taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760646/files/figure.png	https://doi.org/10.5281/zenodo.18760646	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760648/files/figure.png	https://doi.org/10.5281/zenodo.18760648	Fig. 8. Anatomical details of Acipes alicantinus sp. nov. Gilgado & Enghoff. A. Head and first five rings of male specimen from type locality in left lateral view; 30 Nov. 2013; UAH. B. Head of another male specimen from same locality and date in ventral view; UAH. C. Tip of antenna of same specimen as A in dorsal view. D. Rings 6–9 of same specimen as A. E. Telson and last body rings of same specimen in left lateral view. F. First male legs of same specimen as B in posterior view. G. Leg 4 of same specimen as B. H. last leg of same specimen as B. Abbreviations: apo = tibial apophysis; ta = tarsus remnant. Scale bars: A, D–E = 0.2 mm; B–C, G = 0.1 mm; F–H = 0.05 mm.	Fig. 8. Anatomical details of Acipes alicantinus sp. nov. Gilgado & Enghoff. A. Head and first five rings of male specimen from type locality in left lateral view; 30 Nov. 2013; UAH. B. Head of another male specimen from same locality and date in ventral view; UAH. C. Tip of antenna of same specimen as A in dorsal view. D. Rings 6–9 of same specimen as A. E. Telson and last body rings of same specimen in left lateral view. F. First male legs of same specimen as B in posterior view. G. Leg 4 of same specimen as B. H. last leg of same specimen as B. Abbreviations: apo = tibial apophysis; ta = tarsus remnant. Scale bars: A, D–E = 0.2 mm; B–C, G = 0.1 mm; F–H = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760650/files/figure.png	https://doi.org/10.5281/zenodo.18760650	Fig. 9. Gonopods of Acipes alicantinus Gilgado & Enghoff sp. nov. A. Anterior gonopods of holotype in posterior view. MNCN 20.07/2169. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods of holotype in right lateral view. MNCN 20.07/2169. C. Anterior gonopods of male specimen from type locality (30 Nov. 2013) in posterior view. Arrow indicates subapical emargination of coxal process. D. Detail of flange and filamentous tip of posterior gonopods of same specimen as C in left lateral view. Abbreviations: af = apical flange; cxp = coxal process; ft = filamentous tip; tp = telopodite. Scale bars: A–C = 0.1 mm; D = 0.05 mm.	Fig. 9. Gonopods of Acipes alicantinus Gilgado & Enghoff sp. nov. A. Anterior gonopods of holotype in posterior view. MNCN 20.07/2169. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods of holotype in right lateral view. MNCN 20.07/2169. C. Anterior gonopods of male specimen from type locality (30 Nov. 2013) in posterior view. Arrow indicates subapical emargination of coxal process. D. Detail of flange and filamentous tip of posterior gonopods of same specimen as C in left lateral view. Abbreviations: af = apical flange; cxp = coxal process; ft = filamentous tip; tp = telopodite. Scale bars: A–C = 0.1 mm; D = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFCEFD8FFBA147CDFBA4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760638/files/figure.png	https://doi.org/10.5281/zenodo.18760638	Fig. 4. Habitat of Acipes alicantinus sp. nov. Gilgado & Enghoff. A. Scree in Sierra de Mariola, Alicante. B. Scree in Sierra de Aitana, Alicante.	Fig. 4. Habitat of Acipes alicantinus sp. nov. Gilgado & Enghoff. A. Scree in Sierra de Mariola, Alicante. B. Scree in Sierra de Aitana, Alicante.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760656/files/figure.png	https://doi.org/10.5281/zenodo.18760656	Fig. 11. Outline drawing of the gonopods of the cave species from Algarve, Portugal. Redrawn from Enghoff & Reboleira (2013). A–B. Acipes bifilum Enghoff & Reboleira, 2013. A. Anterior gonopods. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods. C–D. Acipes machadoi Enghoff & Reboleira, 2013. C. Posterior gonopods. D. Anterior gonopods. Arrow indicatES subapical emargination of coxal process. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A, D = 0.01 mm; B–C = 0.05 mm.	Fig. 11. Outline drawing of the gonopods of the cave species from Algarve, Portugal. Redrawn from Enghoff & Reboleira (2013). A–B. Acipes bifilum Enghoff & Reboleira, 2013. A. Anterior gonopods. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods. C–D. Acipes machadoi Enghoff & Reboleira, 2013. C. Posterior gonopods. D. Anterior gonopods. Arrow indicatES subapical emargination of coxal process. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A, D = 0.01 mm; B–C = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760671/files/figure.png	https://doi.org/10.5281/zenodo.18760671	Fig. 17. Acipes oligops Gilgado & Enghoff sp. nov. A–C. Paratype male. MNCN 20.07/2167. D–E. Holotype male. NHMD 1184664. A. Head, antenna, and first two legs in right lateral view). B. First pair of legs in posterior view. C. Anterior gonopods in posterior view. Arrow indicates subapical emargination of coxal process. D. A posterior gonopod in right lateral view, inset: distalmost part enlarged. E. A pregonopodal leg (arrows point at lanceolate setae). Abbreviations: af = apical flange, apo = tibial apophysis; cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A = 0.5 mm; B, D–E = 0.05 mm; C = 0.1 mm.	Fig. 17. Acipes oligops Gilgado & Enghoff sp. nov. A–C. Paratype male. MNCN 20.07/2167. D–E. Holotype male. NHMD 1184664. A. Head, antenna, and first two legs in right lateral view). B. First pair of legs in posterior view. C. Anterior gonopods in posterior view. Arrow indicates subapical emargination of coxal process. D. A posterior gonopod in right lateral view, inset: distalmost part enlarged. E. A pregonopodal leg (arrows point at lanceolate setae). Abbreviations: af = apical flange, apo = tibial apophysis; cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A = 0.5 mm; B, D–E = 0.05 mm; C = 0.1 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760668/files/figure.png	https://doi.org/10.5281/zenodo.18760668	Fig. 16. Gonopods and first leg pair of male of Acipes continentalis Enghoff, 1986. A. Distal part of posterior gonopod of A. continentalis from Sierra de Gredos, Ávila, redrawn from Enghoff (1986). B. Tip of posterior gonopod of A. continentalis from Sierra de Guadarrama, Madrid, redrawn from Enghoff and Mauriès (1999). C. Anterior gonopods of A. continentalis from Rascafría, Sierra de Guadarrama in posterior view. UAH. Arrow indicates subapical emargination of coxal process. D. Posterior gonopods of A. continentalis from Sierra de Guadarrama, from same specimen, in right lateral view. Note that the gonopods were bent during manipulation for taking the photographs. E. Tip of posterior gonopod in dorsolateral view of specimen from Guadarrama, different specimen. NHMD. F. Modified first pair of legs of male in anterior view, same specimen as E. Abbreviations: af = apical flange, apo = tibial apophysis, cxp = coxal process, ft = filamentous tip, ta = tarsus remnant, tp = telopodite. Scale bars: A, C–D = 0.1 mm; B, F = 0.05 mm; E = 0.02 mm.	Fig. 16. Gonopods and first leg pair of male of Acipes continentalis Enghoff, 1986. A. Distal part of posterior gonopod of A. continentalis from Sierra de Gredos, Ávila, redrawn from Enghoff (1986). B. Tip of posterior gonopod of A. continentalis from Sierra de Guadarrama, Madrid, redrawn from Enghoff and Mauriès (1999). C. Anterior gonopods of A. continentalis from Rascafría, Sierra de Guadarrama in posterior view. UAH. Arrow indicates subapical emargination of coxal process. D. Posterior gonopods of A. continentalis from Sierra de Guadarrama, from same specimen, in right lateral view. Note that the gonopods were bent during manipulation for taking the photographs. E. Tip of posterior gonopod in dorsolateral view of specimen from Guadarrama, different specimen. NHMD. F. Modified first pair of legs of male in anterior view, same specimen as E. Abbreviations: af = apical flange, apo = tibial apophysis, cxp = coxal process, ft = filamentous tip, ta = tarsus remnant, tp = telopodite. Scale bars: A, C–D = 0.1 mm; B, F = 0.05 mm; E = 0.02 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760660/files/figure.png	https://doi.org/10.5281/zenodo.18760660	Fig. 13. Details of the legs, trunk and telson of a male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos. A. First pair of legs in posterior view. Same specimen as in Fig. 12D. B. Detail of postgonopodal legs and lanceolate setae of same specimen in anterior view. C. Trunk of gonopodal and surrounding rings in lateral view. Same specimen as in Fig. 12C. D. Postgonopodal legs in anterior view. Same specimen as in Fig. 12D. E. Posterior-most body rings of trunk and telson in left lateral view. Same specimen as in Fig. 12C. Abbreviations: apo = tibial apophysis, ta = tarsus remnant. Scale bars: A–B = 0.05 mm; C, E = 0.02 mm; D = 0.1 mm.	Fig. 13. Details of the legs, trunk and telson of a male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos. A. First pair of legs in posterior view. Same specimen as in Fig. 12D. B. Detail of postgonopodal legs and lanceolate setae of same specimen in anterior view. C. Trunk of gonopodal and surrounding rings in lateral view. Same specimen as in Fig. 12C. D. Postgonopodal legs in anterior view. Same specimen as in Fig. 12D. E. Posterior-most body rings of trunk and telson in left lateral view. Same specimen as in Fig. 12C. Abbreviations: apo = tibial apophysis, ta = tarsus remnant. Scale bars: A–B = 0.05 mm; C, E = 0.02 mm; D = 0.1 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760664/files/figure.png	https://doi.org/10.5281/zenodo.18760664	Fig. 14. Gonopods of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos. A. Anterior gonopods of holotype in posterior view. MNCN 20.07/2106. Arrow indicates subapical emargination of coxal process.B. Anterior gonopods of specimen from same locality and capture as holotype in posterior view. UAH. Arrow indicates subapical emargination of coxal process. C. Anterior gonopods of another specimen from same locality and capture as holotype left lateral view. UAH. D. Posterior gonopods of holotype in right lateral view. MNCN 20.07/2106. E. Posterior gonopods of another specimen from same locality and capture as holotype left lateral view. UAH. F. Detail of the distalmost part of same gonopods as E. Abbreviations: af = apical flange; cxp = coxal process; ft = filamentous tip; tp = telopodite. Scale bars: A–E = 0.1 mm; F = 0.05 mm.	Fig. 14. Gonopods of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos. A. Anterior gonopods of holotype in posterior view. MNCN 20.07/2106. Arrow indicates subapical emargination of coxal process.B. Anterior gonopods of specimen from same locality and capture as holotype in posterior view. UAH. Arrow indicates subapical emargination of coxal process. C. Anterior gonopods of another specimen from same locality and capture as holotype left lateral view. UAH. D. Posterior gonopods of holotype in right lateral view. MNCN 20.07/2106. E. Posterior gonopods of another specimen from same locality and capture as holotype left lateral view. UAH. F. Detail of the distalmost part of same gonopods as E. Abbreviations: af = apical flange; cxp = coxal process; ft = filamentous tip; tp = telopodite. Scale bars: A–E = 0.1 mm; F = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760666/files/figure.png	https://doi.org/10.5281/zenodo.18760666	Fig. 15. Gonopods of Acipes celtibericus Gilgado & Enghoff sp. nov. From Moncayo Massif, Zaragoza. UAH. A. Anterior gonopods in posterior view. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods in right lateral view. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bar = 0.1 mm.	Fig. 15. Gonopods of Acipes celtibericus Gilgado & Enghoff sp. nov. From Moncayo Massif, Zaragoza. UAH. A. Anterior gonopods in posterior view. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods in right lateral view. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bar = 0.1 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760648/files/figure.png	https://doi.org/10.5281/zenodo.18760648	Fig. 8. Anatomical details of Acipes alicantinus sp. nov. Gilgado & Enghoff. A. Head and first five rings of male specimen from type locality in left lateral view; 30 Nov. 2013; UAH. B. Head of another male specimen from same locality and date in ventral view; UAH. C. Tip of antenna of same specimen as A in dorsal view. D. Rings 6–9 of same specimen as A. E. Telson and last body rings of same specimen in left lateral view. F. First male legs of same specimen as B in posterior view. G. Leg 4 of same specimen as B. H. last leg of same specimen as B. Abbreviations: apo = tibial apophysis; ta = tarsus remnant. Scale bars: A, D–E = 0.2 mm; B–C, G = 0.1 mm; F–H = 0.05 mm.	Fig. 8. Anatomical details of Acipes alicantinus sp. nov. Gilgado & Enghoff. A. Head and first five rings of male specimen from type locality in left lateral view; 30 Nov. 2013; UAH. B. Head of another male specimen from same locality and date in ventral view; UAH. C. Tip of antenna of same specimen as A in dorsal view. D. Rings 6–9 of same specimen as A. E. Telson and last body rings of same specimen in left lateral view. F. First male legs of same specimen as B in posterior view. G. Leg 4 of same specimen as B. H. last leg of same specimen as B. Abbreviations: apo = tibial apophysis; ta = tarsus remnant. Scale bars: A, D–E = 0.2 mm; B–C, G = 0.1 mm; F–H = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760654/files/figure.png	https://doi.org/10.5281/zenodo.18760654	Fig. 10. Gonopods and first pair of male legs of Acipes andalusius Enghoff & Mauriès, 1999. A. Anterior gonopods in posterior view of specimen from PB-2, Jaén, redrawn from Enghoff & Mauriès (1999). Arrow indicating subapical emargination of coxal process. B. Anterior gonopods in posterior view of male specimen from PB-4, Jaén. UAH. Arrow indicating subapical emargination of coxal process. C. Distal part of posterior gonopods of specimen from PB-2, redrawn from Enghoff & Mauriès (1999). D. Tip of posterior gonopod of specimen from PB-4. UAH. E. Posterior gonopods in left lateral view of male specimen from PB-4. UAH. F. First pair of male legs of male specimen from PB-4. UAH. Abbreviations: af = apical flange; apo = tibial apophysis; cxp = coxal process; ft = filamentous tip; ta = tarsus remnant; tp = telopodite. Scale bars: A–C, E–F = 0.1 mm; D = 0.05 mm.	Fig. 10. Gonopods and first pair of male legs of Acipes andalusius Enghoff & Mauriès, 1999. A. Anterior gonopods in posterior view of specimen from PB-2, Jaén, redrawn from Enghoff & Mauriès (1999). Arrow indicating subapical emargination of coxal process. B. Anterior gonopods in posterior view of male specimen from PB-4, Jaén. UAH. Arrow indicating subapical emargination of coxal process. C. Distal part of posterior gonopods of specimen from PB-2, redrawn from Enghoff & Mauriès (1999). D. Tip of posterior gonopod of specimen from PB-4. UAH. E. Posterior gonopods in left lateral view of male specimen from PB-4. UAH. F. First pair of male legs of male specimen from PB-4. UAH. Abbreviations: af = apical flange; apo = tibial apophysis; cxp = coxal process; ft = filamentous tip; ta = tarsus remnant; tp = telopodite. Scale bars: A–C, E–F = 0.1 mm; D = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760650/files/figure.png	https://doi.org/10.5281/zenodo.18760650	Fig. 9. Gonopods of Acipes alicantinus Gilgado & Enghoff sp. nov. A. Anterior gonopods of holotype in posterior view. MNCN 20.07/2169. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods of holotype in right lateral view. MNCN 20.07/2169. C. Anterior gonopods of male specimen from type locality (30 Nov. 2013) in posterior view. Arrow indicates subapical emargination of coxal process. D. Detail of flange and filamentous tip of posterior gonopods of same specimen as C in left lateral view. Abbreviations: af = apical flange; cxp = coxal process; ft = filamentous tip; tp = telopodite. Scale bars: A–C = 0.1 mm; D = 0.05 mm.	Fig. 9. Gonopods of Acipes alicantinus Gilgado & Enghoff sp. nov. A. Anterior gonopods of holotype in posterior view. MNCN 20.07/2169. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods of holotype in right lateral view. MNCN 20.07/2169. C. Anterior gonopods of male specimen from type locality (30 Nov. 2013) in posterior view. Arrow indicates subapical emargination of coxal process. D. Detail of flange and filamentous tip of posterior gonopods of same specimen as C in left lateral view. Abbreviations: af = apical flange; cxp = coxal process; ft = filamentous tip; tp = telopodite. Scale bars: A–C = 0.1 mm; D = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760646/files/figure.png	https://doi.org/10.5281/zenodo.18760646	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760634/files/figure.png	https://doi.org/10.5281/zenodo.18760634	Fig. 2. Sampling technique and habitat in alluvial MSS in Arroyo de Valdelapuerta, Tamajón, Guadalajara, type locality of Acipes oligops Gilgado & Enghoff sp. nov. A. Removal of Subterranean Sampling Device (SSD) lid. B. Retrieval of pitfall trap from the SSD. C–D. Removal of the SSD tube (0.5 m) at the end of sampling period. E. Structure of the debris and voids in the alluvial MSS as seen after removal of the tube (microhabitat of Acipes oligops sp. nov.). F. Temporal watercourse (Arroyo de Valdelapuerta) and surrounding vegetation (habitat of Acipes oligops).	Fig. 2. Sampling technique and habitat in alluvial MSS in Arroyo de Valdelapuerta, Tamajón, Guadalajara, type locality of Acipes oligops Gilgado & Enghoff sp. nov. A. Removal of Subterranean Sampling Device (SSD) lid. B. Retrieval of pitfall trap from the SSD. C–D. Removal of the SSD tube (0.5 m) at the end of sampling period. E. Structure of the debris and voids in the alluvial MSS as seen after removal of the tube (microhabitat of Acipes oligops sp. nov.). F. Temporal watercourse (Arroyo de Valdelapuerta) and surrounding vegetation (habitat of Acipes oligops).	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760632/files/figure.png	https://doi.org/10.5281/zenodo.18760632	Fig. 1. Sampling technique and landscape in the Iberian System, habitat of Acipes celtibericus Gilgado & Enghoff sp. nov. A. Subterranean Sampling Device (SSD) used to sample in the MSS in Sierra de la Demanda before being covered. B. Glacier circus in Mencilla, Sierra de la Demanda, Burgos. C. Beech forest in Mencilla, Sierra de la Demanda. The pile of stones marks the position of the SSD. D. Scree in the Moncayo massif, Zaragoza.	Fig. 1. Sampling technique and landscape in the Iberian System, habitat of Acipes celtibericus Gilgado & Enghoff sp. nov. A. Subterranean Sampling Device (SSD) used to sample in the MSS in Sierra de la Demanda before being covered. B. Glacier circus in Mencilla, Sierra de la Demanda, Burgos. C. Beech forest in Mencilla, Sierra de la Demanda. The pile of stones marks the position of the SSD. D. Scree in the Moncayo massif, Zaragoza.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B77DFFC5FDFCFD7041CFFBDF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760636/files/figure.png	https://doi.org/10.5281/zenodo.18760636	Fig. 3. Habitat of Acipes continentalis Enghoff, 1986. A. Falda del Cerro de la Muela, Sierra de Guadarrama, Segovia, 1605 m. B. Landscape in the highest sampling point, scree between Cabeza de Hierro Mayor and Menor (peak at the right), Sierra de Guadarrama, Madrid. C. Landscape of La Maliciosa, Sierra de Guadarrama, Madrid. D. Landscape in La Cabrera, Sierra de Guadarrama, Madrid.	Fig. 3. Habitat of Acipes continentalis Enghoff, 1986. A. Falda del Cerro de la Muela, Sierra de Guadarrama, Segovia, 1605 m. B. Landscape in the highest sampling point, scree between Cabeza de Hierro Mayor and Menor (peak at the right), Sierra de Guadarrama, Madrid. C. Landscape of La Maliciosa, Sierra de Guadarrama, Madrid. D. Landscape in La Cabrera, Sierra de Guadarrama, Madrid.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B776FFCEFD9AFB8F41FFF858.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760646/files/figure.png	https://doi.org/10.5281/zenodo.18760646	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B776FFCEFD9AFB8F41FFF858.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760654/files/figure.png	https://doi.org/10.5281/zenodo.18760654	Fig. 10. Gonopods and first pair of male legs of Acipes andalusius Enghoff & Mauriès, 1999. A. Anterior gonopods in posterior view of specimen from PB-2, Jaén, redrawn from Enghoff & Mauriès (1999). Arrow indicating subapical emargination of coxal process. B. Anterior gonopods in posterior view of male specimen from PB-4, Jaén. UAH. Arrow indicating subapical emargination of coxal process. C. Distal part of posterior gonopods of specimen from PB-2, redrawn from Enghoff & Mauriès (1999). D. Tip of posterior gonopod of specimen from PB-4. UAH. E. Posterior gonopods in left lateral view of male specimen from PB-4. UAH. F. First pair of male legs of male specimen from PB-4. UAH. Abbreviations: af = apical flange; apo = tibial apophysis; cxp = coxal process; ft = filamentous tip; ta = tarsus remnant; tp = telopodite. Scale bars: A–C, E–F = 0.1 mm; D = 0.05 mm.	Fig. 10. Gonopods and first pair of male legs of Acipes andalusius Enghoff & Mauriès, 1999. A. Anterior gonopods in posterior view of specimen from PB-2, Jaén, redrawn from Enghoff & Mauriès (1999). Arrow indicating subapical emargination of coxal process. B. Anterior gonopods in posterior view of male specimen from PB-4, Jaén. UAH. Arrow indicating subapical emargination of coxal process. C. Distal part of posterior gonopods of specimen from PB-2, redrawn from Enghoff & Mauriès (1999). D. Tip of posterior gonopod of specimen from PB-4. UAH. E. Posterior gonopods in left lateral view of male specimen from PB-4. UAH. F. First pair of male legs of male specimen from PB-4. UAH. Abbreviations: af = apical flange; apo = tibial apophysis; cxp = coxal process; ft = filamentous tip; ta = tarsus remnant; tp = telopodite. Scale bars: A–C, E–F = 0.1 mm; D = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B777FFCCFD95FED34616FD8F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760656/files/figure.png	https://doi.org/10.5281/zenodo.18760656	Fig. 11. Outline drawing of the gonopods of the cave species from Algarve, Portugal. Redrawn from Enghoff & Reboleira (2013). A–B. Acipes bifilum Enghoff & Reboleira, 2013. A. Anterior gonopods. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods. C–D. Acipes machadoi Enghoff & Reboleira, 2013. C. Posterior gonopods. D. Anterior gonopods. Arrow indicatES subapical emargination of coxal process. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A, D = 0.01 mm; B–C = 0.05 mm.	Fig. 11. Outline drawing of the gonopods of the cave species from Algarve, Portugal. Redrawn from Enghoff & Reboleira (2013). A–B. Acipes bifilum Enghoff & Reboleira, 2013. A. Anterior gonopods. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods. C–D. Acipes machadoi Enghoff & Reboleira, 2013. C. Posterior gonopods. D. Anterior gonopods. Arrow indicatES subapical emargination of coxal process. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A, D = 0.01 mm; B–C = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760646/files/figure.png	https://doi.org/10.5281/zenodo.18760646	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760658/files/figure.png	https://doi.org/10.5281/zenodo.18760658	Fig. 12. Head and first rings of male specimens of Acipes celtibericus sp. nov. Gilgado & Enghoff from Mencilla, Sierra de la Demanda, Burgos. A. Specimen from same locality and capture as holotype, in alcohol. Right lateral view. UAH. B. Same specimen, dry, in left lateral view. UAH. C. Specimen from same locality and capture as holotype. Head and first rings in left lateral view. UAH. D. Another specimen from same locality and capture as holotype. Head in ventral view. UAH. E. Another specimen from same locality and capture as holotype. Head in dorsofrontal view. UAH. F. Detail of labrum and prelabral setae of same specimen as in E. Scale bars: A–B, D = 0.5 mm; C, E = 0.2 mm; F = 0.1 mm.	Fig. 12. Head and first rings of male specimens of Acipes celtibericus sp. nov. Gilgado & Enghoff from Mencilla, Sierra de la Demanda, Burgos. A. Specimen from same locality and capture as holotype, in alcohol. Right lateral view. UAH. B. Same specimen, dry, in left lateral view. UAH. C. Specimen from same locality and capture as holotype. Head and first rings in left lateral view. UAH. D. Another specimen from same locality and capture as holotype. Head in ventral view. UAH. E. Another specimen from same locality and capture as holotype. Head in dorsofrontal view. UAH. F. Detail of labrum and prelabral setae of same specimen as in E. Scale bars: A–B, D = 0.5 mm; C, E = 0.2 mm; F = 0.1 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760660/files/figure.png	https://doi.org/10.5281/zenodo.18760660	Fig. 13. Details of the legs, trunk and telson of a male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos. A. First pair of legs in posterior view. Same specimen as in Fig. 12D. B. Detail of postgonopodal legs and lanceolate setae of same specimen in anterior view. C. Trunk of gonopodal and surrounding rings in lateral view. Same specimen as in Fig. 12C. D. Postgonopodal legs in anterior view. Same specimen as in Fig. 12D. E. Posterior-most body rings of trunk and telson in left lateral view. Same specimen as in Fig. 12C. Abbreviations: apo = tibial apophysis, ta = tarsus remnant. Scale bars: A–B = 0.05 mm; C, E = 0.02 mm; D = 0.1 mm.	Fig. 13. Details of the legs, trunk and telson of a male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos. A. First pair of legs in posterior view. Same specimen as in Fig. 12D. B. Detail of postgonopodal legs and lanceolate setae of same specimen in anterior view. C. Trunk of gonopodal and surrounding rings in lateral view. Same specimen as in Fig. 12C. D. Postgonopodal legs in anterior view. Same specimen as in Fig. 12D. E. Posterior-most body rings of trunk and telson in left lateral view. Same specimen as in Fig. 12C. Abbreviations: apo = tibial apophysis, ta = tarsus remnant. Scale bars: A–B = 0.05 mm; C, E = 0.02 mm; D = 0.1 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760664/files/figure.png	https://doi.org/10.5281/zenodo.18760664	Fig. 14. Gonopods of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos. A. Anterior gonopods of holotype in posterior view. MNCN 20.07/2106. Arrow indicates subapical emargination of coxal process.B. Anterior gonopods of specimen from same locality and capture as holotype in posterior view. UAH. Arrow indicates subapical emargination of coxal process. C. Anterior gonopods of another specimen from same locality and capture as holotype left lateral view. UAH. D. Posterior gonopods of holotype in right lateral view. MNCN 20.07/2106. E. Posterior gonopods of another specimen from same locality and capture as holotype left lateral view. UAH. F. Detail of the distalmost part of same gonopods as E. Abbreviations: af = apical flange; cxp = coxal process; ft = filamentous tip; tp = telopodite. Scale bars: A–E = 0.1 mm; F = 0.05 mm.	Fig. 14. Gonopods of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos. A. Anterior gonopods of holotype in posterior view. MNCN 20.07/2106. Arrow indicates subapical emargination of coxal process.B. Anterior gonopods of specimen from same locality and capture as holotype in posterior view. UAH. Arrow indicates subapical emargination of coxal process. C. Anterior gonopods of another specimen from same locality and capture as holotype left lateral view. UAH. D. Posterior gonopods of holotype in right lateral view. MNCN 20.07/2106. E. Posterior gonopods of another specimen from same locality and capture as holotype left lateral view. UAH. F. Detail of the distalmost part of same gonopods as E. Abbreviations: af = apical flange; cxp = coxal process; ft = filamentous tip; tp = telopodite. Scale bars: A–E = 0.1 mm; F = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760666/files/figure.png	https://doi.org/10.5281/zenodo.18760666	Fig. 15. Gonopods of Acipes celtibericus Gilgado & Enghoff sp. nov. From Moncayo Massif, Zaragoza. UAH. A. Anterior gonopods in posterior view. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods in right lateral view. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bar = 0.1 mm.	Fig. 15. Gonopods of Acipes celtibericus Gilgado & Enghoff sp. nov. From Moncayo Massif, Zaragoza. UAH. A. Anterior gonopods in posterior view. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods in right lateral view. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bar = 0.1 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760644/files/figure.png	https://doi.org/10.5281/zenodo.18760644	Fig. 6. Distribution of the species of Acipes in 10 × 10 km UTM squares in the Iberian Peninsula. Main mountain systems with Acipes Attems, 1937 records are labelled. Blue squares: Acipes celtibericus Gilgado & Enghoff sp. nov. Purple square: Acipes oligops sp. nov. Gilgado & Enghoff. Dark green squares: previous records of Acipes continentalis Enghoff, 1986. Light green squares: new records of Acipes continentalis. Dark red squares: Acipes andalusius Enghoff & Mauriès, 1999. Light pink square: new record of Acipes alicantinus sp. nov Gilgado & Enghoff. Dark pink square: previous record of Acipes alicantinus sp. nov.(as A. andalusius in the literature). Orange square: Acipes machadoi Enghoff & Reboleira, 2013. Yellow square: Acipes bifilum Enghoff & Reboleira, 2013. Scale bar = 200 km.	Fig. 6. Distribution of the species of Acipes in 10 × 10 km UTM squares in the Iberian Peninsula. Main mountain systems with Acipes Attems, 1937 records are labelled. Blue squares: Acipes celtibericus Gilgado & Enghoff sp. nov. Purple square: Acipes oligops sp. nov. Gilgado & Enghoff. Dark green squares: previous records of Acipes continentalis Enghoff, 1986. Light green squares: new records of Acipes continentalis. Dark red squares: Acipes andalusius Enghoff & Mauriès, 1999. Light pink square: new record of Acipes alicantinus sp. nov Gilgado & Enghoff. Dark pink square: previous record of Acipes alicantinus sp. nov.(as A. andalusius in the literature). Orange square: Acipes machadoi Enghoff & Reboleira, 2013. Yellow square: Acipes bifilum Enghoff & Reboleira, 2013. Scale bar = 200 km.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B774FFD1FD8AFDF6467FFCB7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760668/files/figure.png	https://doi.org/10.5281/zenodo.18760668	Fig. 16. Gonopods and first leg pair of male of Acipes continentalis Enghoff, 1986. A. Distal part of posterior gonopod of A. continentalis from Sierra de Gredos, Ávila, redrawn from Enghoff (1986). B. Tip of posterior gonopod of A. continentalis from Sierra de Guadarrama, Madrid, redrawn from Enghoff and Mauriès (1999). C. Anterior gonopods of A. continentalis from Rascafría, Sierra de Guadarrama in posterior view. UAH. Arrow indicates subapical emargination of coxal process. D. Posterior gonopods of A. continentalis from Sierra de Guadarrama, from same specimen, in right lateral view. Note that the gonopods were bent during manipulation for taking the photographs. E. Tip of posterior gonopod in dorsolateral view of specimen from Guadarrama, different specimen. NHMD. F. Modified first pair of legs of male in anterior view, same specimen as E. Abbreviations: af = apical flange, apo = tibial apophysis, cxp = coxal process, ft = filamentous tip, ta = tarsus remnant, tp = telopodite. Scale bars: A, C–D = 0.1 mm; B, F = 0.05 mm; E = 0.02 mm.	Fig. 16. Gonopods and first leg pair of male of Acipes continentalis Enghoff, 1986. A. Distal part of posterior gonopod of A. continentalis from Sierra de Gredos, Ávila, redrawn from Enghoff (1986). B. Tip of posterior gonopod of A. continentalis from Sierra de Guadarrama, Madrid, redrawn from Enghoff and Mauriès (1999). C. Anterior gonopods of A. continentalis from Rascafría, Sierra de Guadarrama in posterior view. UAH. Arrow indicates subapical emargination of coxal process. D. Posterior gonopods of A. continentalis from Sierra de Guadarrama, from same specimen, in right lateral view. Note that the gonopods were bent during manipulation for taking the photographs. E. Tip of posterior gonopod in dorsolateral view of specimen from Guadarrama, different specimen. NHMD. F. Modified first pair of legs of male in anterior view, same specimen as E. Abbreviations: af = apical flange, apo = tibial apophysis, cxp = coxal process, ft = filamentous tip, ta = tarsus remnant, tp = telopodite. Scale bars: A, C–D = 0.1 mm; B, F = 0.05 mm; E = 0.02 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B76EFFD7FDC3FED3472DFDCB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760646/files/figure.png	https://doi.org/10.5281/zenodo.18760646	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B76EFFD7FDC3FED3472DFDCB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760668/files/figure.png	https://doi.org/10.5281/zenodo.18760668	Fig. 16. Gonopods and first leg pair of male of Acipes continentalis Enghoff, 1986. A. Distal part of posterior gonopod of A. continentalis from Sierra de Gredos, Ávila, redrawn from Enghoff (1986). B. Tip of posterior gonopod of A. continentalis from Sierra de Guadarrama, Madrid, redrawn from Enghoff and Mauriès (1999). C. Anterior gonopods of A. continentalis from Rascafría, Sierra de Guadarrama in posterior view. UAH. Arrow indicates subapical emargination of coxal process. D. Posterior gonopods of A. continentalis from Sierra de Guadarrama, from same specimen, in right lateral view. Note that the gonopods were bent during manipulation for taking the photographs. E. Tip of posterior gonopod in dorsolateral view of specimen from Guadarrama, different specimen. NHMD. F. Modified first pair of legs of male in anterior view, same specimen as E. Abbreviations: af = apical flange, apo = tibial apophysis, cxp = coxal process, ft = filamentous tip, ta = tarsus remnant, tp = telopodite. Scale bars: A, C–D = 0.1 mm; B, F = 0.05 mm; E = 0.02 mm.	Fig. 16. Gonopods and first leg pair of male of Acipes continentalis Enghoff, 1986. A. Distal part of posterior gonopod of A. continentalis from Sierra de Gredos, Ávila, redrawn from Enghoff (1986). B. Tip of posterior gonopod of A. continentalis from Sierra de Guadarrama, Madrid, redrawn from Enghoff and Mauriès (1999). C. Anterior gonopods of A. continentalis from Rascafría, Sierra de Guadarrama in posterior view. UAH. Arrow indicates subapical emargination of coxal process. D. Posterior gonopods of A. continentalis from Sierra de Guadarrama, from same specimen, in right lateral view. Note that the gonopods were bent during manipulation for taking the photographs. E. Tip of posterior gonopod in dorsolateral view of specimen from Guadarrama, different specimen. NHMD. F. Modified first pair of legs of male in anterior view, same specimen as E. Abbreviations: af = apical flange, apo = tibial apophysis, cxp = coxal process, ft = filamentous tip, ta = tarsus remnant, tp = telopodite. Scale bars: A, C–D = 0.1 mm; B, F = 0.05 mm; E = 0.02 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B76EFFD7FDC3FED3472DFDCB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760644/files/figure.png	https://doi.org/10.5281/zenodo.18760644	Fig. 6. Distribution of the species of Acipes in 10 × 10 km UTM squares in the Iberian Peninsula. Main mountain systems with Acipes Attems, 1937 records are labelled. Blue squares: Acipes celtibericus Gilgado & Enghoff sp. nov. Purple square: Acipes oligops sp. nov. Gilgado & Enghoff. Dark green squares: previous records of Acipes continentalis Enghoff, 1986. Light green squares: new records of Acipes continentalis. Dark red squares: Acipes andalusius Enghoff & Mauriès, 1999. Light pink square: new record of Acipes alicantinus sp. nov Gilgado & Enghoff. Dark pink square: previous record of Acipes alicantinus sp. nov.(as A. andalusius in the literature). Orange square: Acipes machadoi Enghoff & Reboleira, 2013. Yellow square: Acipes bifilum Enghoff & Reboleira, 2013. Scale bar = 200 km.	Fig. 6. Distribution of the species of Acipes in 10 × 10 km UTM squares in the Iberian Peninsula. Main mountain systems with Acipes Attems, 1937 records are labelled. Blue squares: Acipes celtibericus Gilgado & Enghoff sp. nov. Purple square: Acipes oligops sp. nov. Gilgado & Enghoff. Dark green squares: previous records of Acipes continentalis Enghoff, 1986. Light green squares: new records of Acipes continentalis. Dark red squares: Acipes andalusius Enghoff & Mauriès, 1999. Light pink square: new record of Acipes alicantinus sp. nov Gilgado & Enghoff. Dark pink square: previous record of Acipes alicantinus sp. nov.(as A. andalusius in the literature). Orange square: Acipes machadoi Enghoff & Reboleira, 2013. Yellow square: Acipes bifilum Enghoff & Reboleira, 2013. Scale bar = 200 km.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B76CFFD4FD84FED3407AFDC6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760656/files/figure.png	https://doi.org/10.5281/zenodo.18760656	Fig. 11. Outline drawing of the gonopods of the cave species from Algarve, Portugal. Redrawn from Enghoff & Reboleira (2013). A–B. Acipes bifilum Enghoff & Reboleira, 2013. A. Anterior gonopods. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods. C–D. Acipes machadoi Enghoff & Reboleira, 2013. C. Posterior gonopods. D. Anterior gonopods. Arrow indicatES subapical emargination of coxal process. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A, D = 0.01 mm; B–C = 0.05 mm.	Fig. 11. Outline drawing of the gonopods of the cave species from Algarve, Portugal. Redrawn from Enghoff & Reboleira (2013). A–B. Acipes bifilum Enghoff & Reboleira, 2013. A. Anterior gonopods. Arrow indicates subapical emargination of coxal process. B. Posterior gonopods. C–D. Acipes machadoi Enghoff & Reboleira, 2013. C. Posterior gonopods. D. Anterior gonopods. Arrow indicatES subapical emargination of coxal process. Abbreviations: af = apical flange, cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A, D = 0.01 mm; B–C = 0.05 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B76CFFDBFD97FDAE44EFFDAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760646/files/figure.png	https://doi.org/10.5281/zenodo.18760646	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	Fig. 7. Habitus of Acipes Attems, 1937 species studied in the present work. A. Living specimen of Acipes andalusius Enghoff & Mauriès, 1999 from cave PB-4. Photograph by Toni Pérez (GEV). B. Dissected male specimen of Acipes andalusius from PB-4 cave; UAH. C. Male specimen of Acipes alicantinus Gilgado & Enghoff sp. nov. from Aitana mountain; MNCN 20.07/2169. D. Male specimen of Acipes celtibericus Gilgado & Enghoff sp. nov. from Mencilla, Sierra de la Demanda, Burgos; MNCN 20.07/2106. E. Male specimen of Acipes continentalis Enghoff, 1986 from Rascafría, Sierra de Guadarrama, Madrid. UAH. F. Male paratype of Acipes oligops sp. nov. Gilgado & Enghoff, MNCN 20.07/2167. Scale bar = 2 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
03CF87D4B76CFFDBFD97FDAE44EFFDAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18760671/files/figure.png	https://doi.org/10.5281/zenodo.18760671	Fig. 17. Acipes oligops Gilgado & Enghoff sp. nov. A–C. Paratype male. MNCN 20.07/2167. D–E. Holotype male. NHMD 1184664. A. Head, antenna, and first two legs in right lateral view). B. First pair of legs in posterior view. C. Anterior gonopods in posterior view. Arrow indicates subapical emargination of coxal process. D. A posterior gonopod in right lateral view, inset: distalmost part enlarged. E. A pregonopodal leg (arrows point at lanceolate setae). Abbreviations: af = apical flange, apo = tibial apophysis; cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A = 0.5 mm; B, D–E = 0.05 mm; C = 0.1 mm.	Fig. 17. Acipes oligops Gilgado & Enghoff sp. nov. A–C. Paratype male. MNCN 20.07/2167. D–E. Holotype male. NHMD 1184664. A. Head, antenna, and first two legs in right lateral view). B. First pair of legs in posterior view. C. Anterior gonopods in posterior view. Arrow indicates subapical emargination of coxal process. D. A posterior gonopod in right lateral view, inset: distalmost part enlarged. E. A pregonopodal leg (arrows point at lanceolate setae). Abbreviations: af = apical flange, apo = tibial apophysis; cxp = coxal process, ft = filamentous tip, tp = telopodite. Scale bars: A = 0.5 mm; B, D–E = 0.05 mm; C = 0.1 mm.	2026-02-12	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.		Zenodo	biologists	Gilgado, José D.;Enghoff, Henrik;Ortuño, Vicente M.			
