taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03CECD466B515D4BFF14FBD91CCB24D6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608296/files/figure.png	https://doi.org/10.5281/zenodo.16608296	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B515D4BFF14FBD91CCB24D6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608306/files/figure.png	https://doi.org/10.5281/zenodo.16608306	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B515D4BFF14FBD91CCB24D6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608310/files/figure.png	https://doi.org/10.5281/zenodo.16608310	FIGURE 7. A: Forewing veination diagram of Borysthenes species, labeling longitudinal veins in regular, veinlets in italic, and C3 cell in bold; nodal line: an imaginary transverse line at the level of 1st r-m, 1st m-cu and icua; B: abdomen segments of Borysthenes xuzhiae sp. nov., ventral view, labeling sternites VI, VII, VIII, and genital segment (IX).	FIGURE 7. A: Forewing veination diagram of Borysthenes species, labeling longitudinal veins in regular, veinlets in italic, and C3 cell in bold; nodal line: an imaginary transverse line at the level of 1st r-m, 1st m-cu and icua; B: abdomen segments of Borysthenes xuzhiae sp. nov., ventral view, labeling sternites VI, VII, VIII, and genital segment (IX).	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B515D4BFF14FBD91CCB24D6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608298/files/figure.png	https://doi.org/10.5281/zenodo.16608298	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D48FF14FF2D18F52510.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608296/files/figure.png	https://doi.org/10.5281/zenodo.16608296	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D48FF14FF2D18F52510.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608306/files/figure.png	https://doi.org/10.5281/zenodo.16608306	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D48FF14FF2D18F52510.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608300/files/figure.png	https://doi.org/10.5281/zenodo.16608300	FIGURE 3. Male genitalia of Borysthenes spp. A–H: B. pantherinus sp. nov., I–P: B. opacus sp. nov., Q–X: B. undulatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	FIGURE 3. Male genitalia of Borysthenes spp. A–H: B. pantherinus sp. nov., I–P: B. opacus sp. nov., Q–X: B. undulatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D48FF14FF2D18F52510.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608298/files/figure.png	https://doi.org/10.5281/zenodo.16608298	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D48FF14FF2D18F52510.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608304/files/figure.png	https://doi.org/10.5281/zenodo.16608304	FIGURE 5. Female genitalia of Borysthenes spp. A, B: B. opacus sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C, D: B. undulatus sp. nov., (C) overview, left lateral, (D) posterior vagina, ventral. Scale bar: 0.5mm.	FIGURE 5. Female genitalia of Borysthenes spp. A, B: B. opacus sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C, D: B. undulatus sp. nov., (C) overview, left lateral, (D) posterior vagina, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D4EFF14F9D61F3425B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608296/files/figure.png	https://doi.org/10.5281/zenodo.16608296	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D4EFF14F9D61F3425B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608298/files/figure.png	https://doi.org/10.5281/zenodo.16608298	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D4EFF14F9D61F3425B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608302/files/figure.png	https://doi.org/10.5281/zenodo.16608302	FIGURE 4. Female genitalia of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C–E: B. delicatus sp. nov., (C) overview, left lateral, (D) posterior vagina, right lateral, (E) posterior vagina, ventral; F, G: B. pantherinus sp. nov., (F) overview, left lateral, (G) posterior vagina, ventral. Abbreviations: bc: bursa copulatrix; dr: ductus receptaculi; ov: ovipositor; pv: posterior vagina. Scale bar: 0.5mm.	FIGURE 4. Female genitalia of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C–E: B. delicatus sp. nov., (C) overview, left lateral, (D) posterior vagina, right lateral, (E) posterior vagina, ventral; F, G: B. pantherinus sp. nov., (F) overview, left lateral, (G) posterior vagina, ventral. Abbreviations: bc: bursa copulatrix; dr: ductus receptaculi; ov: ovipositor; pv: posterior vagina. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D4EFF14F9D61F3425B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608306/files/figure.png	https://doi.org/10.5281/zenodo.16608306	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B525D4EFF14F9D61F3425B7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608310/files/figure.png	https://doi.org/10.5281/zenodo.16608310	FIGURE 7. A: Forewing veination diagram of Borysthenes species, labeling longitudinal veins in regular, veinlets in italic, and C3 cell in bold; nodal line: an imaginary transverse line at the level of 1st r-m, 1st m-cu and icua; B: abdomen segments of Borysthenes xuzhiae sp. nov., ventral view, labeling sternites VI, VII, VIII, and genital segment (IX).	FIGURE 7. A: Forewing veination diagram of Borysthenes species, labeling longitudinal veins in regular, veinlets in italic, and C3 cell in bold; nodal line: an imaginary transverse line at the level of 1st r-m, 1st m-cu and icua; B: abdomen segments of Borysthenes xuzhiae sp. nov., ventral view, labeling sternites VI, VII, VIII, and genital segment (IX).	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B545D4CFF14F8B41E412593.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608296/files/figure.png	https://doi.org/10.5281/zenodo.16608296	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B545D4CFF14F8B41E412593.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608298/files/figure.png	https://doi.org/10.5281/zenodo.16608298	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B565D4DFF14F9501FD524FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608296/files/figure.png	https://doi.org/10.5281/zenodo.16608296	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B565D4DFF14F9501FD524FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608298/files/figure.png	https://doi.org/10.5281/zenodo.16608298	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	FIGURE 2. Male genitalia of Borysthenes spp. A–H: B. xuzhiae sp. nov., I–P: B. mysterius sp. nov., Q–X: B. delicatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B565D4DFF14F9501FD524FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608302/files/figure.png	https://doi.org/10.5281/zenodo.16608302	FIGURE 4. Female genitalia of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C–E: B. delicatus sp. nov., (C) overview, left lateral, (D) posterior vagina, right lateral, (E) posterior vagina, ventral; F, G: B. pantherinus sp. nov., (F) overview, left lateral, (G) posterior vagina, ventral. Abbreviations: bc: bursa copulatrix; dr: ductus receptaculi; ov: ovipositor; pv: posterior vagina. Scale bar: 0.5mm.	FIGURE 4. Female genitalia of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C–E: B. delicatus sp. nov., (C) overview, left lateral, (D) posterior vagina, right lateral, (E) posterior vagina, ventral; F, G: B. pantherinus sp. nov., (F) overview, left lateral, (G) posterior vagina, ventral. Abbreviations: bc: bursa copulatrix; dr: ductus receptaculi; ov: ovipositor; pv: posterior vagina. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B565D4DFF14F9501FD524FD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608306/files/figure.png	https://doi.org/10.5281/zenodo.16608306	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B585D41FF14FF2D19A123DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608296/files/figure.png	https://doi.org/10.5281/zenodo.16608296	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B585D41FF14FF2D19A123DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608300/files/figure.png	https://doi.org/10.5281/zenodo.16608300	FIGURE 3. Male genitalia of Borysthenes spp. A–H: B. pantherinus sp. nov., I–P: B. opacus sp. nov., Q–X: B. undulatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	FIGURE 3. Male genitalia of Borysthenes spp. A–H: B. pantherinus sp. nov., I–P: B. opacus sp. nov., Q–X: B. undulatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B585D41FF14FF2D19A123DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608302/files/figure.png	https://doi.org/10.5281/zenodo.16608302	FIGURE 4. Female genitalia of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C–E: B. delicatus sp. nov., (C) overview, left lateral, (D) posterior vagina, right lateral, (E) posterior vagina, ventral; F, G: B. pantherinus sp. nov., (F) overview, left lateral, (G) posterior vagina, ventral. Abbreviations: bc: bursa copulatrix; dr: ductus receptaculi; ov: ovipositor; pv: posterior vagina. Scale bar: 0.5mm.	FIGURE 4. Female genitalia of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C–E: B. delicatus sp. nov., (C) overview, left lateral, (D) posterior vagina, right lateral, (E) posterior vagina, ventral; F, G: B. pantherinus sp. nov., (F) overview, left lateral, (G) posterior vagina, ventral. Abbreviations: bc: bursa copulatrix; dr: ductus receptaculi; ov: ovipositor; pv: posterior vagina. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B585D41FF14FF2D19A123DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608306/files/figure.png	https://doi.org/10.5281/zenodo.16608306	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B5B5D46FF14FE9C184426CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608296/files/figure.png	https://doi.org/10.5281/zenodo.16608296	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B5B5D46FF14FE9C184426CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608300/files/figure.png	https://doi.org/10.5281/zenodo.16608300	FIGURE 3. Male genitalia of Borysthenes spp. A–H: B. pantherinus sp. nov., I–P: B. opacus sp. nov., Q–X: B. undulatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	FIGURE 3. Male genitalia of Borysthenes spp. A–H: B. pantherinus sp. nov., I–P: B. opacus sp. nov., Q–X: B. undulatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B5B5D46FF14FE9C184426CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608304/files/figure.png	https://doi.org/10.5281/zenodo.16608304	FIGURE 5. Female genitalia of Borysthenes spp. A, B: B. opacus sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C, D: B. undulatus sp. nov., (C) overview, left lateral, (D) posterior vagina, ventral. Scale bar: 0.5mm.	FIGURE 5. Female genitalia of Borysthenes spp. A, B: B. opacus sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C, D: B. undulatus sp. nov., (C) overview, left lateral, (D) posterior vagina, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B5B5D46FF14FE9C184426CB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608306/files/figure.png	https://doi.org/10.5281/zenodo.16608306	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B5C5D44FF14F9A81E882503.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608296/files/figure.png	https://doi.org/10.5281/zenodo.16608296	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	FIGURE 1. Habitus of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) dorsal, (B) left lateral. C, D: B. mysterius sp. nov., (C) dorsal, (D) left lateral. E, F: B. pantherinus sp. nov., (E) dorsal, (F) left lateral. G, H: B. delicatus sp. nov., (G) dorsal, (H) left lateral. I, J: B. opacus sp. nov., (I) dorsal, (J) left lateral. K, L: B. undulatus sp. nov., (K) dorsal, (L) left lateral. Scale bar: 1mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B5C5D44FF14F9A81E882503.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608300/files/figure.png	https://doi.org/10.5281/zenodo.16608300	FIGURE 3. Male genitalia of Borysthenes spp. A–H: B. pantherinus sp. nov., I–P: B. opacus sp. nov., Q–X: B. undulatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	FIGURE 3. Male genitalia of Borysthenes spp. A–H: B. pantherinus sp. nov., I–P: B. opacus sp. nov., Q–X: B. undulatus sp. nov. A, I, Q: genitalia with pygofer removed, left lateral view; B, J, R: aedeagus, dorsal; C, K, S: aedeagus, ventral; D, L, T: aedeagus, right lateral; E, M, U: anal segment, dorsal; F, N, V: gonostyli, dorsal; G, O, W: pygofer, left lateral; H, P, X: pygofer, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B5C5D44FF14F9A81E882503.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608304/files/figure.png	https://doi.org/10.5281/zenodo.16608304	FIGURE 5. Female genitalia of Borysthenes spp. A, B: B. opacus sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C, D: B. undulatus sp. nov., (C) overview, left lateral, (D) posterior vagina, ventral. Scale bar: 0.5mm.	FIGURE 5. Female genitalia of Borysthenes spp. A, B: B. opacus sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C, D: B. undulatus sp. nov., (C) overview, left lateral, (D) posterior vagina, ventral. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B5C5D44FF14F9A81E882503.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608306/files/figure.png	https://doi.org/10.5281/zenodo.16608306	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	FIGURE 6. Habitat of Borysthenes spp. A: Type locality of Borysthenes pantherinus sp. nov. and B. opacus sp. nov., arrowed specific location where specimens were collected; B: habitus of B. opacus sp. nov.; C: habitus of B. pantherinus sp. nov.; D: habitus of B. undulatus sp. nov.; E: habitus of B. delicatus sp. nov.; F: aggregation of B. xuzhiae sp. nov. on rock surface, photographed during daylight, arrowed representatives of individuals; G: habitus of B. xuzhiae sp. nov.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
03CECD466B5C5D44FF14F9A81E882503.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16608302/files/figure.png	https://doi.org/10.5281/zenodo.16608302	FIGURE 4. Female genitalia of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C–E: B. delicatus sp. nov., (C) overview, left lateral, (D) posterior vagina, right lateral, (E) posterior vagina, ventral; F, G: B. pantherinus sp. nov., (F) overview, left lateral, (G) posterior vagina, ventral. Abbreviations: bc: bursa copulatrix; dr: ductus receptaculi; ov: ovipositor; pv: posterior vagina. Scale bar: 0.5mm.	FIGURE 4. Female genitalia of Borysthenes spp. A, B: B. xuzhiae sp. nov., (A) overview, left lateral, (B) posterior vagina, ventral; C–E: B. delicatus sp. nov., (C) overview, left lateral, (D) posterior vagina, right lateral, (E) posterior vagina, ventral; F, G: B. pantherinus sp. nov., (F) overview, left lateral, (G) posterior vagina, ventral. Abbreviations: bc: bursa copulatrix; dr: ductus receptaculi; ov: ovipositor; pv: posterior vagina. Scale bar: 0.5mm.	2025-07-21	Lyu, Tianlang		Zenodo	biologists	Lyu, Tianlang			
