taxonID	type	description	language	source
03CE87F40468694213ADFB6FFC9EFF28.taxon	type_taxon	Type species: Tetraonyx flavipennis Solier, 1851: 282, by monotypy (nec. flavipennis Guérin Ménéville, 1843: 22). Replaced by Tetraonyx xanthopterus Gemminger 1879: 293.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40468694213ADFB6FFC9EFF28.taxon	discussion	Taxonomic comments. Solier (1851) described Tetraonyx flavipennis within the genus Tetraonyx, placing it in a separate section named Picnoseus due to its morphological differences from all other known species, particularly in the prothorax (Lacordaire 1859). Today, such sections are considered equivalent to subgenera (Cazzañiga, pers. comm.). The recognition of Picnoseus at the generic level originates with Fairmaire & Germain (1863). The homonymy and replacement name for the type species of Picnoseus are discussed in the " Taxonomic Discussion " section under Picnoseus xanthopterus.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40468694213ADFB6FFC9EFF28.taxon	diagnosis	Diagnosis and comparisons. Picnoseus is characterized by the short antennae, with antennomere XI extending posteriorly to the pronotal base or slightly beyond the humeral angle; antennomeres IV – X becoming progressively wider; antennomere XI elongate (2 × longer than preceding antennomeres), its apex asymmetrically and abruptly tapered (Picnoseus type). It is distinguished from related genera by the following combination of characters: from Bokermania by the simple (non-serrate) claws; from Brasiliota by the head and pronotum not being red, the absence of red coloration on the basal quarter of the elytra, and the lack of a black humeral spot; from Denierota and Glaphyrolytt a by its non-metallic body coloration; from Lyttamorpha by the body not being elongate and narrow, the head not large and quadrangular, the thorax not long and narrow, and the tibiae not flattened; from Pseudomeloe by the elytra being non-rugose and completely covering the abdominal tergites; from Pseudopyrota by the head and pronotum not being orange; from Pyrota by the antenna not being moniliform (with antennomere VII not markedly short) and by the metatibial spurs being neither short nor truncate; and from Wagneronota by the elytra lacking three trace venation lines and the protarsi lacking sexual dimorphism.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40468694213ADFB6FFC9EFF28.taxon	description	Description of Picnoseus Small to medium sized species; length 6 – 15 mm. Coloration. Tegument color stated as black assumes variation to dark-brown or dark-red. Head and pronotum entirely black, with dark-brown luster in most species (except P. angelicae sp. nov.). Pro- and mesotibial spurs brown or dark-brow. Metatibial spurs brown with apical third translucent-pale brown. Pretarsal claws brown. Elytra with a combination of two colors, such as: P. circumseptus, P. ernestus sp. nov., P. flavocinctus, P. laevigatus, P. marginalis, P. rubrofasciatus, and P. roigjunienti sp. nov.; tricolors, such as P. gajardoi; or homogeneous (one color), such as: P. angelicae sp. nov., P. barrigatunoni sp. nov., P. nitidipennis, and P. xanthopterus. Tegument. Head and pronotum surface covered by punctures of different sizes and separation between them, except along midline and 2 raised juxtaocular areas; integument among puncture micro- reticulate or luster; with or without longitudinal median groove. Eyes entirely bordered with a thin denudated low rim. Clypeus and labrum with surface covered by dense or sparse punctures; clypeus with posterior half membranous or not. Pronotum anterior and posterior edges finely but distinctly bordered by a thin rim. Legs surface for most part sparsely punctate. Scutellum surface covered by dense or sparse punctures except along midline; integument between punctures lustrous or rugose. Elytra with surface slightly rugose, with 3 longitudinal costae, visible or faint. Setation. Very sparse; black, brown, or grayish; short (setiform) or elongate (piliform); each setae corresponding to a puncture. Clypeus and labrum with setae directed forward. Labrum with its inferior edge bearing elongated setae on their external sides, and shorter and thicker on middle. Maxillary palpomeres II and III with elongate, sparsely or very sparsely with erect setae (2 times longer than those of palpomere IV); palpomere IV with subappressed pilosity and very sparsely with partially erect setae. Labial palpomeres I and II with elongate, sparsely with partially erect setae (1.2 times longer than those of palpomere III); palpomere III bearing subappressed pilosity and scattered partially erect setae. Antennomeres I and II with elongate, sparse, and partially erect setae (longer than those of maxillary palpomeres); antennomeres III – XI with very short, subappressed pilosity, and with a few elongate, thin, and translucent erect setae, homogeneously distributed. Legs with modified setae allocated on ventral surface along protibiae or at distal half (that opposing femur); protarsomeres with pads on apical or entire ventral sides. Elytra with very short, short, or elongate setae, erect and very sparse (most conspicuous on humeral callus and marginal regions). Head. Suborbicular or subtriangular (Fig. 1); with or without subtle median groove from vertex to upper area of fronto-clypeal suture. Temples subparallel (maximum width at temples) or divergent (maximum width at occiput) on sides. Frons smoothly depressed transversely (Fig. 2). Slightly depressed at the level of posterior edge of eyes. Eyes flat or swollen; with anterior edge slightly or moderately broad emarginate; ventral lobe either abruptly narrowing below or gradually broadening. Fronto-clypeal suture arcuate, deeply or slightly marked. Clypeus flat or indistinctly convex, subrectangular, transverse, with parallel or divergent sides. Anterior edge of the clypeus is slightly curved. Labrum transverse, slightly depressed, its width equal to or up to 1.2 times that of clypeus; lateral edge smoothly to markedly rounded, imparting a bilobed appearance, either curved inward or straight; anterior edge concave or straight. Maxillary palpomere I conspicuously inflate, widest at middle, narrowing towards base; palpomeres II and III sub-trapezoidal, II similar to III; palpomere IV twice longer than precedent one, fusiform or subrectangular, with a sensory area on dorsal region (Fig. 3). Labial palpomere I conspicuously inflates widest at middle, narrowing towards base, palpomere II sub-trapezoidal (2 times longer than I); palpomere III narrowed towards apex (maximum width at base) (Fig. 4). Mandible short, markedly curved on external side, mostly covered by clypeus and labrum while resting (Fig. 5); or elongate, broad curved on external side, partially covered by clypeus and labrum while resting (Fig. 6). Antennal insertions contiguous to the eyes. Area in front of antennal insertions slightly elevated. Antennae filiform, short, extending posteriorly to base of pronotum; progressively widened from base to apex (Fig. 7). Antennomeres sub-cylindrical, closed articulated or not, antennomere XI long, 2 times longer than precedent one, with apex asymmetrically and abruptly tapered into a narrow apex- Picnoseus type. Pronotum. Subhexagonal with angulous sides or suborbicular with rounded sides (maximum width at anterior third) (Figs. 8 and 9). Anterior and posterior edges are almost curved. Dorsal surface is slightly declivity towards anterior edge and with a suborbicular depression on basal half. Scutellum. Subtriangular-shaped, wide, distally rounded and slightly depressed; surface with a longitudinal median groove or not; bordered with a thin denuded low rim; totally visible or partially visible (covered by pronotum at anterior half). Legs. Slender, relatively long, not sexually dimorphic. Femur is almost shorter than tibiae. Protibiae with spurs similar in length, slender and spiniform; mesotibial spurs similar in length, slender and spiniform, not parallel but divergent, forming an angle of 20 °; metatibial spurs similar in length, outer spur broadened apically, obliquely truncate, inner spur narrowed, sticklike, rotated inward; or dissimilar in length with outer spur short, obliquely truncate, inner spur thickened or not. Metatibial spurs are pallid apically only. Tarsi sub-cylindrical but laterally compressed, slightly expanded and emarginated distally. Pretarsal claws: dorsal blade evenly curved; ventral blade broad apically. Elytra. Elongate, completely covering abdominal tergites, or shortened, exposing the last 3 abdominal tergites. Hindwings. Recurrent vein (Rv) formed by medial veins (M 1 + 2) and (M 3 + 4) and crossvein (m-m). Hindwing venation is generally well-developed but simpler; in P. angelicae sp. nov., P. barrigatunoni sp. nov., P. laevigatus, and P. spock sp. nov., it is completely degenerated. Unfolded hindwing outline elongate subtriangulate with three edges (Fig. 23): Costal (CM), External (EM), Posterior (PM). Surface divided into three regions (Fig. 23): Remigial (Rr), Vannal (Vr), Jugal (Jr). In Rr, venation stable (reduced / less sclerotized in above four species. Principal structures: Costa (C) short / long, sclerotized, at basal CM. Radius (R) long, moderately / strongly sclerotized, forms Pterostigmatic area (Pa). Pa, R 1, and crossvein (r) form small Radial cell (Rc). Radial sector (Rs) runs along lower R edge; median part disappears, distal part remains as isolated branch (R 4 + 5). Media (M) sclerotized / slightly sclerotized, runs obliquely; generally divides basally into (M 1 + 2) and (M 3 + 4). These form an open Apterum cell (Ap). In Picnoseus, M 1 + 2 may disappear (P. rubrofasciatus), be vestigial (P. nitidipennis), sclerotized (P. gajardoi), or slightly sclerotized (P. ernestus sp n.). Distal part connected by pale, short, straight crossvein (r-m) and to M 3 + 4 by arc-like crossvein (m-m). Thus, M 1 + 2 and M 3 + 4 form the Rv. Both extend apically as separate veins M 2 and M 3. Vr generally has three veins: Cubitus C 1 (sclerotized / slightly sclerotized, sometimes with basal branch), C 2, C 3. Jugal region generally has three Jugal veins: J 1 (always independent, median in Jr, sometimes with basal branch), J 2, J 3 (occasionally vestigial / rudimentary). Abdomen. Pygidium subtriangular, posteriorly emarginate (with triangular notch). Female genitalia (Fig. 34): as other meloid beetles, principal components of female reproductive system consist of internal structures: paired ovaries (OV), a developed calix (Cl), well developed spermathecal capsule (SpthCap) that continued with a spermathecal duct (SpthDuc), accessory gland (Ag) elongate tubular and thin, and well developed, tubular vagina (Vag). External structures: two valvifer (Va) tubular, localizated ventrolaterally; proximally each Va is produced into a ventrolateral basal stem (BsVa). Distally, Va is nearly cylindrical and bear elongated setae and an unjointed, suboval stylus (Sty) with elongated setae towards apex. Male genitalia (Fig. 35): parameres (Pa) narrowed distally, curved ventrally; an indistinctly membranous area; phalobase (Ph) with a general plate-shaped, with a proximally concavity; parameres and phalobase sclerotized or membranous joined. Aedeagus (Ae) with 2 or 3 hooks: dorsal aedeagal hook (DAH) and ventral aedeagal hook (VAH); Pa and Ae together constituting the phallus organs. Spiculum gastrale (SpGs) with developed laterally curved arms or not.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40468694213ADFB6FFC9EFF28.taxon	biology_ecology	Host Plant and Insects. Literature records on the host plants of Picnoseus species are limited to P. nitidipennis and P. circumseptus, which are associated with grasses, and P. xanthopterus, which feeds on the flowers of Cristaria glaucophylla (Malvaceae) (Denier 1918). However, recent data from amateur contributors on platforms such as iNaturalist indicate that some Picnoseus species feed on the flowers or pollen of Aizoaceae (Carpobrotus chilensis), Boraginaceae (Heliotropium krauseanum), Malvaceae (Cristaria spp), Montiaceae (Cistanthe longiscapa, Cistanthe spp.), Solanaceae (Nolana spp.; Sphaeralcea spp., Solanum trinominum), and Cactaceae species. These observations supplement the limited existing literature, which previously only documented associations with grasses and one species of Malvaceae. The flowers of some of the observed plant species are visited by Bombus spp. (Hymenoptera), which suggests the putative presence of phoretic meloid larvae. This inference is based on Bologna & Pinto (2001), who report that the triungulin is phoretic in several meloid genera associated with apoid hosts. These larvae typically attach to bees during flower visits to gain indirect access to larval food resources (Figs. 61 – 66). Natural History. According to specimen labels and amateur contributions from amateur iNaturalist users, Picnoseus activity occurs primarily in the spring, with fewer records in the late summer and autumn. The following species were observed during the specified months: P. angelicae sp. nov. and P. barrigatunoni sp. nov.: middle September; P. circumseptus: middle September to December; P. ernestus sp. nov.: October, with a single record in May; P. gajardoi: September and October; P. laevigatus: September to November; P. marginalis: October to November; P. nigropicta: September; P. nitidipennis: October, November, and December, with a single record in May; P. rubrofasciatus: October and November; P. roigjunienti sp. nov.: September and October; P. spock sp. nov.: September and October, with a single record in July; and P. xanthopterus: September to January.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40468694213ADFB6FFC9EFF28.taxon	distribution	Distribution range. The species of Picnoseus are endemic to southern South America (Figs. 67 ‒ 73), including the South American Transition Zone (Roig-Juñent et al. 2018) and the Central Chilean subregion of the Andean Region (Fig. 82). They are principally found principally in central and northern Chile, and three species extend into northwest Argentina (Figs. 74 – 82).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40461694913ADF9E2FC6FF89C.taxon	description	urn: lsid: zoobank. org: act: 1 C 8 A 4764 - 5421 - 4771 - B 3 EA- 15371 C 066500 (Fig. 47)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40461694913ADF9E2FC6FF89C.taxon	materials_examined	Type material. Holotype ♀ Chile, Copiapó, 15 sep. 1984, leg. J. E. Barriga. Holotype Chile Picnoseus angelicae Campos-Soldini, JEBTC, 112696. Paratype 1 ♀ Chile prov. Caldera, 15 sep. 1984, leg. J. E. Barriga-Tuñón Chile 117091. Paratype, Picnoseus angelicae, Campos-Soldini, LEC.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40461694913ADF9E2FC6FF89C.taxon	diagnosis	Diagnosis. Small- sized species. Body color pattern: head, antennae, mouthparts, pronotum, and legs uniformly brown; abdomen dark-brown; elytra pale brown; scutellum brown on proximal half and pale brown on distal half. Head and pronotum punctuation small, sparse, and shallow; punctures uniformly distributed; integument between them micro-reticulate. Setation short, very sparse, and partially erect on the dorsal sides of head and pronotum; ventrally long. Antennomere I subglobose, bead-like; II cup-shaped; III progressively divergent, widest distally with a broad, often obscure curvature; IV – VII subtriangular; VIII – X proximally narrowed, thereafter of uniform width. Protibial spurs similar in length, outer spur broadened apically, obliquely truncate, apex curved outwards, inner spur narrowed, and sticklike, rotated inward. Elytra with total length 5 mm; convex edges (maximum width at middle: 2.7 mm); suture dehiscent at posterior one-third; elytra distally rounded. Hindwings with reduced venation, see table 2 a (Fig. 24).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40461694913ADF9E2FC6FF89C.taxon	description	Description of Holotype. Small- sized species; length 10 mm. Coloration. Head, antennae, mouthparts, pronotum, and legs uniformly brown. Abdomen dark-brown. Elytra yellow. Scutellum brown proximally, pale- brown distally. Tegument. Head and pronotum covered with small, sparse, and shallow punctures uniformly distributed; integument between them micro-reticulate. Clypeus and labrum with small punctures (1.2 times smaller than those on frons). Scutellum covered with very small punctures, except at distal third. Elytra without visible longitudinal costate. Setation. Pale- brown; short, very sparse, and partially erect on the dorsal sides of head and pronotum. Elytra with short (similar to those on head), erect, and very sparse setae (being most visible on humeral callus and edges). Long ventrally (twice the length of the dorsal ones). Clypeus and labrum with setae very sparse, twice the length of the frons. Protibial ventral surface (that opposes profemur) with an extensive patch of subappressed, transversely directed silky pubescence at distal two-thirds. On the protarsus, distal ventral pads are homogeneously distributed; the proximal surface has thicker, sparse setae. Head. Suborbicular (long 1.6 mm; wide 1.5 mm; maximum width just above eyes); surface with a longitudinal median groove from vertex to posterior area of fronto-clypeal suture. Temple subparallel, rounded posteriorly. Frons slightly transversely depressed before clypeal suture. Eyes slightly emarginated in front of antennal insertion; abruptly tapering below from widest aspect (wide 0.4 mm; long 0.7 mm). Clypeus flat, subrectangular, with subparallel sides (wide 0.2 mm; long 0.7 mm). Labrum transverse, lateral edge evenly rounded; ventrally with a smooth curvature (wide 0.7 mm; long 0.2 mm). Maxillary palpomere IV subrectangular, proximally narrowed then maintaining uniform width distally. Labial palpomere III subrectangular (with same width in both extremes). Mandible partially covered by clypeus and labrum when resting. Antennae with antennomeres I – V loosely articulate, antennomeres VI – XI closely articulated (Fig. 10). Antennomere I subglobose, bead-like; II cup-shaped; III progressively divergent, widest distally with a broad, often obscure curvature; IV – VII subtriangular; VIII – X proximally narrowed, thereafter of uniform width. Antennomeres length: I 0.17 mm; II: 0.07 mm; III: 0.30 mm; IV and VII: 0.25 mm; VIII – X: 0.22 mm; XI: 0.50 mm. Pronotum. Subhexagonal (long 1.5 mm; wide 1.5 mm) maximum width at anterior one-third, with angulous borders; surface with a longitudinal median groove (scarcely visible). Legs. Femora is almost shorter than tibiae: forefemur 1.3 mm, foretibia 1.5 mm; mesofemur 1.8 mm, mesotibia 2.1 mm; metafemur 2.3 mm, metatibia 2.6 mm. Protibial spurs similar in length, outer spur broadened apically, obliquely truncate, apex curved outwards, inner spur narrowed, sticklike, rotated inward. Tarsomere measurement: protarsomeres (I 0.3 mm; II 0.16 mm; III and IV 0.13 mm; and V 0.3 mm); mesotarsomeres (I 0.6 mm; II and III 0.3 mm; IV 0.2 mm; and V 0.5 mm); and metatarsomeres (I 0.7 mm; II 0.4 mm; II 0.3 mm; and IV 0.6 mm). Elytra. Total length 5 mm; with convex edges (maximum width at middle: 2.7 mm); suture dehiscent at posterior one-third; elytra distally rounded. Hindwings with reduced venation, see table 2 a (Fig. 24). Male genitalia. Unknown. Female genitalia. Va plate sclerotized, broad distally; proximal BsVa irregular in outline. Distal end of Va with a subtriangular Sty, its length subequal to distal width of Va. Both structures with long setae.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40461694913ADF9E2FC6FF89C.taxon	etymology	Etymology. Picnoseus angelicae sp. nov. is dedicated to María Angélica Müller, grandmother of MPCS, in recognition of her enduring love, strength, and profound inspiration.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40461694913ADF9E2FC6FF89C.taxon	distribution	Geographic distribution. This species is known only from northern Chile, in the Coquimbo biogeographic province of Central Chile subregion (Fig. 67). The area is a dry semidesert, and the specimens were collected on the coast, which exhibits more humid habitats.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40461694913ADF9E2FC6FF89C.taxon	biology_ecology	Natural History. According to the specimen labels, the adult was collected in spring (September). There is no available information on the host plants associated with this species.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40463694B13ADF9D3FD62F942.taxon	description	urn: lsid: zoobank. org: act: DD 1 B 1 D 19 - 50 AA- 4073 - 885 E- 590 B 7 F 6 B 991 B (Fig. 48)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40463694B13ADF9D3FD62F942.taxon	materials_examined	Type Material. Holotype ♀ Picnoseus barrigatunoni, Algarrobal, Huasco, Chile, Col. 15 / IX / 84, J. E. Barriga. Holotype Chile Picnoseus barrigatunoni Campos-Soldini, JEBTC, 8549.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40463694B13ADF9D3FD62F942.taxon	diagnosis	Diagnosis. Medium- sized species. Body coloration uniformly black, lustrous tegument. The head surface is covered with large, deep and dense punctures, but separated from each other. Pronotum with punctures medium and sparse. Setation black, piliform, erect and very sparse dorsally; sparse ventrally. Antennomere I widest distally, cup-shaped; antennomere II similarly cup-shaped; antennomere III slightly expanded anteriorly; antennomeres IV – VI fusiform, broadest at midlength (distal width about one-third of maximum); antennomeres VII – X subtrapezoidal. Metatibiae with outer spur shorter than inner spur; outer spur broadened apically, obliquely truncate; inner spur narrowed, sticklike, and rotated inward. Elytra with total length 7.5 mm; suture distally imbricated and rounded. Hindwings with reduced venation (Fig. 24).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40463694B13ADF9D3FD62F942.taxon	description	Description of Holotype. Medium- sized species; length 11.5 – 12 mm. Coloration. Black, lustrous tegument. Tegument. Head surface covered with large, deep and dense punctures, but separated from each other; integument between them micro-reticulate. Clypeus and labrum punctuation similar to those on frons. Mandible with external border with punctures similar to those on clypeus. Pronotum with punctures medium and sparse; integument between them microreticulate. Scutellum surface with small puncture; integument between them micro-reticulate. Elytra surface with three longitudinal costae weakly visible. Setation. Black, piliform, erect and very sparse dorsally; sparse ventrally. Foretibiae with proximal third of ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed pubescence at distally half; distally, foretarsus has ventrally homogeneously distributed pads, proximal surface covered with sparse thicker setae. Head. Suborbicular (length 2.3 mm; width 2.6 mm; maximum width just above eyes); surface with a longitudinal median groove from vertex to the upper area of fronto-clypeal suture. Temple subparallel, rounded posteriorly. Frons transversely depressed before clypeal suture. Eye flat; slightly emarginate anteriorly; abruptly narrowing below (wide 0.7 mm; long 1.3 mm). Clypeus flat, with subparallel sides (wide 0.4 mm; long 1.4 mm). Labrum with smoothly rounded lateral sides; ventrally slightly concave (wide 0.4 mm; long 1.1 mm). Maxillary palpomere IV fusiform (maximum width at middle). Labial palpomere III narrowed towards the proximal region (maximum width distally). Mandibles are elongate, robust, broad on the external side, partially covered by the clypeus and labrum while resting. Antennae loosely articulated (Fig. 11). Antennomere I widest distally, cup-shaped; antennomere II similarly cup-shaped; antennomere III slightly expanded anteriorly; antennomeres IV – VI fusiform, broadest at midlength (distal width about one-third of maximum); antennomeres VII – X subtrapezoidal. Antennomeres length: I: 0.37 mm; II: 0.17 mm; III: 0.57 mm; IV: 0.50 mm; V – X: 0.45 mm; XI: 0.75 mm. Pronotum. Subhexagonal (long 2 mm; wide 2.6 mm) maximum width at posterior one-third, with angulous or rounded borders; surface with longitudinal median groove. Legs. Profemur with the same length as protibia (3.3 mm); mesofemur longer than mesotibia (3.8 mm; 3.3 mm respectively); metafemur slightly shorter than metatibiae (4.3 mm; 4.6 mm respectively). Metatibiae with outer spur shorter than inner spur; outer spur broadened apically, obliquely truncate; inner spur narrowed, sticklike, and rotated inward. Tarsomere measurement: foretarsomere (I 0.8 mm; II 0.5 mm; III and IV 0.4 mm; V 0.8 mm); mesotarsomere (I 0.8 mm; II – IV 0.4 mm; V 0.8 mm); and metatarsomere (incompletes). Elytra. Total length 7.5 mm; with divergent sides (maximum width at distal two-third: 6 mm) or with subparallel sides; suture distally imbricated, elytra distally rounded. Hindwings with reduced venation, see table 2 a (Fig. 24). Male genitalia. Unknown. Female genitalia. Va plate sclerotized, subtriangular; proximally the BsVa is pointed with an external angular protuberance; suborbicular Sty shorter than Va.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40463694B13ADF9D3FD62F942.taxon	etymology	Etymology. The species P. barrigatunoni is named in honor of J. E. Barriga-Tuñón, recognizing his enduring passion for entomology.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40463694B13ADF9D3FD62F942.taxon	distribution	Geographic Distribution. This species occurs in Santiago and Coquimbo biogeographic provinces in Chile (Fig. 67).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40463694B13ADF9D3FD62F942.taxon	biology_ecology	Natural History. According to the specimen labels, adults were collected in spring (September). No information is available on the host plants for this species.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40462695513ADF930FED8FBC0.taxon	description	(Fig. 49)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40462695513ADF930FED8FBC0.taxon	materials_examined	Type Materials. Denier (1918) indicates that the type of Tetraonyx circumseptus is deposited at MNHN. After several searches in the Museum collection, we have not been able to find the type specimens (holotype and allotype). They are probably lost or misplaced. However, we have studied two males of P. circumseptus deposited at MLPC, identified by the specialist P. Denier, and seven males deposited at JEBTC. The specimens examined conform in all respects to the original description of P. circumseptus. Material examined. Chile, Malleco: Lonquimay, Coll. J. E. Barriga, 22 / XII / 1994, 2 ♂ JEBTC. Chile: Algarrobal, 1 / X / 1987, Coll. J. E. Barriga, 1 ♂ MLPC. Chile: Algarrobal, 1 / X / 1987, Coll. J. E. Barriga, 1 ♂ JEBTC. Chile, Copiapó: Caldera, Coll. J. E. Barriga, 16 / IX / 1984, 5 ♂ JEBTC; 9 ♂ LEC. Chile: Santiago de Chile, Coll. J. E. Barriga, 15 / IX / 1984, 1 ♂ JEBTC. Chile, Valle de Aconcagua Los Andes, Coll. J. E. Barriga, XI- 1984, 1 ♂ JEBTC.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40462695513ADF930FED8FBC0.taxon	diagnosis	Diagnosis. Elytra black with an irregular reddish-brown spot at base or an irregular reddish-brown spot at base that continued as marginal vittae towards apex. Metatibiae with spurs similar in length, subparallel, outer spur broadened apically, obliquely truncate, inner spur narrowed, sticklike. Antennomere I conspicuously inflated, broadest at midlength, with a subtle anteroapical curvature; antennomere II cup-shaped; antennomere III – VII widest distally; antennomere VIII subtriangular ¸ antennomeres IX – X subparallel-sided, with convergent proximal thirds. Profemur with the same length as protibia, the subsequent legs with femora almost shorter than tibia. Elytra with total length 7.5 mm, with divergent sides (maximum width at distally two-third: 3 mm); distally rounded. Hindwing veins compositions are summarized in table 2 a. Male genitalia with spiculum gastrale bearing external spines continuated with two lobes separated by a broad middle emargination, proximally narrowed and sinistrally curved; aedeagus with three hooks (Fig. 36).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40462695513ADF930FED8FBC0.taxon	description	Redescription. Small- medium sized species; length 7.5 – 12 mm. Coloration. Head and pronotum dark- brown; antennae, mouth parts dark- brown or brown. Clypeus brown at distal third. Legs black or brown, lustrous tegument; or with femur and tibiae dark- brown, Tarsomere brown. Scutellum black, lustrous tegument. Elytra black with an irregular reddish- brown spot at base or an irregular reddish- brown spot at base that continued as marginal vittae towards the apex. Tegument. Head surface with large, deep and sparse punctures except along the median groove; integument among them micro-reticulate. Clypeus with small and deep punctures sparsely distributed, almost absent at inferior third, which is membranous. Labrum punctures similar to those on clypeus but smaller. Pronotum surface with medium-sized, shallow sparse punctures; integument between them lustrous. Scutellum surface micro-reticulate except at distal third which is lustrous. Elytra surface with three weakly defined longitudinal costae. Setation. Mostly black or dark- brown. Head with setae short and erect dorsally, elongate ventrally (twice longer than ventral ones). Pronotum bearing very sparse, short, semi-erect setae. Protibia with ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed silky pubescence at distal two-third. Distally, protarsus (I – IV) has ventrally homogeneously distributed pads; anteriorly the surface is covered with black, thicker and sparse setae. Elytra with very short setae (similar to those on head), erect and very sparse (being most visible on humeral callus and on edges). Head. Suborbicular (length: 1.5 mm; width: wide 1.8 mm; maximum width just above eyes); surface with a longitudinal subtle median groove from the vertex to the upper area of fronto-clypeal suture. Temple subparallel, rounded posteriorly. Frons transversely depressed before clypeal suture. Eyes slightly swollen, with anteriorly slightly emarginated in front of antenal insertion, ventral lobe abruptly narrowing below (wide 0.6 mm; long 0.8). Clypeus swollen, subtrapezoidal or subrectangular (wide 0.4 mm; long 0.8 mm). Labrum with moderately rounded lateral edge, ventral edge concave; or smoothly rounded laterally, straight ventrally (wide 0.4 mm; long 0.9 mm). Maxillary palpomere IV subrectangular, slightly divergent towards distal region (maximum width distally). Labial palpomere III subrectangular (subparallel- sided, maintaining approximately equal width at both proximal and distal edges). Mandible short, markedly curved on the external side, mostly covered by clypeus and labrum while resting; or elongate, broad curved on the external side, partially covered by clypeus and labrum while resting. Antennae loosely articulated (Fig. 12). Antennomere I conspicuously inflated, broadest at midlength, with a subtle anteroapical curvature; antennomere II cup-shaped; antennomere III – VII widest distally; antennomere VIII subtria ngular ¸ antennomeres IX – X subparallel-sided, with convergent proximal thirds. Antennomeres length: I 0.30 mm; II 0.07 mm; III 0.42 mm; IV – X 0.37; XI 0.57 mm. Pronotum. Suborbicular (long 1.5 mm; wide 1.7 mm) maximum width at middle, with rounded borders; surface with a longitudinal median groove, either complete (extending full length) or medially restricted (interrupted at midpoint); slightly curved anteriorly, almost straight basally. Legs. Profemur with the same length as protibia (1.5 mm and 1.6 mm respectively); the subsequent legs with femora almost shorter than tibia, (1.9 mm – 2.1 mm; 2.3 mm – 2.6 mm respectively). Metatibiae with spurs similar in length, subparallel, outer spur broadened apically, obliquely truncate, inner spur narrowed, sticklike. Tarsomere with protarsomere (I 0.6 mm; II 0.16 mm; III and IV 0.13 mm; and V 0.4 mm); mesotarsomere (I 0.5 mm; II 0.3 mm; III and IV 0.16 mm; and V 0.3 mm); and metatarsomere (I 0.6 mm; II and III 0.3 mm; and IV 0.5 mm). Elytra. Total length 7.5 mm; edge progressively divergent sides (maximum width at the distal two-thirds: 3 mm); suture parallel; elytra distally rounded. Hindwing veins compositions are summarized in table 2 a (Fig. 25). Male genitalia (Fig. 36). SpGs bearing external spines continuated with two lobes separated by a broad middle emargination; proximally narrowed and sinistrally curved. Pa distally with elongate apical setae; surface incompletely sclerotized, each lobe with an extensive membranous area medially, except for a narrow, sclerotized median strip running along the ventromedial surface of each arm. Ph with a deep U- shaped concavity; pa and ph fused by a thin sclerotized tegument. Ae distally with three hooks: two DAH (one often inconspicuous), VAH very robust. AeA short and straight. Female genitalia. Unknown. Intraspecific Variation. The specimens herein studied, as well as the specimens observed in the iNaturalist platform, have the elytra completely covering the abdominal tergites or shortened, exposing the last three tergites.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40462695513ADF930FED8FBC0.taxon	discussion	Taxonomic Comments. Denier (1918: 210) included T. circumseptus in Picnoseus and treated it as a variety of P. nitidipennis. Herein, we consider that P. nitidipennis and P. circumseptus stat. rev. are distinct species, based on differences in external appearance, hindwings morphology, and male genitalia characteristics (see description of P. circumseptus and P. nitidipennis).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40462695513ADF930FED8FBC0.taxon	distribution	Geographic Distribution. This species is endemic to Chile, ranging from the subantarctic Maule to the Coquimbo biogeographic provinces (Fig. 67).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40462695513ADF930FED8FBC0.taxon	biology_ecology	Natural History. According to the specimen labels and the information obtained from the iNaturalist platform, this species was observed in spring (from September to November) observed on grasses, visiting Cristaria glaucophylla (Fig. 61) and eating its flowers (Available from: https: // www. inaturalist. org / observations / 140982315; date of access April 2025).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047C695613ADFB8BFC41F868.taxon	description	urn: lsid: zoobank. org: act: 8075 F 3 BF- 4 CFC- 46 EA-BFC 1 - F 7 ED 953 D 09 A 9 (Fig. 50)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047C695613ADFB8BFC41F868.taxon	materials_examined	Type Material. Holotype ♀ Chile prov. Huasco, 1 oct. 1987, Picnoseus ernestus, leg J. E. Barriga, Colección J. E. Barriga-Tuñón, Chile 090868, LEC. Paratype 2 ♀. White printed label Chile prov. Algarrobal, 1 oct. 1987, leg. J. E. Barriga. White printed label Colección, J. E. Barriga Chile 097348, JEBTC. Paratype 1 ♀ Chile prov. Paipote, Almagro a camino, internacional 21 - V- 1991 col. S. Roig, IADIZA. Paratype 1 ♀ Chile prov. Huasco, P. N. Llanos de Challe, Costa, 25 oct. 2011, 27 ° 59 ’ S 71 ° 08 ’, 40 mt, leg. J. E. Barriga-Tuñón et al. LEC.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047C695613ADFB8BFC41F868.taxon	diagnosis	Diagnosis. Medium- sized species. Elytra reddish or dark-reddish, lustrous tegument with black marginal and sutural vittae from base to the apex; both vittae converging distally. Head surface covered with large, deep, sparse or dense punctures; pronotum surface covered with small, shallow, and very sparse punctures. Setation black, piliform or setiform. Dorsum with erect setation; ventral surfaces with recumbent setation. Antennomere I swollen, widest at midlength, distinctly constricted proximally; antennomere II cup-shaped; antennomere III slightly expanded distally, with a weak, often obscure ventral curvature; antennomeres IV – V broader posteriorly; antennomeres VI – X subtrapezoidal. Metatibia spurs are similar in length, not parallel but divergent, forming an angle of 20 degrees; outer spur broadened apically, obliquely truncate; inner spur narrowed, sticklike. Hindwing veins compositions are summarized in table 2 a. Male genitalia with spiculum gastrale bearing two subrectangular arms, joined by an irregular median concavity, proximally narrow and slightly curved to the left; aedeagus distally with two robust spines (Fig. 37).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047C695613ADFB8BFC41F868.taxon	description	Description of Holotype. Medium- sized species; length 12 – 14.5 mm. Coloration. Head, pronotum, antennae, mouthparts and underside black; lustrous tegument. Scutellum dark-brown or black. Elytra reddish or dark- reddish, lustrous tegument with black marginal and sutural vittae from base to the apex; both vittae converging distally. Tegument. Head surface covered with large, deep, sparse or dense punctures; integument between them micro-reticulate. Pronotum surface covered with small, shallow, and very sparse punctures; integument between them luster. Clypeus surface with punctures similar to those on frons. Labrum surface with punctures smaller than those on clypeus. Scutellum with small, uniformly distributed punctures except along the median line and distal third, which are impunctate. Elytra surface with three longitudinal slightly visible costae. Setation. Black; piliform or setiform. Dorsum with erect setation; ventral surfaces with recumbent setation. Clypeus and labrum with setae as long as those on frons. Scutellum glabrous or with very sparse setae, setae very short (1.5 shorter than the rest of the body). Protibia with ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed silky pubescence at distally two-third; male protarsus with ventral pads homogeneously distributed throughout the length; female protarsus with ventral pads confined distally, proximal surface covered with thicker and sparse setae. Elytra with pubescence being most visible on humeral callus and elytral edge. Head. Suborbicular (length: 1.6 mm; width: 1.5 mm; maximum width just above eyes); surface with marked longitudinal median groove from vertex to the upper area of fronto-clypeal suture. Temple subparallel, rounded or straight posteriorly. Frons transversally depressed before the suture. Eye flattened, with moderately broad emargination at anterior border; ventral lobe gradually tapered (wide 0.5 mm; long 1 mm). Clypeus transverse (wide 0.3 mm; long 1.1 mm). Labrum smoothly rounded lateral edge, ventral edge broad concave (wide 0.3 mm; long 1.1 mm). Maxillary palpomere IV fusiform (maximum width at middle). Labial palpomere III narrow distally (maximal width at proximal one-third). Mandible elongate, broad curved on the external side, totally or partially covered by clypeus and labrum while resting. Antennae loosely articulated (Fig. 13). Antennomere I swollen, widest at midlength, distinctly constricted proximally; antennomere II cup-shaped; antennomere III slightly expanded distally, with a weak, often obscure ventral curvature; antennomeres IV – V broader posteriorly; antennomeres VI – X subtrapezoidal. Antennomeres length: I 0.25 mm; II 0.07 mm; III 0.37 mm; IV 0.27 mm; V – X 0.32 mm; XI 0.57 mm. Pronotum. Subtrapezoidal (long 2.1 mm; wide 2 mm), with rounded or angulous borders; anterior and posterior edges slightly curved. Legs. Femora shorter than tibia: fore- femur 2.9 mm, fore- tibia 2.8 mm; meso- femur 3.4 mm, mesa- tibia 3.5 mm; meta- femur 3.8 mm, meta- tibia 3.9 mm. Metatibia spurs are similar in length, not parallel but divergent, forming an angle of 20 degrees; outer spur broadened apically, obliquely truncate; inner spur narrowed, sticklike. Tarsomere measurement: foretarsomere (I 0.6 mm, II and III 0.5 mm, IV 0.4 mm, and V 0.6 mm); mesotarsomere (I 0.9 mm, II 0.6 mm, III 0.5 mm, IV 0.36 mm, and V 0.6 mm); and metatarsomere (I 1.3 mm, II 0.9 mm, III 0.6 mm, and IV 0.7 mm). Elytra. Total length 6.5 mm; with edges progressively divergent (maximum with at distal two-third: 4 mm), suture dehiscent at distal one-third; elytra rounded distally or convergent towards distal one-third. Hindwing veins compositions are summarized in table 2 a (Fig. 26). Male genitalia (Fig. 37). SpGs bearing two subrectangular arms, joined by an irregular median concavity; proximally narrow and slightly curved to the left. Pa distally with elongate apical setae; surface incompletely sclerotized, medially membranous except for a narrow, sclerotized strip that diverges distally. Ph with a deep Vshaped concavity; Pa and Ph fused by a sclerotized tegument. Ae distally with two robust hooks: one DAH and one VAH (the latter more robust). AeA short and straight. Female genitalia. Va plate subtriangular; proximally the BsVa is pointed with an external and internal protuberance; external border with a square- shaped concavity; Sty subrectangular 1.2 x times longer than Va. Intraspecific variations. The specimens herein studied presents: male protarsus with ventral pads homogeneously distributed towards the distal area; female protarsus with ventral pads confined to the distal portion, proximal surface covered with thicker and sparse setae.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047C695613ADFB8BFC41F868.taxon	etymology	Etymology. The species P. ernestus is dedicated to my life partner, Carlos Ernesto Camaño Schettini.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047C695613ADFB8BFC41F868.taxon	distribution	Geographic Distribution. This species is known only of the type locality, in the desert Coquimbo biogeographic province of Chile (Fig. 68).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047C695613ADFB8BFC41F868.taxon	biology_ecology	Natural History. According to the specimens label, adults were collected in spring (September and October). Picnoseus ernestus was observed feeding on flowers of Carpobrotus chilensis (Aizoaceae). Available from: https: // www. inaturalist. org / observations / 198360616; date of access June 2025.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047E695013ADFF7FFA74FDEC.taxon	description	(Fig. 51)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047E695013ADFF7FFA74FDEC.taxon	materials_examined	Type Material. Holotype ♂ Chile, Biobío, Concepción, C. E. Porter, MLPC. Examined. Paratype ♂, from Chile, Santiago, Bío-Bio and Coquimbo: Elqui, MLPC. Examined.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047E695013ADFF7FFA74FDEC.taxon	diagnosis	Diagnosis. Picnoseus flavocinctus can be reliably distinguished by the following characters: body mostly black; elytra bicolored: proximal half pale-brown (each humeral region bearing a suborbicular reddish macula with central dark- reddish spot) and distal half gradually transitioning from reddish to dark- reddish. Head surface covered with large, shallow and dense punctures; uniformly distributed but absent on frons between eyes; pronotum surface covered with large, sparse and shallow punctures. Profemur shorter than protibia; femur and tibiae II and III with the same length. Metatibiae with spurs dissimilar, outer spur distinctly expanded and obliquely truncate, inner spur narrowed, sticklike. Elytra with total length 6.5 mm; with convergent edge at distal two-third (maximum width at proximal one-third); suture dehiscent along the distal two-thirds. Hindwing veins compositions are summarized in table 2 a. Male genitalia with spuculum gastrale with two stick-like lateral arms joined with broad median emargination, bearing two small median spines; proximally narrowed and curved to the right; aedeagus with two hooks (Fig. 38).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047E695013ADFF7FFA74FDEC.taxon	description	Redescription. Medium- sized species; length 12.5 mm. Coloration. Mostly black with lustrous tegument. Elytra bicolored: proximal half pale-brown (each humeral region bearing a suborbicular reddish macula with central dark-reddish spot) and distal half gradually transitioning from reddish to dark-reddish. Scutellum black. Tegument. Head surface covered with large, shallow and dense punctures; uniformly distributed but absent on frons between eyes; integument between punctures micro- reticulate. Clypeus and labrum puncture somewhat smaller than those on frons; the punctures in the clypeus are absent in its distal border, which is membranous and smooth. Pronotum surface covered with large, sparse and shallow punctures; integument among them luster. Elytra surface with three weakly visible longitudinal costae. Scutellum surface covered with very small punctures, like small pit. Setation. Dark- brown and short dorsally; grayish and elongate ventrally. Clypeus and labrum bearing setae similar to those on the frons. Profemur with distal half of ventral surface with pale, thicker and very sparse setae. Portibia with ventral surface (that opposing femur) with pale, thicker and sparse setae at distal-third. Distally, protarsus has ventrally homogeneously distributed pads; proximal surface glabrous. Head. Suborbicular (long 1.5 mm, wide 1.8 mm; maximum width just above eyes). Temple subparallel, straight posteriorly. Frons with a suborbicular depression on frons (between eyes and before clypeal suture). Eye swollen; slightly emarginated in front of antenal insertion; ventral lobe rounded (wide 0.6 mm; long 0.8 mm). Clypeus transverse (wide 0.4 mm; long 0.7 mm). Labrum with moderately rounded lateral edge; ventral edge broadly concave (wide 0.4 mm; long 0.8 mm). Maxillary palpomeres IV fusiform (maximum width at middle). Labial palpomere III subrectangular (maximum width in both extremes, with parallel sides). Mandibles robust, broad curved externally, partially covered by clypeus and labrum while resting. The specimen studied has damaged antennae. Pronotum. Subhexagonal (long 1.6 mm; wide 1.8 mm) maximum width at apical third, with angulous borders; anterior and posterior edges slightly curved. Legs. Profemur shorter than protibia (1.6 mm; 1.4 mm respectively); femur and tibiae II and III with the same length (1.8 mm; 2.49 mm respectively). Metatibiae with spurs dissimilar, outer spur distinctly expanded and obliquely truncate, inner spur narrowed, sticklike. Tarsomere measurement: protarsomere (I 0.3 mm; II, III and IV 0.23 mm; V 0.49 mm); mesotarsomere (I 0.49 mm; II and III 0.33 mm; IV 0.23 mm; and V 0.56 mm); and metatarsomere (I 0.66 mm; II 0.49 mm; III 0.33 mm; and IV 0.56 mm). Elytra. Total length 6.5 mm; with convergent edge at distal two-third (maximum width at proximal one-third); suture dehiscent along the distal two-thirds. Hindwing veins compositions are summarized in table 2 a (Fig. 27). Male genitalia (Fig. 38). Distally, SG with two stick- like lateral arms joined with broad median emargination, bearing two small median spines; proximally narrowed and curved to the right. Pa surface incompletely sclerotized, with median portion membranous except for a narrow median strip which runs along the ventromedial surface of each arms and diverged apically; PhA with a medial V- like concavity; Pa and PhA fused ventrally at their bases with marginal sclerotized tegument. Distally, Ae with two hooks: one DAH robust, and one VAH very robust. Proximally, AeA long and curved. Female genitalia. Unknown.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047E695013ADFF7FFA74FDEC.taxon	distribution	Geographic Distribution. This species is endemic of Chile, and occurs in the subantarctic Maule and central Santiago and Coquimbo biogeographic provinces (Fig. 68).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047E695013ADFF7FFA74FDEC.taxon	biology_ecology	Natural History. Species label points that P. flavocinctus has been collected in spring (September). There is no available information on the host plants associated with this species.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047E695013ADFF7FFA74FDEC.taxon	discussion	Remarks. Previous to this contribution the holotype of P. flavocinctus were not placed in the drawer of the type material deposited at the (MLPC), but were misplaced along with specimens in the general collection. Based on the information included in the original publication and that on the labels, we consider this specimen as the true type.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40479695113ADFD6FFC92F9A4.taxon	description	(Fig. 52)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40479695113ADFD6FFC92F9A4.taxon	materials_examined	Type Material. Lectotype ♂ Chile: Coquimbo, Paihuano, Dr. Rob. Gajardo T. XI. 34, ♂ Picnoseus gajardoi, Denier Cotypus ♂ Museo De La Plata, Lectotype Picnoseus gajardoi Denier, Det. Selander, MLPC 611 / 1. Examined. Material examined. Chile, Copiapó: Caldera, Coll. J. E. Barriga, 15 / IX / 1984, 5 ♀ JEBTC. Chile, Coquimbo: Paihuano, Coll. Rob. Gajardo, XI / 1934, 1 ♀ LEC. Chile, Huasco, Algarrobal, Coll. J. E. Barriga, 15 / IX / 1984, 5 ♀ JEBTC. Chile: Paihuano, Coll. F. Ruiz, 17 / X / 1937, 3 ♂ LEC. Chile, Valle de Elqui: Rivadavia, 800 m X / 1957, 3 ♀ LEC. Other material examined. Chile, Cordillera, Elqui 800 m, 3 not sexed MLPC. Chile: Rivadavia X / 1957, 1 not sexed LEC. Chile only indicates the country MNHN 611 / 1; 1 not sexed.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40479695113ADFD6FFC92F9A4.taxon	diagnosis	Diagnosis. Elytra tri-colored: ochre- yellowish with reddish irregular spot at anteriorly one-third; sutural and marginal black vittae extended from base to the apex, both vittae converging distally. Head surface covered with large, deep and dense punctures; pronotum surface covered with small, shallow, and very sparse punctures. Antennomere I swollen, bead-like, broadest at midlength; antennomere II cup-shaped; antennomere III smoothly divergent distally, with a slight curvature forming a distal lip; antennomeres IV – V similar in shape to III; antennomere VI subtrapezoidal, its proximal third narrower than that of antennomeres VII – IX; antennomeres VII – X subtrapezoidal, with a pronounced proximal constriction and slightly divergent distal margins, broadest distally. Femora shorter than tibia. Metatibiae with spurs similar in length, not parallel but divergent, forming an angle of 20 degrees, outer spur broadened distally, obliquely truncate, and inner spur narrowed, sticklike. Elytra with total length 7.2 mm; subparallel edges (maximum width at proximal two-third: 3 mm); suture dehiscent at distal one-third; elytra narrowed distally. Hindwing veins compositions are summarized in table 2 a. Male genitalia with spiculum gastrale with laterally tapered arms, joined by a central, broad, multidentate emargination; proximally slightly curved to the left; aedeagus with three hooks (Fig. 39).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40479695113ADFD6FFC92F9A4.taxon	description	Redescription. Small- medium sized species; length 8 ‒ 12.5 mm. Coloration. Head and pronotum black; lustrous tegument. Antennae and mouthparts are dark- brown. Clypeus is completely dark- brown or dark- brown with half brown. Legs dark- brown or with femora black, tibiae and tarsus reddish- brown. Scutellum black; lustrous tegument. Elytra tri- colored: ochre- yellowish with reddish irregular spot at anteriorly one-third; sutural and marginal black vittae from base to the apex, both vittae converging distally. Tegument. Head surface covered with large, deep and dense punctures; integument between them luster. Clypeus and labrum with punctures similar to those on frons. Pronotum surface covered with small, shallow, and very sparse punctures; integument between them lustrous or slightly rugose. Elytra surface with three smoothly visible longitudinal costae. Setation. Black, setiform dorsally, piliform ventrally (1.2 times longer than dorsal setae). Clypeus and labrum with setae similar to those on frons. Protibia with ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed silky pubescence at distally two-third. Distally, the protarsus has ventrally homogeneously distributed pads. Head. Subtriangular (long 1.85 mm; wide 1.9 mm; maximum width at occiput) or suborbicular (long 1.8 mm; wide 1.5 mm; maximum width just above eyes); surface with a longitudinal subtle median groove from the vertex to the fronto-clypeal suture. Temple divergent or subparallel; straight or rounded posteriorly, respectively. Frons transverse depressed before suture or suborbicular depressed at middle of frons (between eyes) before suture; slightly depressed at level of the posterior edges of eyes. Eyes flat; moderately broad emarginate; ventral lobe markedly narrowing below (wide 0.7 mm; long 1.1 mm). Clypeus subtrapezoidal, bulged at middle, with a markedly declivity towards the superior edge (wide 0.5 mm; long 1.1 mm). Labrum with moderately rounded lateral edge, ventral edge concave (wide 0.5 mm; long 1 mm). Maxillary palpomere IV subrectangular (maximum width towards both extremes, parallel sides). Labial palpomeres III narrowed toward the distal region (maximum width distally). Mandibles short, markedly curved on the external side, mostly covered by clypeus and labrum while resting. Antennae loosely articulated (Fig. 14). Antennomere I swollen, bead-like, broadest at midlength; antennomere II cup-shaped; antennomere III smoothly divergent distally, with a slight curvature forming a distal lip; antennomeres IV – V similar in shape to III; antennomere VI subtrapezoidal, its proximal third narrower than that of antennomeres VII – IX; antennomeres VII – X subtrapezoidal, with a pronounced proximal constriction and slightly divergent distal margins, broadest distally. Antennomeres length: I 0.20 mm; II 0.10 mm; III 0.37 mm; IV 0.75 mm; VII – X 0.3 mm; XI 0.55 mm. Pronotum. Subhexagonal (long 2.3 mm; wide 2.1 mm) maximum width at apical third, with angulous borders; anterior edge curved; posterior edge almost straight. Legs. Femora shorter than tibia: profemur, protibia (1.99 mm; 2.5 mm respectively), mesofemur, mesotibia (2.9 mm; 3.16 mm respectively); metafemur, metatibia (3.3 mm; 3.6 mm respectively). Metatibiae with spurs similar in length, not parallel but divergent, forming an angle of 20 degrees, outer spur broadened distally, obliquely truncate, and inner spur narrowed, sticklike. Tarsomere measurement: foretarsomere (I 0.66 mm; II 0.46; III 0.33 mm; IV 0.23 mm; V 0.53 mm); mesotarsomere (I 0.93 mm; II 0.66 mm; III 0.699 mm; IV 0.49 mm; and V 0.83 mm); and metatarsomere (I 1.3 mm; II 0. 83 mm; III 0.33 mm; and IV 0.83 mm). Elytra. Total length 7.2 mm; subparallel sides (maximum width at proximal two-third: 3 mm); suture dehiscent at distal one-third; elytra narrowed distally. Hindwing veins compositions are summarized in table 2 a (Fig. 28). Male genitalia (Fig. 39). SpGs forming a plate- like structure with laterally tapered arms, joined by a central, broad, multidentate emargination; proximally slightly curved to the left. Pa incompletely sclerotized, each lobe with an extensive median membranous area. Ph with a medial V- shaped concavity; Pa and Ph fused by a membranous connection (or: membranous tegument). Ae distally with three hooks: two smaller DAH and one very robust VAH. Cerci well- developed, conical. AeA short and straight. Female genitalia. Sclerotized plate of Va subrectangular, internal border with a wide distal concavity and two small proximal concavities; proximally the BsVa rounded and slightly curved inwards; subrectangular Sty as long as Va. Intraspecific Variation. Variation occurs in the body length: small-medium sized species (length: 8 – 12.5 mm).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40479695113ADFD6FFC92F9A4.taxon	distribution	Geographic Distribution. Picnoseus gajardoi presents a wide area of distribution occurring in Monte and Chaco biogeographic provinces in Argentina; and Santiago and Coquimbo biogeographic provinces in Chile (Fig. 68). Habitat in all these biogeographic provinces are arid.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40479695113ADFD6FFC92F9A4.taxon	biology_ecology	Natural History. According to specimen labels, P. gajardoi has been collected in spring (September, October and November); observed on Malvaceae species (Available from https: // www. inaturalist. org / observations / 244844090; date of access April 2025). Presents an altitudinal of 800 m. a. s. l.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40478695313ADF92BFE95FB38.taxon	description	(Fig. 53)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40478695313ADF92BFE95FB38.taxon	materials_examined	Type Material. After several searches in the museum collections, we have not been able to find the type specimen. Probably lost or destroyed. However, we studied voucher specimens deposited at (MLPC), identified by the specialist Denier, and specimens deposited at (LEC), (JEBTC), (IADIZA) and (MZLU) collections. The characters match those of the original description, except for the size differences. Material examined. Chile, Carretera: Algarrobal, III Región, I / X / 1987, Col. J. E. Barriga-Tuñón, 1 ♂ LEC. Chile, Algarrobal: Huasco, 11 / IX / 84, Col. J. E. Barriga-Tuñón, 4 ♂ LEC. Chile, Castilla: Copiapó, 13 / IX / 84, Col. J. E. Barriga-Tuñón 1 ♀ LEC. Chile: Caldera 15 / IX / 84, Copiapó, Col. J. E. Barriga-Tuñón, 2 ♂ LEC. Chile: Algarrobal, J. E Barriga, 01 / X / 1987, 2 ♀ JEBTC and 1 ♀ LEC. Chile: Huasco, Coll. J. E Barriga, 01 / X / 1987, 1 ♀ LEC. Chile, [Copiapó: Minas, Santa Dominga, Coll. Sergio Roig, X / XI / 1992, 3 ♀ IADIZA. Chile, Melipilla, Laguna Esmeralda, Coll. F. Ramírez, 26 - X- 2010, 1 ♀ JEBTC. Chile, Huasco Algarrobal, 15 / IX / 1984 Leg. J. E. Barriga, Coll. J. E. Barrigal, 1 ♀ JEBTC. Chile, Copiapó: Playa Totoralillo, Coll. Sergio Roig, X ‒ 1992, 1 ex IADIZA. Chile, Copiapó, prov. W. Copiapó, 3 / XII / 1999, leg. J. E. Barriga, Picnoseus laevigatus, Denier, MZLU-COL: 00008237. Other material examined. Chile, Algarrobal: Huasco, 11 / IX / 84, Col. J. E. Barriga, 1 ex LEC. Chile, Copiapó, Caldera 15 sep. 1984, Leg. J. E. Barriga, Coll. J. E. Barriga, 2 exx JEBTC. Chile, Huasco: Algarrobal, 26 / XI / 1984, Leg. J. E. Barriga, Coll. J. E. Barriga, 1 ex JEBTC. Chile: Elqui, 5 km SW Charos Bajos, 14 oct. 2005, Leg. J. E. Barriga, 6 exx JEBTC. Chile, Copiapó: Minas Sta. Dominga, X / 1992 S. Roig, 3 exx IADIZA. Chile Copiapó: Playa Totoralillo, X / 1992 S. Roig, 1 ex IADIZA. Chile, Copiapó: Caldera, X / 1992 S. Roig, 1 ex IADIZA. Chile, II Región, Chañaral Calera, X / 1992 S. Roig, 3 exx IADIZA.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40478695313ADF92BFE95FB38.taxon	diagnosis	Diagnosis. Elytra black with yellow irregular spot at anterior one-third that continued as a wide marginal vittae towards the apex. Head and pronotum surface covered with medium, shallow, and dense punctures. Setation black or brown; short and erect dorsally; elongate ventrally. Antennomere I proximally narrowed, broadest distally; antennomere II cup-shaped; antennomere III progressively divergent from base to apex; antennomeres IV – VII smoothly divergent distally; antennomeres VIII – X parallel-sided (apex and base of equal width). Profemur with the same length than protibia, mesofemur and metafemur shorter than mesotibiae and metatibiae. Elytra with total length 5.5 mm; with sinuous sides (maximum width at posterior two-third); suture dehiscent apically. Hindwings with reduced venation, see table 2 a. Male genitalia with spiculum gastrale with four acute spiniform projections joined by a concavity; median concavity broad and U-shaped, narrowing proximally and becoming slightly curved to the right; aedeagus with two hooks (Fig. 40).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40478695313ADF92BFE95FB38.taxon	description	Redescription. Small- medium sized species; length 9 – 12 mm. Coloration. Head, pronotum, and abdomen black, dark- brown or brown; lustrous tegument. Antennae, mouthparts and legs dark- brown, reddish or brown. Clypeus dark- brown or brown. Elytra black or dark- brown with yellow irregular spot at anterior one-third that continued as a wide marginal vittae towards the apex. Scutellum black, or pale- brown with dark-brown on margins; lustrous tegument. Tegument. Head and pronotum surface covered with medium, shallow, and dense punctures. Head integument between punctures micro- reticulate. Pronotum integument between punctures micro- reticulate or slightly rugose. Clypeus punctures similar to those on frons, almost absent in its distal third, which is membranous and glabrous. Labrum punctures smaller than those on clypeus. Elytra without longitudinal visible costae on disk. Scutellum with puncture similar to those on pronotum. Setation. Black or brown; short and erect dorsally; elongate ventrally. Clypeus and labrum with very sparse setae, thinner than those on head. Protibia with ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed silky pubescence at distally two-third. Ventral pads of protarsus are evenly distributed throughout its length. Head. Suborbicular (length 1.5 mm, 1.7 mm; maximum width just above eyes); surface with a longitudinal subtle median groove from the vertex to the fronto-clypeal suture. Temple subparallel, rounded posteriorly. Frons transversely depressed before clypeal suture or with a suborbicular depression on frons (between eyes before clypeal suture). Eyes swollen; anteriorly slightly emarginated in front of antenal insertion; ventral lobe abruptly narrowing below (wide 0.6 mm; long 0.9 mm). Clypeus transverse, slightly bulged (wide 0.5 mm; long 1 mm). Labrum surface with a median depression; lateral edge moderately rounded, their surface smoothly curved inward; ventral edge concave (wide 0.5 mm; long 1.1 mm). Maxillary palpomere IV subrectangular, widest at distal two-third, proximally narrowed; distally with a slight anterodorsal curvature (difficult to see). Labial palpomere subquadrate (with the same width on both extreme). Mandible short, markedly curved on the external sides, covered by clypeus and labrum while resting. Antennae loosely articulated (Fig. 15). Antennomere I proximally narrowed, broadest distally; antennomere II cup-shaped; antennomere III progressively divergent from base to apex; antennomeres IV – VII smoothly divergent distally; antennomeres VIII – X parallel-sided (apex and base of equal width). Antennomeres length: male: I 0.25 mm; II 0.1 mm; III 0.4 mm; IV 0.30 mm; V 0.25 mm; VI 0.27 mm; VII 0.30 mm; VIII – X 0.37 mm; XI 0.5 mm. Female: I 0.37 mm; II 0.10 mm; III 0.25 mm; IV – VI 0.30; VII 0.46 mm; VIII ‒ X 0.49; XI 0.73 mm. Pronotum. Suborbicular (long 1.8 mm; wide 2 mm); maximum width at apical third, with angulous borders; surface with two suborbicular depressions on the posterior third and one suborbicular depression at the middle of the basal edge. Legs. Profemur with the same length than protibia (2.6 mm; 2.5 mm respectively); mesofemur and metafemur shorter than mesotibiae and metatibiae (2.9 mm, 3.3 mm and 3.6 mm, 4.1 mm respectively). Metatibiae spurs dissimilar, inner spur broad expanded, rotated inwards, outer spur obliquely truncate; not parallel but divergent, forming an angle of 20 degrees. Tarsomere measurement: foretarsomere (I 0.49 mm; II and III 0.39; IV 0.23 mm; V 0.66 mm); mesotarsomere (I 0.99 mm; II 0.49 mm; III and IV 0.39 mm; and V 0.66 mm); and metatarsomere (I 1.3 mm; II 0. 66 mm; III 0.49 mm; and IV 0.89 mm). Elytra. Total length 5.5 mm; with sinuous sides (maximum width at posterior two-third); suture dehiscent apically. Hindwings with reduced venation, see table 2 a (Fig. 24). Male genitalia (Fig. 40). SpGs with four acute spiniform projections joined by a concavity; median concavity broad and U-shaped, narrowing proximally and becoming slightly curved to the right. Pa distally with elongate apical setae; surface incompletely sclerotized, each lobe with an extensive membranous area except for a Y shaped sclerotized region. This region consists of a narrow median strip that diverges distally and runs along the ventromedial surface of each lobe. Ph with a medial U - shaped concavity; Pa and Ph fused ventrally by a marginal sclerotized band. Ae distally with two hooks: one DAH and one robust VAH. AeA short and slightly curved. Female genitalia. Sclerotized pale of Va subtriangular, external border with proximal spine, internal border with two spines; proximally elongate and straight BsVa; suborbicular Sty as lon as Va. Intraspecific variations. Variation occurs in the body length: small- medium sized species (length: 9 – 12 mm); tegument coloration: head, pronotum, and abdomen varying from black to dark- brown or brown, always lustrous; and the elytra macula and vittae varying from black to dark- brown.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40478695313ADF92BFE95FB38.taxon	distribution	Geographic Distribution. Picnoseus laevigatus presents a wide area of distribution occurring in Monte biogeogeographic province in Argentina; and Santiago, Coquimbo and Atacama biogeographic provinces in Chile (Fig. 69). Habitat in all these biogeographic provinces are arid.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40478695313ADF92BFE95FB38.taxon	biology_ecology	Natural History. According to specimen labels and the information obtained from iNaturalist platform P. laevigatus was observed in September, October and November visiting flowers of Cristaria sp. (Malvaceae) eating pollen (Available from https: // www. inaturalist. org / observations / 55749121; date of access April 2025) and on flowers of Nolana spp. (Solanaceae) (Available from https: // www. inaturalist. org / observations / 135990404; date of access April 2025).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047A695E13ADFAD3FE39F85D.taxon	description	(Fig. 54)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047A695E13ADFAD3FE39F85D.taxon	materials_examined	Type Material. Lectotype herein designed. Printed label Museum Paris Chili, F. Videla, C. E. Porter 1912. White labeled P. xanthopterus var. nova marginalis Denier, 1918. Red printed labeled SYNTYPE. White printed labeled MNHN-EC 9080. Paralectotypes, four sex undetermined, same data as Lectotype MNHN-EC 9081 ‒ 84. Examined. Material examined. Chile, Malleco: Lonquimay, Coll. J. E. Barriga, 22 / XII / 1994, 1 ♂ LEC. Chile, Malleco: Lonquimay, Coll. J. E. Barriga, 22 / XII / 1994, 1 ♂ JEBTC. Chile: Huasco, Coll. J. E. Barriga 1 / X / 1987 1 ♂ JEBTC. Chile: Algarrobal, Coll. J. E. Barriga 1 / X / 1987, 1 ♂ JEBC. Chile, Valle de Aconcagua: Los Andes, 3 ♂ LEC. Chile: Coquimbo, Playa Totoralillo, Coll. S Roig. X / 1999, 3 ♂ IADIZA. Chile, Atacama: Caldera, Coll. J. E. Barriga, 15 / IX / 84, 2 ♀ JEBTC. Chile: Santiago de Chile, Coll. J. E. Barriga, 15 / IX / 841 ♀ JEBTC. Chile: Bio bio, Coll. F. Ruiz, I / 1918, 1 ♂ LEC. Chile, Parque del Inglés Radal Chile Curicó].	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047A695E13ADFAD3FE39F85D.taxon	diagnosis	Diagnosis. Head and pronotum, black or dark-reddish; elytra black with reddish irregular spot at basal one-third that continued as marginal vittae towards the apex. Head and pronotum surface covered with large, shallow, and dense punctures, but isolated from each other. Head integument between punctures micro- reticulate. Pronotum integument between punctures lustrous. Setation elongate, black or yellowish. Antennomere I globose (bead-like), constricted proximally; antennomere III and IV gradually divergent distally; antennomere VI divergent distally; antennomeres VII – X subtrapezoidal. Forefemur shorter than foretibiae; mesofemur and metafemur shorter than mesotibiae and metatibiae. Elytra with total length 6 mm, with parallel sides (maximum width at anterior two-third: 3 mm), elytra convergent towards posterior one-third. Hindwing veins compositions are summarized in table 2 b. Male genitalia with spiculum gastrale with a distally multi- dentate border; proximally narrow and slightly dextrally curved; aedeagus with two hooks (Fig. 41).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047A695E13ADFAD3FE39F85D.taxon	description	Redescription. Small- medium sized species; length 9 – 12 mm. Coloration. Head and pronotum, black or dark-reddish, lustrous tegument. Elytra black with reddish irregular spot at basal one-third that continued as marginal vittae towards the apex. Scutellum black or brown, lustrous tegument. Scutellum vestiture pale- brown mixed with dark- brown setae. Tegument. Head and pronotum surface covered with large, shallow, and dense punctures, but isolated from each other. Head integument between punctures micro- reticulate. Pronotum integument between punctures is lustrous. Clypeus and labrum with similar punctures, small, uniformly distributed, and less dense than those on frons; clypeus with inferior third membranous. Elytra without visible longitudinal costae. Scutellum surface with longitudinal median groove or not; puncture similar to those on pronotum. Setation. Black or yellowish; elongate. Labrum bearing elongate setae laterally. Male and female protibiae with ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed pubescence at distal two-third; male protarsus with ventral pads homogeneously distributed; female protarsus with ventral pads confined to the distal portion, proximal surface covered with sparse, thickened setae. Scutellum vestiture setiform. Head. Subtriangular (length 1.6 mm; width 2 mm; maximum width at occiput) or suborbicular (length 1.6 mm; width 1.9 mm; maximum width just above eyes); surface with a longitudinal median groove from the vertex to the upper area of fronto-clypeal suture. Temple divergent, straight posteriorly; or subparallel, curved posteriorly. Frons transversely depressed before clypeal suture. Eyes flat; ventral lobe abruptly tapering below (wide 0.6 mm; long 1.1 mm). Clypeus transversely, slightly bulged (wide 0.5 mm; long 1.1 mm). Labrum surface depressed at center; with moderately rounded lateral edge, ventral edge straight (wide 0.5 mm; long 1 mm). Maxillary palpomere IV subrectangular (with subparallel sides). Labial palpomeres III fusiform (maximum width at middle). Mandibles short, markedly broad curved on the external side, mostly covered by clypeus and labrum while resting; or elongate, broad curved on the external side, partially covered by clypeus and labrum when at rest. Antennae loosely articulated (Fig. 16). Antennomere I globose (bead-like), constricted proximally; antennomere III and IV gradually divergent distally; antennomere VI divergent distally; antennomeres VII – X subtrapezoidal. Antennomeres length: male I 0.25 mm; II 0.10 mm; III 0.37 mm; IV 0.30 mm; V 0.25 mm; VI 0.27 mm; VII 0.30 mm; VIII ‒ X 0.37 mm; XI 0.50 mm. Female I 0.37 mm; II 0.10 mm; III 0.25 mm; IV ‒ VI 0.30 mm; VII 0.35 mm; VIII ‒ X 0.37 mm; XI 0.55 mm. Pronotum. Subhexagonal (long 2 mm; wide 2.3 mm) maximum width at anterior two-third, with angulous borders. Legs. Forefemur shorter than foretibiae (2.9 mm; 3.3 mm, respectively); mesofemur shorter than mesotibiae (3.4 mm; 3.6 mm, respectively); metafemur shorter than metatibiae (3.9 mm; 4.3 mm, respectively). Metatibiae with spurs similar in length, not parallel but divergent, forming an angle of 20 degrees; both spurs slightly expanded, flattened, bluntly rounded. Tarsomere measurement: foretarsomere (I 0.76 mm; II and III 0.49; IV 0.399 mm; V 0.83 mm); mesotarsomere (I 0.99 mm; II 0.66 mm; III and IV 0.49 mm; V 0.83 mm); and metatarsomere (I 1.33 mm; II 0. 99 mm; III 0.66 mm; IV 0.86 mm). Elytra. Total length 6 mm, with sides parallel (maximum width at anterior two-third: 3 mm), converging distally, suture dehiscent apically; elytra convergent towards posterior one-third. Hindwing veins compositions are summarized in table 2 b (Fig. 29). Male genitalia (Fig. 41). SpGs with a distally multi- dentate border; proximally narrow and slightly dextrally curved. Pa incompletely sclerotized, each lobe with an extensive membranous area. Ph with a deep, medial, Ushaped concavity; Pa and Ph sclerotized fused. Ae distally with two robust hooks: one DAH and one robust VAH. AeA short, straigth. Female genitalia. Sclerotized plate of Va subrectangular, internal border with a median small concavity; proximally the BsVa is elongated, narrow proximally pointed, and curved outward; suborbicular Sty a half smaller than Va. Sexual Dimorphism. Occurs in forelegs: male protarsus with ventral pads homogeneously distributed; female protarsus with ventral pads confined to the proximal portion, distal surface covered with thicker sparse setae.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047A695E13ADFAD3FE39F85D.taxon	distribution	Geographic Distribution. Endemic of Chile, in the Maule, Santiago and Coquimbo biogeographic provinces (Fig. 69).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047A695E13ADFAD3FE39F85D.taxon	discussion	Taxonomic Comments. Previously allocated as a variety of P. xanthopterus by Denier (1918). After examination of the types, we concluded that differences in the tegument color and shape of male genitalia supports that P. marginalis and P. xanthopterus are different species.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4047A695E13ADFAD3FE39F85D.taxon	biology_ecology	Natural History. According to specimen labels and the information obtained from iNaturalist platform adults are active from September to January (between spring and summer). Picnoseus marginalis was observed feeding flowers of Sphaeralcea spp. (Solanaceae) (Available from https: // www. inaturalist. org / observations / 258700046; date of access April 2025).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40476695F13ADFF7FFC80F82B.taxon	description	(Fig. 55)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40476695F13ADFF7FFC80F82B.taxon	materials_examined	Type Material. Holotype ♀. White hand writer, C. Bruch, dedit. 1932, Nonogasta: La Rioja Gomez. Red label: Lytta nigropicta, Denier 1932, Holotipo, MLPC. Paratype sex?. White hand writer, Nonogasta: La Rioja Gomez, Cotypus, Paratypus, Lytta nigropicta, Denier 1932 MACN. La Cienaga: Belén, Catamarca, leg. Weiser Wolters 1992, Cotype, Lectotype ♂ Des. R. B. Selander 1988, Fichado, MACN. Examined. Denier (1932) indicates that four cotypes are deposited in Bruch collection at (MACN) collection. After several searches in the museum collection we have not been able to find the complete series of type specimens. We find one male identified by the specialist Denier (1932). The other specimens are probably lost.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40476695F13ADFF7FFC80F82B.taxon	diagnosis	Diagnosis. Head, pronotum, abdomen and legs black, lustrous tegument; elytra yellowish-ochre with a wide dark-brown vitta on the disk and a thin dark-brown sutural vitta. Head and pronotum surface covered with small-size, sparse, shallow punctures. Setation grayish; setiform dorsally, piliform ventrally. Forefemora shorter than foretibia; mesofemora shorter than mesotibiae.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40476695F13ADFF7FFC80F82B.taxon	description	Redescription of Holotype. Small- sized species; length 8.5 ‒ 10 mm. Coloration. Head, pronotum, abdomen and legs black, lustrous tegument; elytra reddish with a wide dark-brown vitta on the disk and a thin dark-brown sutural vitta. Scutellum black. Tegument. Head and pronotum surface covered with small-size, sparse, shallow punctures; integument between them luster. Clypeus similar to those on frons. Labrum with punctures smaller than those on clypeus and less dense. Elytra without longitudinal visible costae. Scutellum with small and dense punctures, integument among them luster. Setation. Grayish; setiform dorsally, piliform ventrally. Head. Suborbicular (length 1.6 mm; width 1.5 mm; maximum width just above eyes); surface with a longitudinal subtle median groove from vertex to the upper area of fronto-clypeal. Frons transversely depressed before suture. Temple subparallel, rounded posteriorly. Eyes flat; interior edge straight; inferior lobe abruptly tapering below (wide 0.6 mm; long 0.8 mm). Clypeus flat, subparallel sides (wide 0.4 mm; long 0.8 mm). Labrum with the distal edge is laterally rounded (wide 0.4 mm; long 0.8 mm). Maxillary palpomere IV progressively divergent towards distal region (maximum width at apex). Labial palpomere III fusiform (maximum width at middle). Mandibles short, markedly curved on the external side, covered by clypeus and labrum while resting. Antennae with antennomeres I – V closely articulated, VI – XI loosely articulated (Fig. 17). Antennomere I inflate, bead-like appearance, narrowing towards the proximally region; antennomere II cup-like appearance; antennomere III smoothly divergent towards distally region (maximum width distally); antennomeres IV – X subtrapezoidal. Pronotum. Suborbicular (long 1.5 mm; wide 1.6 mm) maximum width at anterior one-third, with rounded borders; anterior and posterior edges almost straight. Legs. Forefemora shorter than foretibia (2.8 mm; 2.6 mm, respectively); mesofemora shorter than mesotibiae (3.2 mm; 3.5 mm, respectively); metafemur shorter than metatibiae (3.6 mm; 3.9 mm, respectively). Metatibiae with spurs similar in length, not parallel but divergent, forming an angle of 20 degrees, outer spur broadened apically, obliquely truncate, and inner spur narrowed, sticklike. Tarsomere measurement: foretarsomere (I 0.5 mm; II and III 0.4 mm; IV 0.3 mm; and V 0.5 mm); mesotarsomere (I 0.7 mm; II 0.5 mm; III 0.4 mm; IV 0.3 mm; and V 0.7 mm); and metatarsomere (I 1.1 mm; II 0.9 mm; III 0.6 mm; and IV 0.7 mm). Antennomeres length: I 0.20 mm; II 0.10 mm; III 0.35 mm; IV 0.70 mm; VII ‒ X 0.3 mm; XI 0.50 mm. Elytra. Total length 6.5 mm, with subparallel edges (maximum width at proximal two-third: 2.3 mm); suture dehiscent at distal one-third; elytra convergent at distal one-third. Dissection of the specimen was not authorized by the museum; therefore, the hind wings and genitalia were not examined.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40476695F13ADFF7FFC80F82B.taxon	distribution	Geographic Distribution. The species occurs in the Monte biogeographical province, in Argentina; and Coquimbo bioheographical province in Chile (Fig. 69).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40476695F13ADFF7FFC80F82B.taxon	biology_ecology	Natural History. According with the information obtained from iNaturalist platform, P. nigropicta was observed in September feeding flowers and pollen of Cactaceae species (Available from https: // www. inaturalist. org / observations / 138316069; date of access April 2025) (Fig. 62).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40470695A13ADFF7FFD1FFC8C.taxon	description	(Fig. 56)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40470695A13ADFF7FFD1FFC8C.taxon	materials_examined	Type Material. Lectotype sex?, herein designated. Blue printed labeled Museum Paris, Chili, Germain, Coll. Fairmaire 1906. Green hand writer labeled [Cordillera de Maule, Versant Oriental, 8000 pieds. Blue hand writer labeled: Tetraonyx nitidipennis, R. Zol 1860. White hand writer labeled: P. nitidipennis, F and G, P. Denier visit 1918. White printed labeled: TYPE. Red printed labeled: SYNTYPE. White printed labeled, Syntype Picnoseus nitidipennis (Fairmaire & Germain, 1860, MNHN EC 9097. Examined. Material examined. Chile Coquimbo: Guanaqueros, Coll. J. E. Barriga, 18 / XI / 1990, 3 ♂ JEBTC. Chile, Malleco: Lonquimay, Coll. J. E. Barriga, 22 / XII / 1994, 1 ♂ ENTC. Chile, Paipote: Almagro (salida camino internacional), Coll. S. Roig, 21 / V / 1991, 1 ♂ IADIZA. Chile, Malleco: Malalcahuello, Coll J. E. Barriga, 26 / I / 2007, 1 ♂ LEC. Chile: Desierto de Atacama, Coll L. E. Peña, IX / 22 / 63, 3 ♂ LEC. Other material examined. Chile, Curicó: Parque del Inglés: Radal, 28 / XI / 1993, Leg. J. E. Barriga Colección J. E. Barriga. Chile, 3 not sexed JEBTC. Chile, Ñuble: Puente Marchat, 9 / XII / 1989) | Leg. J. E. Barriga, Colección J. E. Barriga, 1 ex JEBTC.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40470695A13ADFF7FFD1FFC8C.taxon	diagnosis	Diagnosis. Head, mouth parts, antennae and pronotum black, lustrous tegument. Legs black, metatarsus I yellowish proximally. Elytra yellowish- ochre. Head surface covered with large, shallow punctures, dens on frons, sparse toward vertex; pronotum surface covered with large, deep, and very sparse punctures. Setation black, piliform, erect. Antennomere I bead-like; antennomere II cup-shaped; antennomeres III – V gradually widening distally, antennomeres III and IV with a distal anteroapical curvature; antennomeres V – X subtrapezoidal, antennomeres VI and VII with smoothly constricted proximal two-thirds. Profemur and protibiae have the same length; protibiae widened toward the apex, slightly curved apically. Metatibia with spurs similar, slightly expanded, flattened and bluntly rounded. Male genitalia with spiculum gastrale bearing two stick- like lateral arms, joined by a broad emargination; proximally narrowed and slightly curved to the left; aedeagus with two hooks (Fig. 42).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40470695A13ADFF7FFD1FFC8C.taxon	description	Redescription. Small to medium sized species; length 6 – 14.5 mm. Coloration. Head, mouth parts, antennae and pronotum black, lustrous tegument. Legs black, metatarsus I yellowish proximally. Elytra yellowish- ochre. Scutellum black, lustrous tegument. Tegument. Head surface covered with large, shallow punctures, dens on frons, sparse toward vertex; or with small, deep and very sparse punctures uniformly distributed; integument between them lustrous. Clypeus with large, sparse punctures (like on frons). Labrum punctures smaller and less dense than those on clypeus. Pronotum surface covered with large, deep, and very sparse punctures; integument between them lustrous. Elytra with three longitudinal visible costae on disk. Scutellum rugose, smooth distally. Setation. Black, piliform, erect. Protibiae with ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed pubescence at distal two-third; protarsus with ventral pads uniformly distributed along its entire length. Head. Suborbicular (length 1.8 mm; width 8 mm; maximum width just above eyes); surface with longitudinal subtle median groove from vertex to the upper area of the fronto-clypeal suture or not. Temple subparallel, straight or curved posteriorly. Frons transversely depressed before clypeal suture or with a suborbicular depression at center (before clypeal suture, between eyes). Eyes flat; with anteriorly slightly emarginate; ventral lobe abruptly narrowing below (wide 0.5 mm; long 0.7 mm). Clypeus flat, with subparallel sides (wide 0.4 mm; long 0.7 mm). Labrum with moderately rounded lateral edge; ventral edge slightly emarginated (wide 0.4 mm; long 0.8 mm). Maxillary palpomere IV subrectangular (maximum width in both extremes, with parallel sides). Labial palpomere subquadrate, narrowing at proximal one third, divergent towards distal region (maximum width distally). Mandible elongate, broad curved on the external side; partially covered by the clypeus and labrum while resting. Antennae loosely articulate (Fig. 18). Antennomere I bead-like; antennomere II cup-shaped; antennomeres III – V gradually widening distally, antennomeres III and IV with a distal anteroapical curvature; antennomeres V – X subtrapezoidal, antennomeres VI and VII with smoothly constricted proximal two-thirds. Antennomere length: I 0.2 mm, II 0.10 mm, III 0.45 mm, IV – X 0.37 mm, XI 0.50 mm. Pronotum. Subhexagonal (wide 1.5 mm; long 1.8 mm), maximum width at anterior third, with angulous borders; surface with a longitudinal subtle median groove on posterior half or without it. Legs. Profemur and protibiae have the same length (2.1 mm); mesofemur and metafemur are shorter than mesotibia and metatibiae (2.5 mm; 2.9 mm respectively). Protibiae widened toward the apex, slightly curved apically. Metatibia with spurs similar, slightly expanded, flattened and bluntly rounded. Tarsomere measurement: foretarsomere (I 0.49 mm; II and III 0.33; IV 0.16 mm; V 0.56 mm); mesotarsomere (I 0.66 mm; II and III 0.33; IV 0.16 mm; and V 0.56 mm); and metatarsomere (I 0.83 mm; II 0.49 mm; III 0.33 mm; and IV 0.56 mm). Elytra. Total length 7.2 mm, with subparallel edges, then convergent (greatest width at posterior two-third: 2.3 mm); suture dehiscent at basal one-third. Hindwing veins compositions are summarized in table 2 b (Fig. 30). Male genitalia (Fig. 42). SpGs bearing two stick- like lateral arms, joined by a broad emargination; proximally narrowed and slightly curved to the left. Pa incompletely sclerotized, with a membranous area; Ph with a medial V- shaped concavity; Pa and Ph fused by a marginal sclerotized area. Ae bears two robust distal hooks: a DAH with a distal peak, and a ventral VAH. AeA elongated and straight. Female genitalia. Unknown.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40470695A13ADFF7FFD1FFC8C.taxon	distribution	Geographic Distribution. This species occurs from subantarctic Maule, central Chile Santiago and Coquimbo biogeographic provinces (Fig. 70). According to Fairmaire & Germain (1863) P. nitidipennis was collected in Chile at 2500 m. a. s. l (Maule).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40470695A13ADFF7FFD1FFC8C.taxon	biology_ecology	Natural History. According to specimen labels and contributions from the iNaturalist platform, P. nitidipennis was recorded in spring and summer (September, October, November, January, and March), observed on grasses and visiting Carpobrotus chilensis (Aizoaceae) eating the flower (Available from https: // ecuador. inaturalist. org / observations / 198360616; date of access April 2025).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40472696413ADFF7FFDF8FB88.taxon	description	(Fig. 57)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40472696413ADFF7FFDF8FB88.taxon	materials_examined	Type Material. Holotype sex? White hand writer labeled Chile: Prov. Santiago, Dep. La Victoria: El Volcán, C. E. Porter leg. Red hand- writer labeled Picnoseus rubrofasciatus, Denier 1934 Holotipo, MLPC 613 / 1 MLPC. Paratype (sex undetermined) White typewriter labeled Totoral, porv. Copiapó, Chile, 28 - XI- 1987, leg. J. E Barriga, MZLU-COL 00008236, MZLU. Examined. Material examined. Chile, Copiapó, Bahía Inglesa, Coll. Barriga J. E, 06 / XI / 1991, 7 ♂; 1 ♀ JEBTC. Chile, Algarrobal, Coll. Barriga J. E, 01 / 10 / 1987, 1 ♀ JEBTC. Chile: Huasco, Coll. Barriga J. E, 01 / 10 / 1987, 1 ♀ LEC. 6 exx JEBTC, Chile, Huasco: Algarrobal, 26 / XI / 1987, Leg. J. E. Barriga. Other material examined. 2 exx JEBTC, Chile, Huasco: Algarrobal, 15 / XI / 1984, Leg. J. E. Barriga. 2 exx JEBTC, Chile, Copiapó, Totoral 28 / XI / 1987, Leg. J. E. Barriga. 3 exx JEBTC, Chile: Vallenar Algarrobal, 1 / X / 1987, Leg. J. E. Barriga, Colección J. E. Barriga. 3 exx JEBTC, date is not indicated Chile III Región, Canto del Agua, Leg. J. E. Barriga. 1 ex JEBTC, Chile: Huasco Freirina, Leg. J. E. Barriga, IX / 1984. 2 exx JEBTC Huasco: Quebrada Honda, N Carriza Bajo, 30 / XI / 1987, Leg. J. E. Barriga. 1 ex JEBTC, Chile, Copiapó, Totoral 30 / XI / 1987, Leg. J. E. Barriga. 2 ex JEBTC, Chile: Huasco 10 km, N Huasco, 18 / IX / 1991, Leg. J. E. Barriga. 3 exx MEL, Chile: Atacama, 16 km N. Vallenar, 20 / X / 1980, L. Peña det. Lytta unknown det. Picnoseus sp. by Pinto 2006.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40472696413ADFF7FFDF8FB88.taxon	diagnosis	Diagnosis. Head, pronotum and abdomen black. Legs with femora black, tibiae and tarsomere dark-reddish. Elytra is reddish with a broad black sutural vitta extending from the base to near the apex. Head and pronotum surface covered with large, shallow, and sparse punctures. Setation black, piliform, and erect. Antennomere I broadened distally, distinctly constricted proximally; antennomere II cup-shaped; antennomere III gradually widening distally, with an anteroapical curvature; antennomere IV broader posteriorly; antennomeres V – X subtriangular. Femur and tibiae I and II with the same length; metafemur shorter than metatibiae. Metatibiae spurs are similar, slightly expanded, flattened and bluntly rounded. Male genitalias with spiculum gastrale as in the figure 43; aedeagus with two robust hooks (Fig. 43).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40472696413ADFF7FFDF8FB88.taxon	description	Redescription. Medium- sized species; length 11 – 15.5 mm. Coloration. Head, pronotum and abdomen black, lustrous tegument. Antennae black. Legs with femora black, tibiae and tarsomere dark-reddish. Elytra reddish with a broad black sutural vitta extending from the base to near the apex. Tegument. Head and pronotum surface covered with large, shallow, and sparse punctures. Head with integument between punctures, luster or rugose. Pronotum with integument between punctures rugose. Clypeus and labrum punctures similar in both, small and very sparse (smaller than those on frons). Scutellum surface with very small punctures (smaller than those on pronotum) and homogeneously distributed, except on both sides of the median groove; integument between punctures lustrous. Setation. Black, piliform, erect. Clypeus and labrum setae similar to those on frons. Protibiae with ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed pubescence at distally two-third; male protarsus with ventral pads homogeneously distributed; female protarsus with ventral pads confined to the distal portion, proximal surface covered with thicker setae. Head. Suborbicular (length 1.6 mm; width 2.3 mm; maximum width just above the eyes); surface with a longitudinal median groove from the vertex to the upper area of fronto-clypeal suture. Frons transversely depressed. Temple subparallel, straight posteriorly. Eye swollen; anteriorly slightly emarginate; inferior lobe abruptly tapering below (wide 0.6 mm; long 1 mm). Clypeus subtrapezoidal, bulged at middle, with a markedly declivity towards the superior edge (wide 0.4 mm; long 1.2 mm). Labrum with moderately rounded lateral edge; ventral edge almost straight (wide 0.4 mm; long 1.2 mm). Maxillary palpomere IV subrectangular (maximum width in both extremes, with parallel sides). Labial palpomeres III fusiform (maximum width at middle). Mandible robust, broad curved externally, partially covered by clypeus and labrum while resting. Antennae loosely articulated (Fig. 19). Antennomere I broadened distally, distinctly constricted proximally; antennomere II cup-shaped; antennomere III gradually widening distally, with an anteroapical curvature; antennomere IV broader posteriorly; antennomeres V – X subtriangular. Antennomere length: male I 0.25 mm; II 0.12; III 0.50 mm; IV 0.37 mm; V – VII 0.32 mm; VIII – X 0.37 mm; XI 0.67 mm. Female I 0.20 mm; II 0.10 mm; 0.37 mm; IV and V 0.22 mm; VI and VII 0.25 mm; VIII 0.32 mm; IX and X 0.37 mm; XI 0.67 mm. Pronotum. Subhexagonal (wide 2 mm; long 2.4 mm) maximum width at anterior one third, with rounded or angulous borders; surface with longitudinal median groove or not. Scutellum totally or partially covered by pronotum. Legs. Femur and tibiae I and II with the same length (2.1 mm; 3.3 mm respectively); metafemur shorter than metatibiae (3.4 mm; 4.3 mm respectively). Metatibiae spurs subparallel, slightly expanded, flattened and bluntly rounded. Tarsomere measurement: foretarsomere (I 0.66 mm; II 0.66 mm; III and IV 0.49 mm; V 0.66 mm); mesotarsomere (I 0.99 mm; II and III 0.33 mm; IV 0.16 mm; and V 0.66 mm); and metatarsomere (I 1.33 mm; II 0. 83 mm; III 0.66 mm; and IV 0.99 mm). Elytra. Total length 9.5 mm, with markedly divergent edges (greatest width at posterior two-third: 4 mm); suture dehiscent distally. Hindwing veins compositions are summarized in table 2 b (Fig. 31). Male genitalia (Fig. 43). SpGs as in figure 43. Pa incompletely sclerotized, with a membranous area except for a Y- shaped sclerotized region. Ph with a medial U- shaped concavity; Pa and Ph fused by a marginal sclerotized band. Ae with two robust hooks: one DAH and one VAH. AeA straight. Female genitalia. Sclerotized plate of Va subtriangular; proximally the BsVa is pointed with an external and internal protuberance; external border with a square- shaped concavity; subrectangular Sty 1.2 x times longer than Va. Sexual dimorphism. Occurs in: male protarsus with ventral pads homogeneously distributed; female protarsus with ventral pads confined to the distal portion, proximal surface covered with thicker setae.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40472696413ADFF7FFDF8FB88.taxon	distribution	Geographic Distribution. This species occurs in Santiago and Coquimbo biogeographic provinces in Chile (Fig. 70).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40472696413ADFF7FFDF8FB88.taxon	biology_ecology	Natural History. According to specimen labels and contributions from the iNaturalist platform, P. rubrofasciatus was observed in spring (September, October, and November). The specimen was photographed on leaves of an undetermined plant (Available from https: // www. inaturalist. org / observations / 30898406; date of access April 2025).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F40472696413ADFF7FFDF8FB88.taxon	discussion	Remarks. One specimen examined shows one of their elytres less developed and shorter than the other. This specimen could be teratological.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044D696513ADFB4BFAD6FA4C.taxon	description	urn: lsid: zoobank. org: act: 0 DEBEB 33 - 7062 - 4 C 53 - B 9 E 6 - 768 C 78144 A 27 (Fig. 58)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044D696513ADFB4BFAD6FA4C.taxon	materials_examined	Type Material. Holotype. Picnoseus roigjunienti, Chile, Copiapó: Playa Totoralillo, X / 1992 S. Roig, IADIZA. Paratype 1 ♂ Chile, Copiapó: Playa Totoralillo, X / 92 S. Roig. 1 ♂ Chile, II Región: Chañaral, X / 1992. 1 ♂ Chile; II Región, Caldera, X / 1992, 2 ♂ Chile: Copiapó 90 km antes, S. Roig X / 1992, IADIZA. Material examined. Chile, Algarrobal: Huasco 11 – 16 / IX / 1984. Col. J. E Barriga-Tuñón; 7 ♂ LEC.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044D696513ADFB4BFAD6FA4C.taxon	diagnosis	Diagnosis. Small- medium sized species. Head, pronotum, abdomen and legs dark- brown. Metatarsomere I yellow-ochre on basal third. Elytra color, yellow-ochre with dark-brown vitta on disk from base to near apex; the vitta is narrow in the posterior two-third, widened in the anterior one-third. Metatarsomere with tarsus I yellowish-ochre on basal third. Head with medium-sized, dense, shallow punctures, but isolated from each other; pronotum with medium- sized punctures, sparse, shallow. Setae black and setiform. Antennomere I cup-shaped, broadest distally and narrowing proximally (distal width about two-thirds of total length); antennomere II cup-shaped; antennomeres III – V divergent distally; antennomeres VIII – X subrectangular. Profemur with the same length as protibia; mesofemur slightly longer than mesotibia; metafemur shorter than metatibiae. Metatibiae spurs subparallel in length, outer spur broadened apically obliquely truncate, inner spur narrowed, sticklike. Hindwing exhibiting with reduced venation, see table 2 a. Male genitalia as in Fig. 44.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044D696513ADFB4BFAD6FA4C.taxon	description	Description of Holotype. Small- medium sized species; length 9 – 12 mm. Coloration. Head, pronotum, abdomen and legs dark- brown. Metatarsomere I yellow- ochre on basal third. Elytra black with pale-brown vittae on disk from anterior to near posterior region; the vittae is narrow in the proximal third, widened in the distal two-third. Tegument. Head with medium-sized, dense, shallow punctures, but isolated from each other; integument among them micro-reticulate. Pronotum with medium- sized punctures, sparse, shallow; integument among them rugose. Clypeus and labrum punctures similar to those on frons. Elytra with three slightly marked longitudinal costae on disk; sutural costae longitudinal extended from base to the apex. Scutellum with small punctures (smaller than those on pronotum), integument among them lustrous. Setation. Black and setiform. Protibiae with apical half of ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed pubescence, protarsus with ventral pads confined to the apical portion, basal surface subglabrous. Head. Suborbicular (length 1.9 mm; width 2.4 mm; maximum width just above eyes); surface with a longitudinal median groove from vertex to the upper area of fronto-clypeal suture. Frons transversely depressed before clypeal suture. Temple subparallel, straight posteriorly. Eye flat; inferior lobe rounded distally (wide 0.9 mm; long 1.1 mm). Clypeus flat, subrectangular, subparallel sides (wide 0.5 mm; long 1.4 mm). Labrum with rounded lateral edges; ventrally slightly concave (wide 0.5 mm; long 1.2 mm). Maxillary palpomere IV divergent from proximal to the distal region (maximum width distally). Labial palpomere III subrectangular (with the same width in both extremes). Mandibles elongate, robust, almost straighter externally, partially covered by the clypeus and labrum while resting. Antennae closely articulated (Fig. 20). Antennomere I cup-shaped, broadest distally and narrowing proximally (distal width about two-thirds of total length); antennomere II cup-shaped; antennomeres III – V divergent distally; antennomeres VIII – X subrectangular. Antennomere length: I 0.5 mm; II 0.12 mm; III – X 0.37 mm; XI 0.65 mm. Pronotum. Suborbicular (2 mm long; 2.3 mm width; maximum width at anterior two-third); surface with longitudinal median groove from anterior to the posterior region. Legs. Profemur with the same length as protibia (2.9 mm); mesofemur slightly longer than mesotibia (3.3 mm; 3.4 mm respectively); metafemur shorter than metatibiae (3.6 mm; 3.9 mm respectively). Metatibiae spurs subparallel in length, outer spur broadened apically obliquely truncate, inner spur narrowed, sticklike. Tarsomere measurement: protarsomere (I 0.6 mm; II and III 0.4 mm; IV 0.3 mm; V 0.6 mm); mesotarsomere (I 0.8 mm; II and III 0.3 mm; IV 0.3 mm; V 0.8 mm); and metatarsomere (I 1.1 mm; II 0.8 mm; III 0.4 mm; IV 0.6 mm). Elytra. Total length 7.5 mm, greatest width at posterior two-third; suture dehiscent from anterior third. Hindwing exhibiting venation reduction, see table 2 a (Fig. 24). Male genitalia (Fig. 44). SpGs with lateral spiniform expansions, joined by a broad median concavity bearing two small central spines, narrowing proximally and becoming straight. Pa incompletely sclerotized, with a membranous area except for a Y- shaped sclerotized region. Ph with a medial V- shaped concavity; Pa and Ph fused sclerotized marginally. Ae distally with three hooks: two DAH and one VAH. AeA elongated and straight. Female genitalia. Unknown.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044D696513ADFB4BFAD6FA4C.taxon	etymology	Etymology. This species is dedicated to Sergio Alberto Roig-Juñent, a cherished friend, in recognition of his passion for coleopteran and his constant support since the beginning of my academic studies.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044D696513ADFB4BFAD6FA4C.taxon	distribution	Geographic Distribution. This species occurs in Santiago and Coquimbo biogeographic provinces in Chile (Fig. 71).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044D696513ADFB4BFAD6FA4C.taxon	biology_ecology	Natural History. According to the specimen labels P. roigjunienti have been collected in spring (September and October). Adults of these species were observed visiting flowers of Crisatria spp. (Malvaceae) (Fig. 63).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044C696713ADFA0FFB41FD20.taxon	description	urn: lsid: zoobank. org: act: 9 BF 3 EF 62 - 298 E- 4085 - 9818 - 72 B 8 B 013 FC 4 F (Fig. 59)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044C696713ADFA0FFB41FD20.taxon	materials_examined	Type Material. Holotype ♂ Chile: Caldera 15 / IX / 1984, III Región, Col. J. E. Barriga-Tuñón, LEC. Paratype 3 ♂ Chile: Castilla Copiapó, 13 / IX / 1984, Col. J. E Barriga-Tuñón, LEC. 3 ♂ Chile, Algarrobal: Huasco 11 / IX / 1984, Col. J. E. Barriga-Tuñón, LEC. Material examined. 1 ♂ Chile, Copiapó: Playa Totoralillo IX / 1992, S. Roig, IADIZA. 1 ♂, country labeled only, Chile MLPC.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044C696713ADFA0FFB41FD20.taxon	diagnosis	Diagnosis. Small to medium- sized species. Head, pronotum, and abdomen black; antennae reddish. Legs with femora and tibiae dark-reddish; tarsomere reddish. Elytra black each with: broad yellow discal vitta extending continuously from humeral region to posterior edge, and complete marginal yellow vitta running parallel to lateral edge from anterior to posterior; both vittae converging distally. Setation black, piliform. Antennomere I suboval, broadest distally, with an anteroapical curvature overhanging antennomere II; antennomere II cup-shaped; antennomeres III – V divergent from base to apex, broadest at the distal third; antennomeres VI – X subrectangular. Femur and tibiae I and II with the same length; metafemur slightly shorter than metatibiae. Metatibia with spurs dissimilar, outer spur truncate apically, inner spur spiniform with apex broadened. Hindwing veins compositions are summarized in table 2 b. Male genitalia with spiculum gastrale progressively divergent distally (border irregular); proximally narrowed slightly curved leftward; aedeagus with two hooks (Fig. 45).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044C696713ADFA0FFB41FD20.taxon	description	Description of Holotype. Small to medium- sized species; length 9 – 12 mm. Coloration. Head, pronotum, and abdomen black, lustrous tegument. Antennae reddish. Legs with femora and tibiae dark- reddish; Tarsomere reddish. Elytra black each with: broad yellow discal vitta extending continuously from humeral region to posterior margin, and complete marginal yellow vitta running parallel to lateral edge from anterior to posterior; both vittae converging distally. Sutural vitta orbicularly expanded in anterior third, forming a large macula that gradually tapers posteriorly. Tegument. Head and pronotum with big, dense and shallow punctures, but isolated from each other; integument among them luster. Clypeus and labrum with dense punctures but smaller than those on frons. Elytra with imperceptible longitudinal costae on disk. Scutellum punctures similar to those on labrum. Setation. Black and piliform. Protibiae with apical half of ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed pubescence; protarsus with ventral pads confined to the distal portion, proximal surface covered with thicker setae. Head. Suborbicular (long: 1.9 mm; width 2.3 mm; maximum width just above eyes); surface with a longitudinal median groove from vertex to the upper area of fronto-clypeal suture. Frons transversely depressed before clypeal suture. Temple subparallel, straight posteriorly. Eye flat; anteriorly slightly emarginate; inferior lobe markedly narrow distally (wide 0.6 mm; long 1 mm). Clypeus slightly bulged, subrectangular, with subparallel sides (wide 0.5 mm; long 1.1 mm). Labrum smoothly curved inwards, with rounded lateral edges, ventrally slightly concave (wide 0.5 mm; long 1.2 mm). Maxillary palpomere IV subtrapezoidal (maximum width towards distal region), with apicoventral curvature. Labial palpomere III subrectangular (with the maximum width in both extremes). Mandibles elongate, robust, almost straighter on the external side; partially covered by the clypeus and labrum when at rest. Antennae with antennomeres I – VII loosely articulated, antennomeres VIII – XI closely articulated (Fig. 21). Antennomere I suboval, broadest distally, with an anteroapical curvature overhanging antennomere II; antennomere II cup-shaped; antennomeres III – V divergent from base to apex, broadest at the distal third; antennomeres VI – X subrectangular. Antennomere length: I 0.25 mm; II 0.10 mm; III 0.52 mm; IV ‒ X 0.37 mm; XI 0.85 mm. Pronotum. Subhexagonal (long 1.8 mm; width 2 mm), maximum width at apical third, with angulous borders; anterior and posterior edges almost straight. Legs. Femur and tibiae I and II with the same length (2.3 mm; 2.6 mm respectively); metafemur slightly shorter than metatibiae (2.9 mm; 3.1 mm respectively). Metatibia with spurs dissimilar, outer spur truncate apically, inner spur spiniform with apex broadened. Tarsomere measurement: foretarsomere (I 0.5 mm; II – IV 0.3 mm; V 0.6 mm); mesotarsomere (I 0.6 mm; II – IV 0.4; V 0.6 mm); metatarsomere (I 1.1 mm; II 0.6 mm; III 0.4 mm; 0.8 mm). Elytra. Total length 7.5 mm (greatest width at posterior two-third: 4 mm). Hindwing veins compositions are summarized in table 2 b (Fig. 34). Male genitalia (Fig. 45). SpGs progressively divergent distally (border irregular); proximally narrowed slightly curved leftward. Pa incompletely sclerotized, with a membranous area. Ph with a medial U- shaped concavity; Pa and Ph fused by a marginal sclerotized band. Ae distally with two robust hooks: a DAH and a markedly more robust VAH. AeA elongate and curved. Female genitalia. Unknown.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044C696713ADFA0FFB41FD20.taxon	etymology	Etymology. The species is named after the fictional personage of the Star Trek movie, Sr. Spock (Leonard Simon Nimoy), Science officer and first officer of the Enterprise Starship.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044C696713ADFA0FFB41FD20.taxon	distribution	Geographic Distribution. This species occurs in Santiago and Coquimbo biogeographic provinces in Chile (Fig. 72).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044C696713ADFA0FFB41FD20.taxon	biology_ecology	Natural History. According to the specimen label the specimens have been collected in spring (September). Adults were observed visiting flowers of Heliotropium krauseanum (Boragineaceae) (Fig. 64).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044E696013ADFCABFE5FF80C.taxon	description	(Fig. 60)	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044E696013ADFCABFE5FF80C.taxon	materials_examined	Type material: After several searches in the museum collections, we have not been able to find the type specimens. Probably is lost. However, we studied voucher specimens deposited at (JEBTC) collection, identified by Ramírez and voucher specimens deposited at (LEC and MLPC). The characters of the specimen match those of the original description.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044E696013ADFCABFE5FF80C.taxon	discussion	Taxonomic comments. Solier (1851) described the species Tetraonyx flavipennis within the genus Tetraonyx, marking its name with a cross (†) to indicate its status as new to him. However, Guérin-Ménéville (1843) had already described a species under the same name, T. flavipennis, in the same genus. Consequently, the two names constitute a case of primary homonymy, rendering Tetraonyx flavipennis Solier, 1851 an unavailable name (Cazzañiga, pers. comm.). Seven years later, Lacordaire (1859) recognized that Solier’s and Guérin-Ménéville’s taxa were distinct species, explaining why Solier (1851) had placed his in a separate section, Picnoseus. Nevertheless, Tetraonyx flavipennis Solier, 1851 remains unavailable. Gemminger (1870) subsequently proposed the replacement name Tetraonyx xanthopterus, which must be used instead.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044E696013ADFCABFE5FF80C.taxon	materials_examined	Material examined. Chile Guanaqueros prov. Elqui, 18 / IX / 1990, leg. J. E. Barriga, Col. J. E. Barriga Chile 31613 1 ♂ JEBTC. Chile, Guanaqueros, Coll. J. E. Barriga, 18 / XI / 1990, 1 ♀ LEC. Chile, Malleco: Lonquimay, Coll. J. E. Barriga 22 / XII / 1994, 1 ♀ LEC. Chile, Malleco: Malalcahuello, Coll J. E. Barriga, 2 / I / 2007, 3 ♀ JECTB. Other material examined. 6 exx JEBTC, Chile, Los Andes, Coll. Ruiz F. 1918, MLPC. Chile, Malleco: Lonquimay, Coll. J. E. Barriga, 22 / XII / 1994. 1 ex IADIZA, Chile: Paipote Almagro salida a camino internacional, 21 / V / 1991 Col. S. Roig-Juñent.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044E696013ADFCABFE5FF80C.taxon	diagnosis	Diagnosis. Head and pronotum black, lustrous tegument; elytra yellowish- ochre. Head with big, dense, shallow punctures, but isolated from each other; pronotum with punctures along the edge and median groove; integument between them micro- reticulate; with two suborbicular, rugose areas located centrally on either side of the median groove; or with big-sized punctures homogeneously distributed. Setation piliform, grayish. Antennomere I broadest at midlength, with an anteroapical curvature that extends over segment IV, forming an apical lip; antennomere II cup-shaped; antennomere III broadest distally; antennomeres IV – X subtrapezoidal. Profemur and protibiae with the same length; mesafemur shorter than mesotibiae; metafemur and metatibiae with the same length. Metatibiae spurs similar in length, outer spur truncated distally, inner broader proximally (like spatula) rotated inwards. Hindwing veins compositions are summarized in table 2 b. Male genitalia with spiculum gastrale bearing external spines continuated with two lobes separated by a broad middle emargination; proximally slightly curved to the left; aedeagus with two hooks (Fig. 46).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044E696013ADFCABFE5FF80C.taxon	description	Redescription. Medium- sized species; length 12 – 15.5 mm. Coloration. Head and pronotum black, lustrous tegument; antennomeres I – IV and maxillary palpomere brown, antennomeres V – XI dark- brown. Legs dark- brown. Elytra yellowish-ochre. Scutellum black, lustrous tegument. Tegument. Head with big, dense, shallow punctures, but isolated from each other; integument among them micro-reticulate; frons with small punctures (smaller than vertex), integument among them rugose. Clypeus with punctures similar to those on frons except on the inferior half that is membranous. Labrum with puncture larger and dense than on clypeus. Pronotum with punctures along the edges and on the median groove; integument between them micro- reticulate; with two suborbicular, rugose areas located centrally on either side of the median groove; or with big- sized punctures homogeneously distributed; integument between them lustrous. Elytra slightly rugose, with three slightly marked costae on disk; sutural costae longitudinal extended from base to the apex. Setation. Piliform, grayish. Clypeus and labrum with setae similar to those on head (but somewhat shorter). Male and female protibiae with ventral surface (that opposing femur) with an extensive patch of subappressed, transversely directed pubescence; male protarsus with ventral pads homogeneously distributed; female protarsus with ventral pads confined to the distal portion, proximal surface covered with thicker and sparse setae. Head. Suborbicular (length 1.5 mm; width 1.8 mm; maximum width just above eyes); surface with longitudinal subtle median groove from vertex to the upper area of fronto-clypeal suture. Frons transversely depressed before the suture, or with marked suborbicular depression at middle of the frons (between eyes). Temple subparallel, rounded posteriorly. Eyes flat; anteriorly broad emarginate; inferior lobe narrowed apically (wide 0.6 mm; long 0.9 mm). Clypeus bulged, transversal, with subparallel sides (wide 0.5 mm; long 0.9 mm). Labrum with rounded lateral edges; ventrally slightly concave (wide 0.3 mm; long 1.1 mm). Maxillary palpomere IV subrectangular (sides subparallel). Labial palpomere III subrectangular but narrow distally. Mandible short, widely curved on the external side, mostly covered by clypeus and labrum while resting or elongated partially covered by clypeus while resting. Antennae loosely articulated (Fig. 22). Antennomere I broadest at midlength, with an anteroapical curvature that extends over segment II, forming an apical lip; antennomere II cup-shaped; antennomere III broadest distally; antennomeres IV – X subtrapezoidal. Antennomere length: I 0.37 mm; II 0.12 mm; III 0.5 mm; IV – X 0.45 mm; XI 0.62 mm. Pronotum. Subhexagonal (length 1.8 mm; width 1.9 mm) maximum width at apical third, with angulous borders; surface with a slightly marked median groove. Legs. Profemur and protibiae with the same length (2.33 mm); mesafemur shorter than mesotibiae (2.5 mm; 2.9 mm respectively); metafemur and metatibiae with the same length (3.9 mm). Metatibiae spurs similar in length, outer spur truncated distally, inner broader proximally (like spatula) rotated inwards. Tarsomere measurement: forestarsomere (I 0.7 mm; II – IV 0.4 mm; V 0.7 mm); mesotarsomere (I 0.7 mm; II and III 0.5 mm; IV 0.3, V 0.7 mm); metatarsomere (I 1.3 mm; II 0.8 mm; III 0.6 mm, IV 0.9). Elytra. Total length 7 mm (greatest width at middle: 3 mm); with progressively divergent edges from the base to the apex; suture dehiscent at distal two-third. Hindwing veins are summarized in table 2 b (Fig. 35). Male genitalia (Fig. 46). SpGs bearing external spines continuated with two lobes separated by a broad middle emargination; proximally slightly curved to the left. Pa incompletely sclerotized, with a membranous area except for a Y- shaped sclerotized region. Pa and Ph fused by a marginal sclerotized band. Ae distally with two hooks: a robust DAH, with marked peak, and a very robust VAH. AeA short and straight. Female genitalia. Va subrectangular, each with a short proximal BsVa. Distal portion of each Va nearly cylindrical, bearing a separate, suboval Sty; Sty about half as long as Va.	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044E696013ADFCABFE5FF80C.taxon	distribution	Geographic Distribution. This species occurs in the northern Maule and south central coast of Coquimbo biogeographical provinces (Fig. 73).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
03CE87F4044E696013ADFCABFE5FF80C.taxon	biology_ecology	Natural History. According to specimen labels, P. xanthopterus were recorded in spring (September, October and November). Adults of this species were collected on flowers of Malvaceae: Cristaria glaucophylla Cav. (Malvaceae) (Denier 1918), an endemic plant of central and northern Chile. Also was observed feeding in flowers of Solanum trinominum (Solanaceae) (Figs. 65 and 66) (Available from https: // ecuador. inaturalist. org / observations / 139105617; date access April 2025).	en	Campos-Soldini, Maria P., Safenraiter, Melania E., Roig-Juñent, Sergio A. (2026): Taxonomic revision of the southamerican genus Picnoseus Solier (Coleoptera: Meloidae), including Pyrotini biogeographical considerations. Zootaxa 5777 (1): 1-45, DOI: 10.11646/zootaxa.5777.1.1, URL: https://doi.org/10.11646/zootaxa.5777.1.1
