taxonID	type	description	language	source
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	description	urn: lsid: zoobank. org: act: D 1829548 - 6533 - 4834 - 835 A-CB 37 A 394 CA 64 Figs 7 C, 8 – 9	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	diagnosis	Diagnosis Acipes alicantinus sp. nov. differs from the other Acipes species by the following combination of characters: absence of eyes (eyes present in A. atlanticus, A. celtibericus sp. nov., A. continentalis, A. decolor Enghoff, 1983, A. franzi (Loksa, 1967), A. lateralis Enghoff, 1983, A. oligops sp. nov., A. portosantoensis Enghoff, 1983, A. serratus Enghoff, 1983 and A. waldeni); flange in posterior gonopods present (absent in A. bifilum); first pair of legs of male modified, with no claw visible (not modified and with visible claw in A. machadoi); tibiae of first pair of legs of male slender, more than twice as long as wide, subapical emargination of coxal process of anterior gonopods subtle and distalmost part of anterior gonopods, narrower than prior to the emargination, flange of posterior gonopods lanceolate, accessory claw in midbody legs not visible (tibiae of first pair of male legs around 1.5 times as long as wide, coxal process of anterior gonopods similar in width along all their length, flange of posterior gonopods truncate or rounded, accessory claw of midbody legs visible in A. andalusius).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	etymology	Etymology The specific epithet ' alicantinus ', refers to the province of Alicante, where this species was found, adjective.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	materials_examined	Type material Holotype SPAIN • ♂; Alicante, Confrides, Sierra de Aitana, MSS; 38 ° 39 ′ 26 ′′ N, 0 ° 17 ′ 54 ′′ W; 1350 m a. s. l.; 16 May 2013 (trap set on 1 Nov. 2012); J. D. Gilgado, V. M. Ortuño et al. leg.; MNCN 20.07 / 2169. Paratypes SPAIN • 5 ♂♂, 2 ♀♀; same collection data as for holotype; MNCN 20.07 / 2170 - 76 • 2 ♂♂, 1 ♀; same data as for preceding; 24 Mar. 2012 (trap set on 26 Jan. 2012); NHMD. Other material examined SPAIN • 8 ♂♂, 1 ♀; same data as for holotype; 27 Jul. 2012 (trap set on 24 Apr. 2012); NHMD • 2 ♂♂, 6 juveniles; same data as for holotype; 16 May 2013 (trap set on 1 Nov. 2012); UAH • 5 ♂♂; same data as for holotype; 30 Nov. 2013 (trap set on 16 May 2013); UAH • 1 ♂; Cocentaina, Sierra de Mariola, Montcabrer, near Mas de Llopis, MSS; 38 ° 45 ′ 22 ′′ N, 0 ° 28 ′ 57 ′′ W; 1210 m a. s. l.; 19 Mar. 2012 (trap set on 13 Jan. 2012); J. D. Gilgado, V. M. Ortuño et al. leg.; NHMD • 2 ♂♂, 3 ♀♀; same data as for preceding; 14 Jan. 2014 (trap set on 9 May 2013); UAH.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	description	Description Light brown to pale amber colour in preserved specimens (Fig. 7 C). Males 7 – 11 mm long, and 0.68 mm in maximum vertical diameter, with 29 – 40 podous rings including collum + 2 – 4 apodous rings + telson. Measures taken on five randomly chosen males. Females 11 – 13 mm long, and 0.7 mm in maximum height, with 38 – 43 podous rings including collum + 2 apodous rings + telson. Measures taken on five randomly chosen females. HEAD. Males with parrot beak modification of cardo and stipes (Fig. 8 A – B). Labrum with 3 teeth and 6 + 6 labral and 2 + 2 supralabral setae (Fig. 8 B). Cephalic capsule with a few smaller setae laterally, 1 + 1 short setae on frons, and 1 + 1 long setae on vertex. Eyeless. Antenna with antennomere 5 with a distal ring of thick sensilla basiconica, antennomere 6 with half such ring of thinner sensilla (sensilla absent ventrally), and antennomere 7 with a dorsodistal row of smaller sensilla basiconica (Fig. 8 A, C). Antennomere lengths (mm): 0.05, 0.22, 0.22, 0.19, 0.21, 0.16, 0.06, 0.04. TRUNK. Collum with 4 + 4 setae. Rings of anterior part of body with 5 + 5 metazonital setae, increasing to ca 14 at midbody rings, and 16 + 16 at final body rings (Fig. 8 D – E). Length of midbody setae around 22 % of midbody ring height. 6 to 7 striae in ventral half of the ring. Telson: epiproct with 1 + 1 long dorsal, 3 + 3 lateral, and 1 + 1 ventral setae (Fig. 8 E). Hypoproct with 1 + 1 setae, and paraprocts with 2 setae each. LEGS. First pair of male legs modified, with no visible claw (Fig. 8 F). Tibiae of first pair of legs of male slender, more than twice as long as wide, and with two stout apophyses, and tarsus remnant reduced, almost absent. Legs of first half of body of males with up to six lanceolate, transversely striate setae on femur (1 – 2), postfemur (2) and tibia (2) (Fig. 8 G). Number of lanceolate setae decreasing towards and after midbody legs, absent in posteriormost third of the body (Fig. 8 H). Lanceolate setae absent in females. Accessory claw in midbody legs not visible, but present in anteriormost legs (Fig. 8 G – H). ANTERIOR GONOPODS. Coxal process (cxp) with subapical emargination subtle, almost absent, distalmost part narrower than prior to the emargination. Telopodites (tp) around a third of the length of coxal processes (Fig. 9 A, C). POSTERIOR GONOPODS. Curved, distal part anteriorly serrulate, including a filamentous tip (ft), and distal lanceolate flange (af) with apical margin serrulate (Fig. 9 B, D).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	distribution	Type locality The type locality is a colluvial debris slope at the base of Peña Cacha in the Sierra de Aitana massif, Alicante (Fig. 4 A). This mountain range has the highest peak in Alicante (1558 m a. s. l.). The bedrock consists of Eocene limestones (Alfaro et al. 2006), whose fragmentation during the cold Quaternary periods formed the debris (Marco 2000). The climate features extremes, with summer temperatures exceeding 30 ° C and winter snow cover. Annual precipitation ranges between 800 and 900 mm. The debris forms scree and colluvial fans, partially covered by sparse soil and vegetation. Sierra de Aitana represents a high Mediterranean mountain environment of significant biogeographic interest (Marco 2001). It harbours refugial Eurosiberian species (e. g., Taxus baccata L., Rhamnus alpina L.) and marks the northern range limit for several Baetic endemics (e. g., Genista longipes Pau, Vella spinosa Boiss). The dominant shrubs are Salvia lavandulifolia subsp. mariolensis (Figuerola) Alcaraz & De la Torre, Erinacea anthyllis Link, and Hormathophylla spinosa L.; the debris is colonized by Hedera helix subsp. rhizomatifera McAll., grasses (Festuca L. spp., Arrhenatherum P. Beauv. spp.), and mosses. Acipes alicantinus sp. nov. was also found in a scree slope in the Sierra de Mariola Natural Park, a rugged area spanning Alicante and Valencia provinces, formed by cretaceous limestone bedrock. Its highest peak, Montcabrer (1390 m a. s. l.), is the third highest in Alicante. The Mediterranean climate exhibits high microclimatic diversity, with frequent winter snow at higher elevations and significant cryptoprecipitation (fog, dew) in temperate seasons. This humidity supports relict flora, such as the “ Micro-reserve of the Teixera d'Agres ”, which contains the southernmost Taxus baccata forest in Europe (Generalitat Valenciana 2006; Gualda Gómez 1988).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	discussion	Remarks Four individuals of Acipes alicantinus sp. nov. from Sierra de Aitana were previously recorded in the literature as Acipes andalusius by Enghoff & Reboleira (2013) and Jiménez-Valverde et al. (2015). The present records of Acipes alicantinus from Mont Cabrer, Cocentaina, 20 km apart from the type locality, represent a new, second locality of the species. These two sites are 250 km apart from the type locality of its closest relative, Acipes andalusius sp. nov.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	type_taxon	Type species Acipes atlanticus Attems, 1937.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	diagnosis	Diagnosis (modified from Enghoff 1983) When present, ommatidia in a more or less regular row. First pair of male legs incrassate (except in A. machadoi) with one or two tibial apophyses (except in A. machadoi). Anterior gonopods with separate, apically dilated coxal apophyses, telopodites without setae. Posterior gonopods with a slender, curved shaft carrying a ventral rugose furrow, apically with a dorsal or lateral flange (no flange in A. waldeni Enghoff, 1983 and A. bifilum). Vulvae without setae. Operculum much longer than bursa, rugose, apically emarginate. Bursa subspherical, with a subspherical to subcylindrical receptaculum seminis.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	description	Discussion Morphology The genus Acipes presents relatively simple gonopods, which provides few characters for species delimitation. Anterior gonopods are quite similar across species, leaving as main characters the relative length / width of the coxal processes, their distal emargination, the shape of their tip, and the relative length of telopodites. Posterior gonopods present more variability across species, with characters such as their relative length / width, the presence or absence of a distal flange with different shapes, serrulated or not, the presence or absence of a filamentous tip of different lengths, serrulated or not, etc. Some species, such as A. bifilum, A. lateralis, or A. waldeni present clearly distinct posterior gonopods; but those of the remaining species are more similar among them. The species A. alicantinus sp. nov., A. andalusius, A. celtibericus sp. nov. A. continentalis, A. oligops sp. nov., A. portosantoensis and A. serratus present quite similar posterior gonopods (with a serrulated filamentous tip and a flange) and they are similar, to a lesser extent, to A. atlanticus and A. franzi and A. machadoi (flange or filamentous tip more variable). Based only on posterior gonopod morphology, it is tempting to sketch a species group that includes most of the Iberian species, except A. bifilum, and the Macaronesian species A. serratus. However, the similarity of posterior gonopods alone is probably not enough evidence for phylogenetic proximity. Despite the general simplicity and similarity of Acipes gonopods, there are several other characters that can be used in combination with them for species delimitation. The most relevant ones are the shape of the first pair of male legs, presence and number of eyes, length of setae, number of rings and body size (Enghoff 1983, 1986; Enghoff & Mauriès 1999; Enghoff & Reboleira 2013). The three here described species share a similar gonopod morphology with the Iberian species A. continentalis and A. andalusius as explained below. Acipes celtibericus sp. nov. is most similar to A. continentalis (size, colour, number of ommatidia, shape of first pair of legs of male, posterior gonopods with filamentous tip and serrulate flange, accessory claw in midbody legs, etc.), but differs in the shape of the coxal processes of anterior gonopods, and the shape of the flange of posterior gonopods. The gonopods of Acipes celtibericus sp. nov. also shows resemblances with the geographically nearest species, A. oligops, A. andalusius and A. alicantinus sp. nov. (serrulate apical margin of flange, filamentous tip), but A. celtibericus differs in several other characters such as the first pair of male legs and number of ommatidia. As mentioned above, their gonopods also seem to show more similarities with Macaronesian species, such as A. serratus, than with the troglobiont continental Portuguese species. Acipes oligops sp. nov., is similar to A. continentalis and A. celtibericus sp. nov. in the general shape of its gonopods (presence of a serrulated filamentous tip and flange), but resembles more A. continentalis (flange lanceolate, similar emargination of coxal processes of anterior gonopods). In addition to being clearly distinct from congeneric species in the structure of the first pair of legs, Acipes oligops is also notable for its very small number of ommatidia (2 to 3 per side, Fig. 17 A.) and its size: it is thinner than all other known adult males of Acipes, but has a relatively high number (40) of podous rings: adult males of the smallest Macaronesian species, A. serratus Enghoff, 1983, with 40 podous rings thus have a body diameter of 0.45 – 0.53 mm (Enghoff 1983: fig. 59), whereas the holotype of A. oligops only measures 0.36 mm in diameter. Acipes alicantinus sp. nov. is morphologically very similar to Acipes andalusius, but it shows several differences that justify its description as a new species. The first pair of legs of male Acipes alicantinus is slenderer (Figs 8 F, 10 F; Enghoff 1983: fig 61). The shape of the distal part of coxal processes of anterior gonopods is different, with a more acute ending in Acipes alicantinus (Fig. 9 A, C), while it is blunter in Acipes andalusius (Fig. 10 A – B). Acipes alicantinus has a more lanceolate and narrower flange of the posterior gonopod (Fig. 9 B, D), while it is rounder or more truncate in Acipes andalusius (Fig. 10 C – E). Acipes alicantinus also differs in its body size from Acipes andalusius (Fig. 7 B – C). The two species overlap in their size ranges, but Acipes alicantinus is shorter and with fewer body rings than Acipes andalusius (see Enghoff & Mauriès 1999; Enghoff & Reboleira 2013) (Fig. 7 B – C).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	distribution	Habitat and distribution The genus Acipes is distributed across Macaronesia and the Iberian Peninsula, but it seems to have a stronger preference for subterranean habitats in the Iberian Peninsula (Enghoff 1983; Enghoff & Mauriès 1999; Enghoff & Reboleira 2013). Troglobiont species are distributed in southern Spain and southern Portugal (A. andalusius, A. bifilum and A. machadoi, cave specialists) (Enghoff & Mauriès 1999; Enghoff & Reboleira 2013) and eastern Spain (A. alicantinus sp. nov., MSS). The species from central Spain inhabit the MSS but they seem not to be MSS specialists. However, limited sampling efforts in both subterranean and epigean environments suggest that future discoveries could potentially alter these observed patterns. In Macaronesia, species are found exclusively in epigean habitats (Enghoff 1983; Kime & Enghoff 2017). Two Macaronesian species are associated with deadwood (Enghoff 1983; Kime & Enghoff 2017), and at least one of the new records of A. continentalis was from this microhabitat. This suggests that deadwood could serve as a suitable habitat for Acipes species and should be targeted for future samplings. Central Iberian species (A. continentalis, A. celtibericus sp. nov., and A. oligops sp. nov.) occur in the MSS and, except Acipes celtibericus, also in surface habitats. The absence of the latter on the surface might be attributed to the lack of sampling in these habitats (only MSS was sampled in the Iberian System). Its morphology suggests a lifestyle similar to that of A. continentalis. The reduced number of ommatidia and the reduced size of A. oligops compared to other Acipes suggests that this species may be predominantly edaphic or subterranean. The capture of only one individual after several years of sampling in the MSS, and only one additional individual on the surface after four hours of visual searching, indicates that it is a species with a low abundance. Another possibility is that the temporal watercourse (Fig. 2) may not be its primary microhabitat. The species might inhabit the soils surrounding the watercourse, entering it only occasionally. The landscape where Acipes oligops was found (slate debris within a limestone landscape, Fig. 2) differs from those where A. continentalis and A. celtibericus were found (mostly siliceous, quartzitic or granitic terrain in the mountains, Figs 1, 3). Three southern Iberian species (A. andalusius, A. bifilum, and A. machadoi) are restricted to caves, and they can be classified as troglobionts due to their exclusive presence in hypogean habitats and lack of eyes (Enghoff & Reboleira 2013). Acipes alicantinus sp. nov. is also a blind subterranean species, so it can be considered as an MSS specialist troglobiont as well. The difference in size between A. andalusius and A. alicantinus may be related to the different size of the voids they inhabit (larger in caves, smaller in the MSS).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B776FFCEFD9AFB8F41FFF858.taxon	description	Figs 7 A – B, 10 Records in the literature The species was described from 20 specimens collected in a cave in Peal del Becerro, Jaén, Spain (Enghoff & Mauriès, 1999), but the name of the cave (PB- 2) was only mentioned later by Enghoff & Reboleira (2013). In this same publication they mentioned a personal communication from the collector, who stated that the species was also present in a nearby cave (PB- 4). Eight further specimens were reported from collections made in the decade of 1990 by Mauriès (2013), five of them from the cave “ Cueva de la Pasada de Algeciras ”, Hornachuelos, Córdoba, Spain, and three other specimens from “ Pasada Jeira ” (most probably misspelling of “ Pasada de Algeciras ”, thus same locality).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B776FFCEFD9AFB8F41FFF858.taxon	materials_examined	Material examined SPAIN • 1 ♂, 1 subadult ♂, 1 ♀; Jaén, Peal del Becerro, Cave PB- 4, ca 400 m from entrance; 37 ° 47 ′ 56.8 ′′ N, 2 ° 53 ′ 41.5 ′′ W; 1010 m a. s. l.; 15 Aug. 1994; A. Tinaut leg.; UAH.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B776FFCEFD9AFB8F41FFF858.taxon	discussion	Remarks The studied male specimen from PB- 4 shows some minor differences from that of the original description from PB- 2. The apical flange is less truncated and more rounded in its apical margin than shown in the drawing of the original description (Fig. 10 C – D). The first pair of male legs was not drawn in the original description, but it is stated to be identical to those of A. serratus. However, in the studied specimen the first pair of legs is less modified, the tarsal remnant is slightly more developed and shows a visible terminal claw (Fig. 10 F), not drawn in the original description of A. serratus (see Enghoff 1983: fig. 61). The specimens from Córdoba (Sierra Morena) were tentatively assigned to Acipes aff. andalusius by Mauriès (2013), but Reboleira & Enghoff (2017) considered them as Acipes andalusius.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B777FFCCFD95FED34616FD8F.taxon	description	Fig. 11 A – B Records in the literature This species was described from two specimens collected from the cave “ Gruta da Senhora ”, Moncarapacho, Algarve, Portugal (Enghoff & Reboleira 2013), and it is only known from the type series.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B777FFCCFD95FED34616FD8F.taxon	discussion	Remarks Enghoff & Reboleira (2013) identified several morphological similarities between this species and Acipes machadoi, such as their high number of podous rings and their short anterior gonopod telopodites. Based on these traits and the geographical proximity of the two species, the authors suggested a potential close relationship between them. However, Enghoff & Reboleira (2013) also noted significant differences between the posterior gonopods of these two species (Fig. 11 B – C). They also indicated that those of A. bifilum do not resemble those of any other species.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	description	urn: lsid: zoobank. org: act: 9 FF 8003 B-B 3 FC- 4 F 07 - 8064 - 9552 FF 5 F 5 B 09 Figs 7 D, 12 – 15	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	diagnosis	Diagnosis Acipes celtibericus sp. nov. differs from the other Acipes species by the following combination of characters: at least four ommatidia per side (whereas A. alicantinus sp. nov., A. andalusius, A. bifilum and A. machadoi are anophthalmous; and A. oligops sp. nov. has only two or three), a developed accessory claw in the midbody legs (absent in A. decolor and A. portosantoensis), anterior gonopods with shallow subapical emargination (deep emargination in A. atlanticus), posterior gonopods with a filamentous tip (absent in A. franzi and A lateralis), and a truncate and serrulate flange (flange absent in A. waldeni, and lanceolate in A. continentalis and A. serratus).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	etymology	Etymology The specific epithet ' celtibericus ', refers to the pre-Roman people (Celtiberians) who inhabited the region of central Iberian Peninsula where this species was found, adjective.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	materials_examined	Type material Holotype SPAIN • ♂; Burgos, Pineda de la Sierra, Sierra de Mencilla in Sierra de la Demanda, beech forest, MSS; 42 ° 11 ′ 55 ′′ N, 3 ° 18 ′ 41 ′′ W; 1470 m a. s. l.; 18 Jun. 2018 (trap set on 26 Jun. 2017); V. M. Ortuño, J. D. Gilgado and E. Ledesma leg.; MNCN 20.07 / 2106. Paratypes SPAIN • 5 ♂♂, 5 ♀♀; same collection data as for holotype; MNCN 20.07 / 2107 - 2116 • 5 ♂♂, 5 ♀♀; same data as for preceding; UAH • 5 ♂♂, 5 ♀♀; same data as for preceding; 22 Jun. 2019 (trap set on 18 Jun. 2018); NHMD. Other material examined SPAIN • 33 ♂♂, 9 ♀♀, 8 juveniles; same collection data as for holotype; UAH • 333 ♂♂, 100 ♀♀, 21 juveniles; same data as for preceding; 22 Jun. 2019 (trap set on 18 Jun. 2018), UAH • 106 ♂♂, 37 ♀♀, 10 juveniles; same data as for preceding; 3 Aug. 2020 (trap set on 22 Jun. 2019); UAH • 1 ♂, 1 subadult ♂, 2 ♀♀, 1 juvenile; Pineda de la Sierra, Sierra de Mencilla in Sierra de la Demanda, Glaciar Circus, MSS in bare scree; 42 ° 11 ′ 21 ′′ N, 3 ° 18 ′ 37 ′′ W; 1750 m a. s. l.; 18 Jun. 2018 (trap set on 26 Jun. 2017); V. M. Ortuño, J. D. Gilgado and E. Ledesma leg.; UAH • 1 ♂, 1 ♀, 1 juvenile; same data as for preceding; 22 Jun. 2019 (trap set on 18 Jun. 2018); UAH • 1 ♀; Zaragoza, Añón de Moncayo, Moncayo Massif, Collado Bellido, MSS in bare Scree; 41 ° 46 ′ 1 ′′ N, 01 ° 47 ′ 01 ′′ W; 1540 m a. s. l.; 12 Dec. 2012 (trap set on 26 Sep. 2012); V. M. Ortuño, J. D. Gilgado, E. Ledesma and E. Cuesta leg.; NHMD • 1 ♂, 1 juvenile; same data as for preceding; 25 Mar. 2013 (trap set on 12 Dec. 2012); NHMD • 1 ♂; same data as for preceding; 12 Mar. 2014 (trap set on 23 Oct. 2013); UAH.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	description	Description Pale brown to dark amber colour in preserved specimens (Figs 7 D, 12 A – B). Males 10.5 – 12.5 mm long, and 0.52 – 0.63 mm in maximum ring vertical diameter, with 35 – 38 podous rings including collum + 2 apodous rings + telson. Measures taken on 10 randomly chosen males. Females 11.5 – 14.5 mm long, and 0.58 – 0.73 mm in maximum height, with 38 – 40 podous rings including collum + 2 (1) apodous rings + telson. Measures taken on 10 randomly chosen females. HEAD. Males with parrot beak modification of cardo and stipes (Fig. 12 A – D). Labrum with 3 teeth and 6 + 6 labral and 2 + 2 supralabral setae (Fig. 12 D – F). Cephalic capsule with several smaller setae laterally, 1 + 1 setae on frons, and 1 + 1 long setae on vertex (Fig. 12 A – C, E). One row of 4 – 6 blackish ommatidia (Fig. 12 A), but some can be displaced (poorly visible on SEM (Fig. 12 C). Antenna with antennomere 5 with a distal ring of thick sensilla basiconica, antennomere 6 with half such ring (sensilla absent ventrally), and antennomere 7 with a dorsodistal row of shorter sensilla basiconica (Fig. 12 C, E). Antennomere lengths (mm): 0.05, 0.17, 0.16, 0.13, 0.16, 0.13, 0.05, 0.04. TRUNK. Collum with 4 + 4 setae (Fig. 12 C). Second ring with 4 + 4 setae, increasing up to 12 + 12 long setae on the last rings of the body (Figs 12 C, 15 C, E). Length of midbody setae around 18 % of midbody ring height. Around 10 striae in ventral half of the rings. Telson: epiproct with 2 + 2 long dorsal, 1 + 1 lateral, and 1 + 1 ventral setae. Hypoproct with 1 + 1 setae, and paraprocts with 2 setae each (Fig. 15 E). LEGS. First pair of legs modified (Fig. 13 A). Tibiae of first pair of legs of male robust, twice as long as wide, and with two curved, tusk-like apophyses, the proximal one larger than the distal one. Tarsus remnant visible, with terminal claw present. All legs of males, except in posteriormost third of body, with lanceolate, transversely striate setae on femur (2), postfemur (2) and tibia (2) (Fig. 13 B, D). Number of lanceolate setae decreasing towards and after midbody legs. Lanceolate setae absent in females. Accessory claw visible in midbody legs (Fig. 13 D). ANTERIOR GONOPODS. Coxal processes (cxp) with shallow subapical emargination, telopodites (tp) slightly more than a third of the length of coxal processes. Distal part of coxal processes more acute in type locality (Fig. 14 A – C) than in the specimens from Moncayo Massif (Fig. 15 A). POSTERIOR GONOPODS. Curved, distal part anteriorly serrulate, including a relatively long filamentous tip (ft), and a truncate apical flange (af) serrulate in its apical margin (Figs 14 C, E – F, 15 B).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	distribution	Type locality The Sierra de Mencilla, in the northwestern Sierra de la Demanda (Burgos, Spain), reaches 1932 m a. s. l. at Pico Mencilla. It is composed mainly of Paleozoic slates, quartzitic sandstones, and conglomerates (Fuentes Cabrera 1981). Its morphology is asymmetrical: the southern slopes descend smoothly towards the Arlanzón valley, while the northern slopes are steep and host large block fields and colluvial cones such as “ La Concha ” (Fuentes Cabrera 1981). During the Quaternary, small glaciers developed on its northern side, leaving cirques and moraines around Pineda de la Sierra. Cosmogenic dating has shown that fossil debris-covered glaciers on the northern slope of Mencilla persisted from the Last Glacial Maximum (~ 17.8 ka) to the Oldest Dryas (~ 16.5 ka), and possibly survived into the Holocene under protective debris mantles (Fernández-Fernández et al. 2017). The climate is cold and continental montane, with annual precipitation ranging between 700 and 1200 mm depending on altitude, and frequent snowfall in winter (Fuentes Cabrera 1981). Biogeographically, the Sierra de Mencilla lies at the interface of Eurosiberian and Mediterranean sectors, which enhances species diversity. Forests of Pyrenean oak (Quercus pyrenaica Willd.) dominate mid-elevations, with beech (Fagus sylvatica L.) on humid north-facing slopes, while higher zones support Scots pine (Pinus sylvestris L.), heathlands, and montane grasslands (Fuentes Cabrera 1981). Acipes celtibericus sp. nov. was also found in bare scree on the northern slope of the Moncayo Massif. This mountain is the highest peak of the Iberian System (2314 m a. s. l.), and it is located between the provinces of Zaragoza (Aragón) and Soria (Castilla y León). Its geology is characterized by siliceous materials, including angular blocks of Mesozoic and Palaeozoic quartzites and sandstones fractured by ice, accumulated through solifluction processes during the cold periods of the Quaternary (Pellicer 1980). The climate is temperate and humid, with notable differences between slopes: the northern slope, wetter due to interception of Atlantic fronts via the Ebro valley, supports forests typical of Atlantic climates, contrasting with the surrounding Mediterranean environment, making Moncayo an “ Atlantic island ” (Ballarín 1985; Navarro 1989). Annual precipitation exceeds 1000 mm on the wetter slopes, sustaining diverse vegetation including trees such as European beech (Fagus sylvatica), Scots pine (Pinus sylvestris), and black pine (Pinus uncinata Ramond ex A. DC.), even on stony screes at 1540 m a. s. l. where soil is scarce (Ballarín 1985; Navarro 1989).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	discussion	Remarks Acipes celtibericus sp. nov. was found in two mountains of the Iberian System, 130 km apart (Fig. 6). While the posterior gonopods in both populations are identical, the coxal apophysis of anterior gonopods of males from Moncayo Massif shows a more rounded distal part after the subapical emargination (Fig. 14 A), similar to that of A. continentalis (Fig. 16 C). This species has so far only been found in the MSS, but its presence in surface environments, including soil or deeper soil layers, cannot be discarded. The two populations were discovered in different macrohabitats. In the Mencilla Mountain, the species was significantly more abundant in a beech forest at 1470 m a. s. l. than in bare scree at 1750 m a. s. l .. However, in the Moncayo Massif, it was exclusively found in bare scree at 1540 m a. s. l., with no individuals captured in the beech forest at 1420 m a. s. l.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76EFFD7FDC3FED3472DFDCB.taxon	description	Figs 7 E, 16 Records in the literature This species was originally described from a single specimen collected at the “ Pico ” mountain pass (“ Puerto del Pico ”), located in Cuevas del Valle, Sierra de Gredos, Ávila, Spain (Enghoff 1986). Years later, Enghoff & Mauriès (1999) examined 59 specimens collected during the 1960 s and reported the presence of the species in the Sierra de Guadarrama, in the Navacerrada Mountain Pass, Cercedilla, and close to this locality in Peñalara Mountain, both in Madrid, Spain.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76EFFD7FDC3FED3472DFDCB.taxon	materials_examined	Material examined Other material SPAIN • 1 ♂; Madrid, Cercedilla, Schmidt trail, deadwood; 40 ° 47 ′ 15 ′′ N, 4 ° 03 ′ 02 ′′ W; 1830 m a. s. l.; 27 Mar. 2017; E. Recuero leg.; UAH • 1 ♂; Madrid, La Cabrera, cleared forest of Quercus ilex; 40 ° 51 ′ 25.32 ′′ N, 3 ° 37 ′ 19.25 ′′ W; 1030 m a. s. l.; 27 Dec. 2020; D. Cabanillas leg.; UAH • 1 ♀; Madrid, Rascafría, Sierra de Guadarrama National Park (henceforth SGNP), Alto de los Poyales, MSS; 40 ° 53 ′ 40.48 ′′ N, 3 ° 56 ′ 10.13 ′′ W; 1890 m a. s. l.; 14 Oct. 2016 (trap set on 9 Jun. 2016); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; UAH. • 1 ♀; Madrid, Rascafría, SGNP, Scree, between peaks Cabeza de Hierro Mayor and Cabeza de Hierro Menor, MSS; 40 ° 47 ′ 54.79 ′′ N, 3 ° 56 ′ 6.45 ′′ W; 2300 m a. s. l.; 14 Oct. 2016 (trap set on 9 Jun. 2016); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; UAH. • 1 ♀, 1 juvenile; Madrid, Cercedilla, SGNP, Cerro Ventoso, Scree, MSS; 40 ° 47 ′ 28.83 ′′ N, 4 ° 3 ′ 24.32 ′′ W; 1870 m a. s. l.; 17 Sept. 2015 (trap set on 9 Jun. 2015); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; NHMD • 1 ♀; same data as for preceding; 21 Sept. 2016 (trap set on 9 Jun. 2016); UAH • 3 ♂♂; Madrid, Rascafría, SGNP, Collado de Valdemartín, Scree, MSS; 40 ° 47 ′ 43.48 ′′ N, 3 ° 57 ′ 20.81 ′′ W; 2155 m a. s. l.; 6 Nov. 2015 (trap set on 3 Jul. 2015); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; NHMD • 2 ♂♂; Madrid, Rascafría, SGNP, Collado de la Flecha, La Gelecha, Scree, MSS; 40 ° 55 ′ 25.95 ′′ N, 3 ° 55 ′ 16.92 ′′ W; 1865 m a. s. l.; 9 Jun. 2016 (trap set on 6 Oct. 2015); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; UAH • 1 ♀; Madrid, Manzanares el Real, SGNP, La Maliciosa, Collado del Piornal, scree, MSS; 40 ° 46 ′ 17.26 ′′ N, 3 ° 58 ′ 16.14 ′′ W, 2100 m a. s. l.; 22 Sep. 2015 (trap set on 9 Jun. 2015); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; NHMD • 1 ♂; same data as for preceding; 26 May 2016 (trap set on 22 Sept. 2015); UAH • 1 ♂, 2 ♀♀; Segovia, Real Sitio de San Ildefonso, SGNP, Falda del Cerro de la Muela, Cancho del Río Peces, MSS; 1605 m a. s. l.; 17 Sept. 2015 (trap set on 20 May 2015); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; NHMD • 7 ♂♂, 23 ♀♀; same as for preceding; 1 May 2016 (trap set on 17 Sept. 2015); UAH • 1 ♂, 15 ♀♀; same data as for preceding; 21 Sep. 2016 (trap set on 1 May 2016); UAH • 1 juvenile; Segovia, Real Sitio de San Ildefonso, SGNP, Falda del Montón de Trigo, Corrales de la Majada Minguete; MSS; 40 ° 47 ′ 45.00 ′′ N, 4 ° 03 ′ 57.98 ′′ W; 1820 m a. s. l.; 17 Sep. 2015 (trap set on 20 May 2015); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; NHMD • 7 ♂♂, 11 ♀♀; same data as for preceding; 26 May 2016 (trap set on 17 Sep. 2015); UAH • 1 ♀; same data as for preceding; 21 Sep. 2016 (trap set on 26 May 2016); UAH • 7 ♂, 2 ♀♀; Segovia, Real Sitio de San Ildefonso, SGNP, Loma del Noruego, Canchal de La Pedriza, MSS; 40 ° 48 ′ 52.12 ′′ N, 3 ° 58 ′ 20.87 ′′ W; 1790 m a. s. l.; 22 Sep. 2016 (trap set on 1 May 2016); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; UAH • 1 ♀, 1 juvenile; Segovia, Navafría, SGNP, Pine forest beside Navafría shelter, MSS (buried pitfall trap); 40 ° 59 ′ 12.92 ′′ N, 3 ° 48 ′ 51.60 ′′ W; 1780 m a. s. l.; 22 Sep. 2015 (trap set on 24 Jun. 2015); V. M. Ortuño, J. D. Gilgado, E. Ledesma et al. leg.; NHMD • 5 ♂♂, 2 ♀♀; same data as for preceding; 13 Jun. 2016 (trap set on 22 Sep. 2015); UAH • 1 ♂, 1 ♀; same data as for preceding; 40 ° 59 ′ 12.84 ′′ N, 3 ° 48 ′ 52.57 ′′ W; UAH.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76EFFD7FDC3FED3472DFDCB.taxon	discussion	Remarks This species seems to have a relatively wide elevational range, from 930 to 2300 m a. s. l., and is found in surface habitats, under stones and in deadwood, and MSS. This species has been recorded in Sierra de Gredos (holotype) and Sierra de Guadarrama mountain ranges, and the new records indicate that it might be widespread in the latter (Fig. 6). The type specimens of A. continentalis from Gredos and the specimens from Guadarrama (100 km distance) also show some differences according to Enghoff & Mauriès (1999), such as the different development of the terminal claw of the modified first pair of legs of the male in the Guadarrama specimens, apical flange (af) protruding parallel to the filamentous tip (Fig. 16 A – B, D – E), and apical flange of Guadarrama specimens without spines (but spines can be observed under SEM, Fig. 16 E). Enghoff (1986) already questioned that this developed claw in the first leg pair of the holotype could be an individual aberration; however, more material from the type locality should be examined to test it.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76CFFD4FD84FED3407AFDC6.taxon	description	Fig. 11 C – D Records in the literature This species is known only from one male specimen which was captured in the cave “ Gruta do Vale Telheiro ”, Loulé, Algarve, Portugal (Enghoff & Reboleira 2013).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76CFFD4FD84FED3407AFDC6.taxon	discussion	Remarks This is the largest species of Acipes, both in size and number of rings (Enghoff & Reboleira 2013).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76CFFDBFD97FDAE44EFFDAF.taxon	description	urn: lsid: zoobank. org: act: 84 E 0 CD 53 - 129 F- 4 EAB-B 4 F 9 - D 7 FA 7 F 2 A 72 A 0 Figs 7 F, 17	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76CFFDBFD97FDAE44EFFDAF.taxon	diagnosis	Diagnosis Acipes oligops sp. nov. differs from A. andalusius, A. alicantinus sp. nov., A. bifilum, and A. machadoi by the presence of ommatidia, and from the other eyed species of Acipes by having only two to three ommatidia on each side of the head (vs 4 – 15 in A. atlanticus, 4 – 6 in A. celtibericus sp. nov., 5 – 8 in A. continentalis, 5 – 9 in A. decolor, 6 – 8 in A. franzi, 6 – 9 in A. lateralis, 5 – 10 in A. portosantoensis, 4 – 9 in A. serratus, and 4 – 12 in A. waldeni). Acipes oligops sp. nov. further differs from all these species by the unique shape of the first pair of legs of the male (except A. decolor which is known only from females), with a single stout, claw-like tibial apophysis, vs apophysis absent in A. machadoi; apophysis narrow in A. portosantoensis; two apophyses in A. alicantinus sp. nov., A. andalusius, A. atlanticus, A. bifilum, A. celtibericus sp. nov., A. continentalis, A. franzi, A. serratus, and A. waldeni; apophysis similar in proportions to A. lateralis, but the latter having a prominent tarsus, vs. almost absent in A. oligops sp. nov. Moreover, Acipes oligops sp. nov. differs from other species in gonopod shape (except A. decolor, male unknown): posterior gonopod with a filamentous tip and a lanceolate flange, serrulate in its distalmost part (filamentous tip absent in A. franzi and A. lateralis; flange absent in A. bifilum and A. waldeni; flange smooth in A. atlanticus, A. machadoi, and A. serratus; flange clearly truncated in A. celtibericus sp. nov. and A. portosantoensis, truncated or rounded in A. andalusius); anterior gonopod with a very slender coxal apophysis with a slight subapical emargination leaving a rounded distalmost part narrower than the rest of the coxal process (marked emargination but distalmost part as wide as the rest of coxal process in A. continentalis; almost no emargination and distalmost part not rounded in A. alicantinus sp. nov.).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76CFFDBFD97FDAE44EFFDAF.taxon	etymology	Etymology The specific epithet (Greek) means ' few eyes' and refers to the low number of ommatidia. Noun in apposition.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76CFFDBFD97FDAE44EFFDAF.taxon	materials_examined	Type material Holotype SPAIN • ♂; Guadalajara, Tamajón, Sierra del Ocejón, Arroyo de Valdelapuerta, MSS (alluvial); 41 ° 1 ′ 20 ′′ N, 3 ° 15 ′ 1 ′′ W; 1050 m a. s. l.; 9 Jun. 2013 (trap set on 11 Oct. 2012); J. D. Gilgado and V. M. Ortuño leg.; NHMD 1184664. Paratype SPAIN • 1 ♂; same locality as for holotype but under a stone on the surface of a temporal watercourse; 16 Mar. 2024; J. D. Gilgado and V. Martínez-Pillado leg.; MNCN 20.07 / 2167.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76CFFDBFD97FDAE44EFFDAF.taxon	description	Description Little to no pigmentation, whitish, ivory to amber colour (Figs 7 F, 17 A). Females unknown, description based on two males. Measurements: 7.6 – 8.7 mm; midbody vertical diameter 0.36 – 0.4 mm. 34 – 39 podous rings including collum + 1 – 3 apodous rings + telson. HEAD (FIG. 17 A). Males with parrot beak modification of cardo and stipes. Labrum with 3 teeth and 5 + 5 labral and 2 + 2 supralabral setae. Cephalic capsule with 1 + 1 setae on frons, and 1 + 1 long setae on vertex. Two clearly separated black ommatidia on each side of the head (three on left side in paratype) (Fig. 17 A). Antenna with antennomere 5 with a distal ring of thick sensilla basiconica, antennomere 6 with half such ring (sensilla absent ventrally), and antennomere 7 with a dorsodistal row of shorter sensilla basiconica. Antennomere lengths (mm): 0.08, 0.38, 0.30, 0.24, 0.31, 0.25, 0.11, 0.05. TRUNK. Collum with 4 + 4 setae. Midbody rings with 8 + 8 setae, increasing up to 10 + 10 on the last rings of the body. Length of midbody setae around 16 – 23 % of midbody ring height, setae longest on anterior part of body. Seven to eight visible striae in ventral half of the rings. Telson: epiproct with 2 + 2 long dorsal, 1 + 1 lateral, and 1 + 1 ventral setae. Hypoproct with 1 + 1 setae, and paraprocts with 2 setae each. LEGS. First pair modified (Fig. 17 B). Stout, podomeres distinct; prefemur, femur, postfemur and tibia forming a curved limb terminating in a very stout, claw-like tibial apophysis (apo); tarsus remnant (ta) tiny, with several spikes. All legs, except in posteriormost part of body, with up to five lanceolate, transversely striate setae on femur (1), postfemur (2) and tibia (2) (Fig. 17 E, arrows). Accessory claw strongly reduced but present in midbody legs (Fig. 17 E). ANTERIOR GONOPODS (FIG. 17 C). Very slender coxal processes (cxp) (12 times as long as wide) with only slight subapical lateral emarginations leaving a rounded distalmost part narrower than the rest of coxal process, and very slender telopodites (tp). Telopodites approximately ¼ of the length of coxal processes. POSTERIOR GONOPODS (FIG. 17 D). slender and curved, with a long, slender, spinose filamentous tip (ft) and a lanceolate apical flange (af) with a serrulate distal margin.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76CFFDBFD97FDAE44EFFDAF.taxon	distribution	Type locality The temporary watercourse of the Arroyo de Valdelapuerta in Tamajón, Guadalajara, is a narrow streambed carved into a calcareous environment but composed mainly of slates. The area is renowned for its distinctive geomorphological features, most notably extensive karstified formations of the Ciudad Encantada type, developed on Cretaceous dolomites and limestones (Carcavilla et al. 2025). According to the Köppen climate classification, the Tamajón region has a temperate, dry continental climate, intermediate between arid and humid conditions (Rivas Goday & Esteban 1944). It is characterized by long, harsh winters, with mean annual temperatures ranging from 9 to 14 ° C and pronounced thermal amplitudes. Frosts are frequent, and precipitation is scarce, typically between 350 and 600 mm per year. From a biogeographic perspective, the site lies within a transitional zone between the Mediterranean Iberian and Celtiberian-Alcarrean sectors (Rivas Martínez 1987). Vegetation in the gully surrounding the stream consists mainly of Juniperus thurifera L. and Quercus ilex L., with scattered Juniperus oxycedrus L., basophilic scrub (lavender, thyme, etc.), and several rupicolous and sub-rupicolous species growing on the rocky outcrops (de la Fuente García 1982).	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
03CF87D4B76CFFDBFD97FDAE44EFFDAF.taxon	discussion	Remarks Acipes oligops sp. nov. seems to be closely related to A. continentalis and A. celtibericus sp. nov., but its two known specimens show the same distinct combination of characters that warrant its recognition as a new species (small size, low number of ommatidia, and especially the unique shape of the first pair of legs). The location and habitat (lithology, vegetation, elevation, etc.) are also different from those known for A. continentalis and A. celtibericus sp. nov. The s hape of the first pair of legs in males clearly differs from all other Acipes species, being most similar to A. portosantoensis Enghoff, 1983, but with much stouter tibial apophysis.	en	Gilgado, José D., Enghoff, Henrik, Ortuño, Vicente M. (2026): The genus Acipes Attems, 1937 in the Iberian Peninsula (Diplopoda, Julida, Blaniulidae), with the description of three new species from Spain. European Journal of Taxonomy 1039: 251-283, DOI: 10.5852/ejt.2026.1039.3195, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3195/14187
