Vladimir V (2020): Phylogenetic position and taxonomic rearrangement of Davidina (Lepidoptera: Nymphalidae), an enigmatic butterfly genus new for Europe and America. Zoological Journal of the Linnean Society 190 (3): 1036-1053, DOI: 10.1093/zoolinnean/zlaa104, URL: https://academic.oup.com/zoolinnean/article/190/3/1036/5907879
03CF1E33936EFF85FEEECD0E5B139D35.taxon	distribution	Distribution: C. Korea. 4 e. Davidina mongolica hallasanensis (Murayama, 1991) (Oeneis urda hallasanensis). Nature and Insects 26 (3): 20. TL: Mt. Hallasan, Cheju-do Island, S. Korea. TM -?. Distribution: Korea: Cheju-do Island. 5. Davidina nanna (Ménétriés, 1858)	en	Lukhtanov, Vladimir A, Dubatolov, Vladimir V (2020): Phylogenetic position and taxonomic rearrangement of Davidina (Lepidoptera: Nymphalidae), an enigmatic butterfly genus new for Europe and America. Zoological Journal of the Linnean Society 190 (3): 1036-1053, DOI: 10.1093/zoolinnean/zlaa104, URL: https://academic.oup.com/zoolinnean/article/190/3/1036/5907879
03CF1E33936EFF85FEEECD0E5B139D35.taxon	description	5 a. Davidina nanna dzhulukuli (Korshunov, 1998) (Oeneis anna dzhulukuli). In Korshunov, Novye opisaniya i dopolneniya dlya knigi ‘ Dnevnye babochki Aziatskoi chasti Rossii’: 27, t. 17 (4). TL: Taboshak Mt., Kuraiski Chrebet, Altai, Russia. TM in SZM. Distribution: Russia: S. E. Altai (Kurai, S. Tchuja and Sailjugem Mountains, Ukok-Plateau), W. Tuva. 5 b. Davidina nanna anna (Austaut, 1911) (Oeneis nanna var. anna). Ent. Zeitschr. 24: 243. TL: ‘ Arasagun-Gol’ (N. Mongolia). TM in?. = brunhilda A. Bang-Haas, 1912 (Oeneis brunhilda). Iris 26: 105, t. 6, f. 1. TL: ‘ aus dem Sajan-Gebiete’. Acording to the syntype labels the type-locality is ‘ Sajan, Arasagun-Gol’ (N. Mongolia) and ‘ Sajan, Arsyn’ (Russia, S. Tuva). STs in ZMHU. = burjatica Korshunov et Nikolaev, 2002 (Oeneis anna burjatica). In Korshunov, 2002, Bulavousye Cheshuekrylaye Severnoi Azii (Butts N Asia): 319. D i s t r i b u t i o n: R u s s i a: S. T u v a, S. B u r y a t i a. N. Mongolia. 5 c. Davidina nanna nanna (Ménétriés, 1858) (Chionobas nanna). In Bull. Class. phys. - math. Acad. imp. Sci. St. Petersb. 17 (12 / 14): 216. TL: ‘ Amour’ [Amur], Russia. HT in ZISP. = hulda Staudinger, 1887 (Oeneis hulda). Mém. Lép. Rom. 3: 149, t. 16, f. 8. TL: Pokroffka, Amur, Russia. LT in ZMHU, designated by F. J. Gross (1970), Mitt. Münch. Ent. Ges. 58: 20, Abb. 29. = coriacea Seitz, 1909 (Oeneis nanna coriacea). In Seitz, Grossschmetterlinge der Erde 1: 121, t. 40 g. TL: ‘ Apfelgebirge’, S. Siberia, Russia. TM in SMF. Distribution: Russia: Transbaikalia, Amur basin. N. E. Mongolia. N. E. China. 5 d. Davidina nanna jakutski (Korshunov, 1998) (Oeneis jakutski). Novye opisaniya i dopolneniya dlya knigi ‘ Dnevnye babochki Aziatskoi chasti Rossii’: 27, t. 17 (3). TL: Yakutsk, botsad [botanical garden], Yakutia, Russia. TM in SZM. Distribution: Russia: Yakutia. 5 e. Davidina nanna dzhugdzhuri (Sheljuzhko, 1929) (Oeneis dzhugdzhuri). Mitt. Münch. Ent. Ges. 19: 349, t. 28, figs. 3 – 4. TL: ‘ in der Bergkette Dzgugdzhur (Grenze der Provinzen Amur und Jakutsk), an den Quellen des Flusses Dzhelinda’ [Yakutia], Russia. HT in coll. Sheljuzhko (Kiev University, Ukraine). Distribution: Russia: Southern and Eastern Yakutia, northern part of Amurskaya oblast’. 5 f. Davidina nanna taimyrica (Lukhtanov & Eitschberger, 2001). (Oeneis nanna taimyrica). In: Lukhtanov & Eitschberger (2001) Catalogue of the genera Oeneis and Davidina (Bauer E, Frankenbach T, eds) Butterflies of the World, Supplement 4: 33. TL: Russia, Taimyr Peninsula, Putorana plateau, Ayan Lake, near Ayan River. HT in Zoological Museum of Moscow University. Distribution: Russia: Northern Siberia (Taimyr Peninsula). 6. Davidina sculda (Eversmann, 1851) 6 a. Davidina sculda sculda (Eversmann, 1851) [Hipparchia (Chionobas) sculda]. Bull. Soc. Nat. Mosc. 24: 612. TL: ‘ environs de Kiachta de la Sibérie orientale’ (Buryatia, S. Siberia, Russia). STs in ZISP. = sculda Keferstein, 1851, Stett. ent. Ztg. 12 (9): 274 (a nomen nudum) (Chionabas [sic]) = velleda Austaut, 1911 (Oeneis velleda). Int. Ent. Zeitschr. 5: 360. TL: Siberia. TM in?. Distribution: Russia: Altai, Tuva, Sayan Mts, S. Buryatia. N. E. Mongolia. 6 b. Davidina sculda pseudosculda (Korshunov, 1977) (Oeneis pseudosculda). Insects of Mongolia 5: 662, fig, 1. TL: Sudzykte, Central Aimak, Mongolia. HT in ZISP. Distribution: C. Mongolia. 6 c. Davidina sculda pumila (Staudinger, 1892) (Oeneis Sculda Ev. var. Pumila Stgr.). Mém. Lép. Rom. 6: 201. TL: ‘ Pokr. [offka] ’ and ‘ am oberen Amur’ (Transbaikalia region, Russia). STs in ZMHU. Distribution: Russia: Transbaikal (Tchita region), Amur region. N. E. Mongolia. N. E. China. 6 d. Davidina sculda vadimi (Korshunov, 1995) (Oeneis sculda vadimi). In Korshunov & Gorbunov, Dnevnye babochki Asiatskoi chasti Rossii: 138. TL: Severobaikalsk, Verkhneangarski Khrebet, N. Transbaikal, Russia. HT in SZM. Distribution: Russia: N. Transbaikal, Yakutia. 7. Davidina tarpeia (Pallas, 1771) 7 a. Davidina tarpeia tarpeia (Pallas, 1771) (Papilio tarpeia). Reise versch. Prov. russ. Reiches 1: 171 (tarpeja), 480 (tarpeia). TL: ‘ In campis aridis ad Volgam copio fus Maio’, Volga region, Russia. TM lost. = celimene (Cramer, 1782) (Papilio celimene). Pap. Exot. 4: 169, t. 375 E, F. TL: ‘ Siberie’. = vacuna Grum-Grshimailo, [1891] (Oeneis Vacuna). Horae Soc. Ent. Ross. 25: 458. TL: ‘ in montibus ad Dongar-tschen’, S. of Xining, Qinghai, China. HT in BMNH. = rozhdestvenskyi Korb et Yakovlev, 1997 (Oeneis tarpeia rozhdestvenskyi). Zoosystematica Rossica 5 (2): 313. TL: Shtabka, Altai, Russia. HT in ZISP. = ukokana Korb, 1998 (Oeneis tarpeia ukokana) Alexanor 20: 144, figs 1 – 3. TL: ‘ Südaltai, Ukok, Dschasator’, Russia. HT in ZISP. = baueri Lukhtanov & Eitschberger, 2000 (Oeneis tarpeia baueri). Schmetterlinge der Erde 11: 8. TL: Mezen’ r. [iver], vic. Sush’, Tuva, Siberia, Russia, 1100 – 1200 m. HT in McGuire Center for Lepidoptera and Biodiversity, University of Florida. Distribution: Russia: E. and S. E. Europe, N. Caucasus (Kislovodsk, Teberda, Elbrus), S. Ural, S. W. Siberia (W. Siberian Lowland), Altai, Tuva. N. and E. Kazakhstan. W. Mongolia. NW. China. Note. There are two different spellings of this name in the original description and in subsequent works. Here we select tarpeia as the most common spelling in historical and contemporary works. 7 b. Davidina tarpeia grossi (Eitschberger & Lukhtanov, 1994) (Oeneis tarpeia grossi). Atalanta 25: 164, t. Va, figs 1 – 3. TL: ‘ 20 km NW Selenduma’, Buryatia, S. Siberia, Russia. HT in ZISP. Distribution: Russia: Buryatia, Chita region. N. Mongolia. 8. Davidina uhleri (Reakirt, 1866) 8 a. Davidina uhleri varuna (W. H. Edwards, 1882) (Chionobas varuna). Canadian Ent. 14: 205. TL: ‘ plains of Dacotah Terr. ’, Canada. LT in CM. ab. dennisi Gander, 1927. Canadian Ent. 59: 285 – 286 (1927). TL: Beulah, Manitoba. HT in CNC. Distribution: Canada: British Columbia, Alberta, Saskatchewan, Manitoba. USA: Montana, Dakotas, Nebraska, W. Minnesota. 8 b. Davidina uhleri nahanni (Dyar, 1904) (Oeneis nahanni). Proc. Ent. Soc. Washington 6: 142. TL: Nahanni Mts., Dist. of Mackenzie, Canada. HT in USNM. Distribution: Canada: North-West Territories (Mackenzie Mts). 8 c. Davidina uhleri cairnesi (Gibson, 1920) (Oeneis cairnesi). Rept. Canadian Arctic Exped. 3 (1): 15. TL: White River dist., Yukon Territory, Canada. HT in CNC. = kluanensis (Hassler, 2002) (Oeneis nanna kluanensis). Nachricht. Entomol. Ver. Apollo 22: 197 – 205. TL: SW-Yukon, NW Burwash Landing, Quill Creek Area. HT in SMF. Distribution: Canada: Yukon, northwestern corner of the North-West Territories. USA: N. E. Alaska. 8 d. Davidina uhleri uhleri (Reakirt, 1866) (Chionobas Uhleri). Proc. Ent. Soc. Philadelphia 6: 143. TL: ‘ Rocky Mts., Colorado Territory’, USA, restricted to vic. Georgetown, Clear Creek Co., Colorado by F. M. Brown (1953), American Mus. Novitates (1625): 6. TM in FMNH. = obscura (W. H. Edwards, 1892) (Chionobas Uhleri var. Obscura). Butt. N. America 3: 294. TL: Clear Creek Canyon, Jefferson Co., Colorado. HT in CM. Distribution: USA: Colorado east of the Continental Divide. 8 e. Davidina uhleri reinthali (F. M. Brown, 1953) (Oeneis uhleri reinthali). American Mus. Novitates (1625): 10 – 11. TL: Gothic, Colorado. HT in AMNH. Distribution: USA: western slope of the Continental Divide in Colorado. 9. Davidina urda (Eversmann, 1847) 9 a. Davidina urda urda (Eversmann, 1847) (Hipparchia urda). Bull. Soc. Nat. Mosc. 20 (3): 69, t. 2, f. 1 – 4. TL: ‘ Dauria’, Russia. TM in ZISP. = umbra Staudinger, 1892 (Oen. urda Ev. var. Umbra). Iris 5: 335. TL: ‘ Südamur’, Russia. TM in ZMHU. = laeta Austaut, 1908 (Oeneis urda Evers. var. laeta). Ent. Zeitschr. 22: 44. TL: ‘ Sayan’. TM -?. = trybomi Bryk, 1946 (Oeneis urda tribomi). Arkiv Zool. 38 A (3): 25, t. 4, fig. C 3. TL: ‘ Krasnojarsk’, [S. Siberia, Russia]. HT in NRS. = rasa Bryk, 1946, Arkiv Zool. 38 A (3): 24 (an infrasubspecific name). ab. albidior Austaut, 1908. Ent. Zeitschr. 22: 44. TL: ‘ Sayan’. TM -? ab. banghaasi Austaut, 1908. Ent. Zeitschr. 22: 44. TL: ‘ Sayan’. TM -? Distribution: Russia: N. Altai (Tongosh Mountains), S. Siberia, Transbaikal, Amur and Primorye regions, S. and Central Yakutia. N. Mongolia. N. E. China. 9 b. Davidina urda tschiliensis (O. Bang-Haas, 1933) (Oeneis urda tschiliensis). Ent. Zeitschr. 47: 98. TL: Hsingan mont., Tunkia-jingze, Prov. Tschili, China sept. HT is probably in ZMHU. Distribution: N. E. China: Hebei. 9 c. Davidina urda monteviri Bryk, 1946 (Oeneis urda monteviri). Arkiv Zool. 38 A (3): 24, t. 4, fig. A 3. TL: ‘ Shinten’, Korea. HT in NRS (?). Distribution: N. Korea.	en	Lukhtanov, Vladimir A, Dubatolov, Vladimir V (2020): Phylogenetic position and taxonomic rearrangement of Davidina (Lepidoptera: Nymphalidae), an enigmatic butterfly genus new for Europe and America. Zoological Journal of the Linnean Society 190 (3): 1036-1053, DOI: 10.1093/zoolinnean/zlaa104, URL: https://academic.oup.com/zoolinnean/article/190/3/1036/5907879
