taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426851/files/figure.png	https://doi.org/10.5281/zenodo.15426851	Fig. 2. Habitat of Andreaeobryum macrosporum in Yakutia. A: photo from 970 m elev. B–D, H: at 1070 m, E–F: at 1100 m.	Fig. 2. Habitat of Andreaeobryum macrosporum in Yakutia. A: photo from 970 m elev. B–D, H: at 1070 m, E–F: at 1100 m.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426853/files/figure.png	https://doi.org/10.5281/zenodo.15426853	Fig. 3. Habit of Andreaeobryum macrosporum (photographs taken from living plants, transported to the camp on original rock pieces). A: Plants with mostly open and partly broken capsules, arrowed is the capsule with spores seen among valves.B: Capsules within bicolored calyptrae.C: Tuft with capsules at various stages of maturation, including some still embedded in epigonium; note a bright base of the latter (arrowed, cf. Fig. 16A). D: Tuft with some shoots with julaceous foliage in basal part; note round premature capsule (arrowed). E: Tuft with male plant in the central part of photograph, with perigonia conspicuous due to lighter and brighter green color.	Fig. 3. Habit of Andreaeobryum macrosporum (photographs taken from living plants, transported to the camp on original rock pieces). A: Plants with mostly open and partly broken capsules, arrowed is the capsule with spores seen among valves.B: Capsules within bicolored calyptrae.C: Tuft with capsules at various stages of maturation, including some still embedded in epigonium; note a bright base of the latter (arrowed, cf. Fig. 16A). D: Tuft with some shoots with julaceous foliage in basal part; note round premature capsule (arrowed). E: Tuft with male plant in the central part of photograph, with perigonia conspicuous due to lighter and brighter green color.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437365/files/figure.png	https://doi.org/10.5281/zenodo.15437365	Fig. 4. Andreaeobryum macrosporum Steere & B.M. Murray (from: Russia, Yakutia, Segenyakh Creek, Ignatov & Ignatova 15-599, MHA). A–B – plants with sporophytes, dry; C – opened capsule, dry; D – female plant; E–F – capsules with calyptra; G – closed capsule; H – male plant; I – paraphysis; J – antheridium; K–M – perigonial leaves; N–O – beaked axillary hairs; P – archegonium; Q–R – perichaetial leaves; S–U – primary leaves. Scale bars: 2 mm for A–B, D, H; 1 mm for C, E–G; 0.5 mm for K– M, Q–U; 500 µm for I–J; P; 100 µm for N–O.	Fig. 4. Andreaeobryum macrosporum Steere & B.M. Murray (from: Russia, Yakutia, Segenyakh Creek, Ignatov & Ignatova 15-599, MHA). A–B – plants with sporophytes, dry; C – opened capsule, dry; D – female plant; E–F – capsules with calyptra; G – closed capsule; H – male plant; I – paraphysis; J – antheridium; K–M – perigonial leaves; N–O – beaked axillary hairs; P – archegonium; Q–R – perichaetial leaves; S–U – primary leaves. Scale bars: 2 mm for A–B, D, H; 1 mm for C, E–G; 0.5 mm for K– M, Q–U; 500 µm for I–J; P; 100 µm for N–O.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426855/files/figure.png	https://doi.org/10.5281/zenodo.15426855	Fig. 5. Andreaeobryum macrosporum (from: Russia, Yakutia, Segenyakh Creek, Ignatov & Ignatova 15-599, MHA). O–P: SEM (herbarium specimens, coated without drying in critical point). A – distal leaf cells; B – transverse section at distal part of leaf; C – distal cells of leaf from julaceous shoot; D – transverse section at mid-leaf; E – transverse section at the basal part of leaf; F – median leaf cells; G – basal leaf cells; H–K – stem leaves; L–N – leaves from julaceous shoot; O – dorsal leaf surface; P – ventral leaf surface. Scale bars: 1 mm for H–N; 100 µm for A–G.	Fig. 5. Andreaeobryum macrosporum (from: Russia, Yakutia, Segenyakh Creek, Ignatov & Ignatova 15-599, MHA). O–P: SEM (herbarium specimens, coated without drying in critical point). A – distal leaf cells; B – transverse section at distal part of leaf; C – distal cells of leaf from julaceous shoot; D – transverse section at mid-leaf; E – transverse section at the basal part of leaf; F – median leaf cells; G – basal leaf cells; H–K – stem leaves; L–N – leaves from julaceous shoot; O – dorsal leaf surface; P – ventral leaf surface. Scale bars: 1 mm for H–N; 100 µm for A–G.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426857/files/figure.png	https://doi.org/10.5281/zenodo.15426857	Fig.6. Andreaeobryum macrosporum (A, C, D: LM; B, Da, E–H: SM; I: LSCM, living plants). A: Protonema forming a mass upon rock surface in ‘B’. B: Protonema with protonematal leaves upon rock surface. C: Rhizoids from young innovation ‘Ce’, showing variation in their structure (see text for discussion). D: Rhizoids arising from the wounded shoot (cf. Da); note strongly oblique wall between cells. E–G: male plant, perigonia arrowed; note dimorphism of innovations in ‘G’. H: Shoot with leaveas covered by calcareous material, apparently cemented by moldy bacterial layer upon leaves. I: Perigonium with two young innovations.	Fig.6. Andreaeobryum macrosporum (A, C, D: LM; B, Da, E–H: SM; I: LSCM, living plants). A: Protonema forming a mass upon rock surface in ‘B’. B: Protonema with protonematal leaves upon rock surface. C: Rhizoids from young innovation ‘Ce’, showing variation in their structure (see text for discussion). D: Rhizoids arising from the wounded shoot (cf. Da); note strongly oblique wall between cells. E–G: male plant, perigonia arrowed; note dimorphism of innovations in ‘G’. H: Shoot with leaveas covered by calcareous material, apparently cemented by moldy bacterial layer upon leaves. I: Perigonium with two young innovations.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437397/files/figure.png	https://doi.org/10.5281/zenodo.15437397	Fig. 7. (previous page) Andreaeobryum macrosporum (living, A–L), Takakia lepidozioides (living, M–N) and Sphagnum girgensohnii (living, O–R) axillary hairs with apical pore (AP), position and structure (A–G, J, L, O–Q: LSCM; H–I, K, M, R: LM; N: SM). A: Leaves crowded at shoot apex; note beaked axillary hairs (ah) and putative filamentose primary leaf (PL?). B–I: axillary hairs. C: still not opened axillary hair. E: axillary hair with protoplast fallen out, so looking as branched in LM. J and L: apical parts of axillary hairs. K: a bud, propagated in perigonium of old plant, with few young orbicular leaves and axillary hairs, studied in living state and presented in Figs. C, E, J and L. Note that B,F, J and L illustrate an armature of apical cell, apparently regulating mucilage release. Takakia pictures show mucilage release through the apical pore (AP). Sphagnum ‘P’ and ‘Q’ are close ups of ‘O’.	Fig. 7. (previous page) Andreaeobryum macrosporum (living, A–L), Takakia lepidozioides (living, M–N) and Sphagnum girgensohnii (living, O–R) axillary hairs with apical pore (AP), position and structure (A–G, J, L, O–Q: LSCM; H–I, K, M, R: LM; N: SM). A: Leaves crowded at shoot apex; note beaked axillary hairs (ah) and putative filamentose primary leaf (PL?). B–I: axillary hairs. C: still not opened axillary hair. E: axillary hair with protoplast fallen out, so looking as branched in LM. J and L: apical parts of axillary hairs. K: a bud, propagated in perigonium of old plant, with few young orbicular leaves and axillary hairs, studied in living state and presented in Figs. C, E, J and L. Note that B,F, J and L illustrate an armature of apical cell, apparently regulating mucilage release. Takakia pictures show mucilage release through the apical pore (AP). Sphagnum ‘P’ and ‘Q’ are close ups of ‘O’.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437395/files/figure.png	https://doi.org/10.5281/zenodo.15437395	Fig. 8. Andreaeobryum macrosporum (TEM). Axillary hairs from the area near the end of foot (sections transverse to foot and also representing transverse sections of the axillary hairs in their middle part (‘C–D’), while for some axillary hairs bent to one side the apical part has also been available for study (‘A–B’). A: the apical part of the axillary hair, showing a place of mucilage release (looking like a pore, it is rather a transverse section of a rupture, shown in Figs. 7J and 7L); drops of hygrophobous compounds occur near the ‘pore’; among the complex structure below the pore, there is one most massive conic ‘reservoir’ (marked with asterisk*), 0.8 µm wide and 0.6 µm long, at the distance ca. 1 µm below the pore; its proximity is especially rich in fibrillose network, apparently participated in the regulation of the mucilage discharge. B: close up of A, for surrounding of pore and ‘reservoir’, surrounded by bundles of these fibrils. C–D: part of transverse section of the axillary hair, showing extensive endoplasmatic reticulum (ER) and dictyosome (Di), as well as a compleх trilayered wall of the cells formed axillary hair.	Fig. 8. Andreaeobryum macrosporum (TEM). Axillary hairs from the area near the end of foot (sections transverse to foot and also representing transverse sections of the axillary hairs in their middle part (‘C–D’), while for some axillary hairs bent to one side the apical part has also been available for study (‘A–B’). A: the apical part of the axillary hair, showing a place of mucilage release (looking like a pore, it is rather a transverse section of a rupture, shown in Figs. 7J and 7L); drops of hygrophobous compounds occur near the ‘pore’; among the complex structure below the pore, there is one most massive conic ‘reservoir’ (marked with asterisk*), 0.8 µm wide and 0.6 µm long, at the distance ca. 1 µm below the pore; its proximity is especially rich in fibrillose network, apparently participated in the regulation of the mucilage discharge. B: close up of A, for surrounding of pore and ‘reservoir’, surrounded by bundles of these fibrils. C–D: part of transverse section of the axillary hair, showing extensive endoplasmatic reticulum (ER) and dictyosome (Di), as well as a compleх trilayered wall of the cells formed axillary hair.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426859/files/figure.png	https://doi.org/10.5281/zenodo.15426859	Fig. 9. Andreaeobryum macrosporum, female shoot (LSCM): longitudinal (A) and transverse sections, where values indicate the distance in µm from the apical cell (denoted as 0 in longitudinal section A). Note that unistratose lamina in younger leaves quickly transforms to multistratose. Archegonium (AR) is developed in a lateral position to apical cell (*); numbers on photo indicate a number of leaf conventionally (regarding available leaves at the level of stem apical cell). Axillary hairs (ah) are irregular in shape; it seems that slimy blur at apical cell level is caused by their contents (cf. Fig. 7).	Fig. 9. Andreaeobryum macrosporum, female shoot (LSCM): longitudinal (A) and transverse sections, where values indicate the distance in µm from the apical cell (denoted as 0 in longitudinal section A). Note that unistratose lamina in younger leaves quickly transforms to multistratose. Archegonium (AR) is developed in a lateral position to apical cell (*); numbers on photo indicate a number of leaf conventionally (regarding available leaves at the level of stem apical cell). Axillary hairs (ah) are irregular in shape; it seems that slimy blur at apical cell level is caused by their contents (cf. Fig. 7).	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437399/files/figure.png	https://doi.org/10.5281/zenodo.15437399	Fig. 10 (previous page). Andreaeobryum macrosporum (LSCM), four series of longitudinal sections through stem apical cells. Series a, b and c were done in female shoots (with values indicating distance in µm from the the sections denoted as 0); series d is from julaceous shoot with small orbicular leaves. Series ‘a’ shows young archegonium (AR) next to the apical cell (*), while the latter remains functional. Series ‘b’ and ‘c’ show primary leaves near the stem apical cell. It is not clear if some young leaf-like structures (e.g., ‘leaf’ in front of apical cell in ‘b0’ and ‘b8’) is not a young archegonium. Andreaeobryum macrosporum (LSCM), longitudinal sections of julaceous shoot with small orbicular leaves. Some divisions in apical cell (in ‘8’) are almost longitudinal (compare with Fig. 9–b24). Fig. 11 (next page). Andreaeobryum macrosporum, young leaves near sterile shoot apex (A,C-J: LSCM, B, K-L: LM [K: Shiff staining]). A–D: one group of leaves, showing bulgings in distal cells; note that leaves #2 and #6 are obviously narrower at base than above (cf. with Figs. I–J). E-H: another group of leaves from different views, showing that the apical leaf cell is rather trifacial, not bifacial, forming multistratose leaf lamina since the beginning; leaf #3 is multistratose as well; I-J: two leaves at opposite views; such rounded and concave leaves are the first in julaceous shoots. K: Cytoplasm staining illustrates the thick cell walls in distal parts of young leaves, although not from the earliest stage (compare with leaves #4 and #5 in the ‘A-D’ pictures). L: young leaves with rather apparent trifacial structure.	Fig. 10 (previous page). Andreaeobryum macrosporum (LSCM), four series of longitudinal sections through stem apical cells. Series a, b and c were done in female shoots (with values indicating distance in µm from the the sections denoted as 0); series d is from julaceous shoot with small orbicular leaves. Series ‘a’ shows young archegonium (AR) next to the apical cell (*), while the latter remains functional. Series ‘b’ and ‘c’ show primary leaves near the stem apical cell. It is not clear if some young leaf-like structures (e.g., ‘leaf’ in front of apical cell in ‘b0’ and ‘b8’) is not a young archegonium. Andreaeobryum macrosporum (LSCM), longitudinal sections of julaceous shoot with small orbicular leaves. Some divisions in apical cell (in ‘8’) are almost longitudinal (compare with Fig. 9–b24). Fig. 11 (next page). Andreaeobryum macrosporum, young leaves near sterile shoot apex (A,C-J: LSCM, B, K-L: LM [K: Shiff staining]). A–D: one group of leaves, showing bulgings in distal cells; note that leaves #2 and #6 are obviously narrower at base than above (cf. with Figs. I–J). E-H: another group of leaves from different views, showing that the apical leaf cell is rather trifacial, not bifacial, forming multistratose leaf lamina since the beginning; leaf #3 is multistratose as well; I-J: two leaves at opposite views; such rounded and concave leaves are the first in julaceous shoots. K: Cytoplasm staining illustrates the thick cell walls in distal parts of young leaves, although not from the earliest stage (compare with leaves #4 and #5 in the ‘A-D’ pictures). L: young leaves with rather apparent trifacial structure.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437401/files/figure.png	https://doi.org/10.5281/zenodo.15437401	Fig. 11 (next page). Andreaeobryum macrosporum, young leaves near sterile shoot apex (A,C-J: LSCM, B, K-L: LM [K: Shiff staining]). A–D: one group of leaves, showing bulgings in distal cells; note that leaves #2 and #6 are obviously narrower at base than above (cf. with Figs. I–J). E-H: another group of leaves from different views, showing that the apical leaf cell is rather trifacial, not bifacial, forming multistratose leaf lamina since the beginning; leaf #3 is multistratose as well; I-J: two leaves at opposite views; such rounded and concave leaves are the first in julaceous shoots. K: Cytoplasm staining illustrates the thick cell walls in distal parts of young leaves, although not from the earliest stage (compare with leaves #4 and #5 in the ‘A-D’ pictures). L: young leaves with rather apparent trifacial structure.	Fig. 11 (next page). Andreaeobryum macrosporum, young leaves near sterile shoot apex (A,C-J: LSCM, B, K-L: LM [K: Shiff staining]). A–D: one group of leaves, showing bulgings in distal cells; note that leaves #2 and #6 are obviously narrower at base than above (cf. with Figs. I–J). E-H: another group of leaves from different views, showing that the apical leaf cell is rather trifacial, not bifacial, forming multistratose leaf lamina since the beginning; leaf #3 is multistratose as well; I-J: two leaves at opposite views; such rounded and concave leaves are the first in julaceous shoots. K: Cytoplasm staining illustrates the thick cell walls in distal parts of young leaves, although not from the earliest stage (compare with leaves #4 and #5 in the ‘A-D’ pictures). L: young leaves with rather apparent trifacial structure.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426861/files/figure.png	https://doi.org/10.5281/zenodo.15426861	Fig. 12. Andreaeobryum macrosporum, antheridia and paraphyses (A: LSCM; B–K: LM [B–H: Schiff plus indigo carmine staining]). A: Premature unopened an- theridium. B–C: Antheridia, colors are changing along maturation, depending on pH. D: Stalk, close-up from ‘C’. E–G: Paraphyses, colors are changing along matu- ration, depending on pH; note that cells are relatively thick-walled, but walls in apical cells are usually evenly thickened, although sometimes a ‘beak-pattern’ is observed, cf. ‘E’. H: Open antheridium, thick-walled cells along distal pore are colored; I–J: Recently opened antheridium, with only small part of spermatozoids releasedand someare in queue waiting topass throughthe narrow pore with tickened mouth; K: shoot, photographed through leaves, showing outlines of antheridia: nu- merous and very young in the apical group and one of mature outline and with multicellular long stalk in the axil of leaf somewhat beside the apical group.	Fig. 12. Andreaeobryum macrosporum, antheridia and paraphyses (A: LSCM; B–K: LM [B–H: Schiff plus indigo carmine staining]). A: Premature unopened an- theridium. B–C: Antheridia, colors are changing along maturation, depending on pH. D: Stalk, close-up from ‘C’. E–G: Paraphyses, colors are changing along matu- ration, depending on pH; note that cells are relatively thick-walled, but walls in apical cells are usually evenly thickened, although sometimes a ‘beak-pattern’ is observed, cf. ‘E’. H: Open antheridium, thick-walled cells along distal pore are colored; I–J: Recently opened antheridium, with only small part of spermatozoids releasedand someare in queue waiting topass throughthe narrow pore with tickened mouth; K: shoot, photographed through leaves, showing outlines of antheridia: nu- merous and very young in the apical group and one of mature outline and with multicellular long stalk in the axil of leaf somewhat beside the apical group.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437405/files/figure.png	https://doi.org/10.5281/zenodo.15437405	Fig. 13 (previous page). Andreaeobryum macrosporum (A, D–H: LM; B–C: LSCM) and Andreaea rupestris (I–J: LM), archegonia, showing their variable apical and subapical positions in Andreaeobryum and location at base of pseudopodium in Andreaea. Note pedestaled archegonia both in Andreaeobryum and in Andreaea. Note primary leaves (PL) in between large leaves and beside archegonia, and numerous axillary hairs (ah). The leaf apical cell in ‘Aa’ has a terminal papillae.	Fig. 13 (previous page). Andreaeobryum macrosporum (A, D–H: LM; B–C: LSCM) and Andreaea rupestris (I–J: LM), archegonia, showing their variable apical and subapical positions in Andreaeobryum and location at base of pseudopodium in Andreaea. Note pedestaled archegonia both in Andreaeobryum and in Andreaea. Note primary leaves (PL) in between large leaves and beside archegonia, and numerous axillary hairs (ah). The leaf apical cell in ‘Aa’ has a terminal papillae.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437407/files/figure.png	https://doi.org/10.5281/zenodo.15437407	Fig. 14 (next page). Andreaeobryum macrosporum (A–D: LM, the rest: LSCM), showing position of archegonia side by side with primary leaves. ‘D’ is pedestaled leaf, or a ‘half-leaf, half archegonium’, likely started developing as archegonium, but after fertilization of neigboring archegonium started developing lamina. The series ‘10’–‘160’ shows one leaf axil with one archegonium (AR) and three primary leaves PL1–3, shown in selected pictures). Note that the pedestal of archegonium has leaf-like structure: see ‘126’. Note that the central part of stem has enormously extensive intracellular space, putatively having conduction capacity.	Fig. 14 (next page). Andreaeobryum macrosporum (A–D: LM, the rest: LSCM), showing position of archegonia side by side with primary leaves. ‘D’ is pedestaled leaf, or a ‘half-leaf, half archegonium’, likely started developing as archegonium, but after fertilization of neigboring archegonium started developing lamina. The series ‘10’–‘160’ shows one leaf axil with one archegonium (AR) and three primary leaves PL1–3, shown in selected pictures). Note that the pedestal of archegonium has leaf-like structure: see ‘126’. Note that the central part of stem has enormously extensive intracellular space, putatively having conduction capacity.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426863/files/figure.png	https://doi.org/10.5281/zenodo.15426863	Fig. 15. Andreaeobryum macrosporum, transverse sections of young sporophyte of ca. 200 µm long, still embedded in epigonium (–10 and –20 and their close ups: TEM; 10–144: LSCM; the values indicate the distance from foot tip, µm). Note strongly modified gametophyte cells near sporophyte foot, being rich in starch (white), but without ingrowths. Note that at the level of capsule (126–144) sporophyte cells retain the arrangement of fundamental cross, whereas at the level of foot (68–78) sporophyte outline and cells are rather flattened (cf. with Fig. 23).	Fig. 15. Andreaeobryum macrosporum, transverse sections of young sporophyte of ca. 200 µm long, still embedded in epigonium (–10 and –20 and their close ups: TEM; 10–144: LSCM; the values indicate the distance from foot tip, µm). Note strongly modified gametophyte cells near sporophyte foot, being rich in starch (white), but without ingrowths. Note that at the level of capsule (126–144) sporophyte cells retain the arrangement of fundamental cross, whereas at the level of foot (68–78) sporophyte outline and cells are rather flattened (cf. with Fig. 23).	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426865/files/figure.png	https://doi.org/10.5281/zenodo.15426865	Fig. 16. Andreaeobryum macrosporum (LSCM), longitudinal sections through epigonium (total view in ‘A’) with young sporophyte of ca. 400 µm long inside (the latter is somewhat oblique). Values indicate the distance (in µm) from the section denoted as 0, where placental space with mucilage medium appeared in this series of sections. Sporophyte cells appear after 6 µm (arrowed in ‘6’ and ‘8’). Gametophytic cells adjoining to the foot are narrow. Foot cells have brighter fluorescence due to cell wall ingrowths (cf. Fig. 17 and 18A–D). Note a three-celled beaked axillary hair in ‘0’–‘2’–‘4’, followed by five-celled axillary hair in ‘6’–‘8’–‘12’.	Fig. 16. Andreaeobryum macrosporum (LSCM), longitudinal sections through epigonium (total view in ‘A’) with young sporophyte of ca. 400 µm long inside (the latter is somewhat oblique). Values indicate the distance (in µm) from the section denoted as 0, where placental space with mucilage medium appeared in this series of sections. Sporophyte cells appear after 6 µm (arrowed in ‘6’ and ‘8’). Gametophytic cells adjoining to the foot are narrow. Foot cells have brighter fluorescence due to cell wall ingrowths (cf. Fig. 17 and 18A–D). Note a three-celled beaked axillary hair in ‘0’–‘2’–‘4’, followed by five-celled axillary hair in ‘6’–‘8’–‘12’.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426867/files/figure.png	https://doi.org/10.5281/zenodo.15426867	Fig. 17. Andreaeobryum macrosporum (TEM), longitudinal section through foot of young sporophyte embedded in epigonium and shown in Fig. 16-70’ (note that this section is somewhat beside from the tip of the foot). Note numerous ingrowths of cell walls in the outer cell layer of foot and occasionally in some inner cells in ‘B’, and labyrinths on the foot surface faced to the placental space (cf. also Figs. 18A–D). A: Foot is separated from gametophyte tissue by expanded placental spaced filled by macerated and partly decomposed gametophytic cells; note that walls of gametophytic cells faced to placental space lack any ingrowth, but they are partly irregular in shape, obviously experienced some degradation toward complete maceration. B–D: Foot periphery showing protruding of its surface cells. E–F: foot surface showing some ‘fjords’ of placental space, filled with possible starch-like material.	Fig. 17. Andreaeobryum macrosporum (TEM), longitudinal section through foot of young sporophyte embedded in epigonium and shown in Fig. 16-70’ (note that this section is somewhat beside from the tip of the foot). Note numerous ingrowths of cell walls in the outer cell layer of foot and occasionally in some inner cells in ‘B’, and labyrinths on the foot surface faced to the placental space (cf. also Figs. 18A–D). A: Foot is separated from gametophyte tissue by expanded placental spaced filled by macerated and partly decomposed gametophytic cells; note that walls of gametophytic cells faced to placental space lack any ingrowth, but they are partly irregular in shape, obviously experienced some degradation toward complete maceration. B–D: Foot periphery showing protruding of its surface cells. E–F: foot surface showing some ‘fjords’ of placental space, filled with possible starch-like material.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426869/files/figure.png	https://doi.org/10.5281/zenodo.15426869	Fig. 18. Andreaeobryum macrosporum (TEM). A–D: Longitudinal section through the foot of young (cf. Fig. 16–70) sporophyte embedded in epigonium. E–F: Cells of vaginula (V) and placental space (PC) adjoining to the foot (F) of mature sporophyte (cf. Fig. 23), showing cells of placental space with strongly ‘softened cell walls’, with numerous invaginations. G–H: Ultrastructure of surface foot cells (from specimens in Fig. 19): among chloroplasts (Ch) and endoplasmatic reticulum (ER), the ER whorls occur, presumably implementing autofagous function, utilizing membranes along with the labyrinth (L) expansion.	Fig. 18. Andreaeobryum macrosporum (TEM). A–D: Longitudinal section through the foot of young (cf. Fig. 16–70) sporophyte embedded in epigonium. E–F: Cells of vaginula (V) and placental space (PC) adjoining to the foot (F) of mature sporophyte (cf. Fig. 23), showing cells of placental space with strongly ‘softened cell walls’, with numerous invaginations. G–H: Ultrastructure of surface foot cells (from specimens in Fig. 19): among chloroplasts (Ch) and endoplasmatic reticulum (ER), the ER whorls occur, presumably implementing autofagous function, utilizing membranes along with the labyrinth (L) expansion.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426871/files/figure.png	https://doi.org/10.5281/zenodo.15426871	Fig. 19. Andreaeobryum macrosporum (A: LM, B, D, F: TEM; C, E: LSCM). Longitudinal section of foot of sporophyte at the stage approximately the same as in Fig. 16. A: Change of color of still intact gametophyte cells around the foot and decomposed cells in placental space. B, D, F: details of ‘A’, showing lack of ingrowths in gametophytic cells, strongly developed labyrinths in outermost foot cells and strongly modified cells within placental space. ‘D’ shows ‘fjords’ within the foot, filled with the putatively starch ‘outer labyrinths’, as in Figs. 17E–F. C & E are LSCM picutes from 2 µm section, shown in ‘A’: labyrinths looks spongy, being especially well-developed near haustorium, but quite abundant in the second surface layer of cells in the middle part of the foot; no ingrowth are seen in the closest gametophytic cells, and decomposed cell material in placental cells produces a strong fluorescence.	Fig. 19. Andreaeobryum macrosporum (A: LM, B, D, F: TEM; C, E: LSCM). Longitudinal section of foot of sporophyte at the stage approximately the same as in Fig. 16. A: Change of color of still intact gametophyte cells around the foot and decomposed cells in placental space. B, D, F: details of ‘A’, showing lack of ingrowths in gametophytic cells, strongly developed labyrinths in outermost foot cells and strongly modified cells within placental space. ‘D’ shows ‘fjords’ within the foot, filled with the putatively starch ‘outer labyrinths’, as in Figs. 17E–F. C & E are LSCM picutes from 2 µm section, shown in ‘A’: labyrinths looks spongy, being especially well-developed near haustorium, but quite abundant in the second surface layer of cells in the middle part of the foot; no ingrowth are seen in the closest gametophytic cells, and decomposed cell material in placental cells produces a strong fluorescence.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437409/files/figure.png	https://doi.org/10.5281/zenodo.15437409	Fig. 20. Andreaeobryum macrosporum (LSCM). a20–a68: longitudinal sections through epigonium with young sporophyte. Series a20–a68 shows the stage somewhat 30 µm later than in Fig. 16 (sporophyte is 135 µm B in diameter vs. 100 µm in Fig. 16), while A and B show still later stage (sporophyte ca. 200 µm). Values indicate the distance (in µm) from the first section denoted as 0 (not shown), where placental space with mucilage medium around sporophyte appears discernible. Foot is composed of a larger number of cells and conducting tissue at the seta level is better developed. Haustorial part is more blunt and looks not so efficient in penetration in gametophytic tissue, as compared with that in Fig. 16. A solid mass of cells with small dark bodies near the base of archegonium neck likely represent spermatozoids, which were late for fertilization (red arrows in ‘56’ and ‘68’). ‘20’ shows pre-exothecial cells in the upper third of sporophyte. ‘56’ is a section through the middle of the sporophyte, showing well differentiated amphithecium (am) and endothecium (en) in the upper part of sporopyte. A bar on the right indicates the level, represented in a series of transverse sections in Fig. 26. Potential archesporial tissue (‘ar?’ in ‘68’) is still almost undifferentiated. A-B: transverse sections, showing lobate foot, with outer cells with extensive ingrowths. The section ‘B’ is closer to foot base, so leaf, archegonia and axillary hairs sections at various levels are seen.	Fig. 20. Andreaeobryum macrosporum (LSCM). a20–a68: longitudinal sections through epigonium with young sporophyte. Series a20–a68 shows the stage somewhat 30 µm later than in Fig. 16 (sporophyte is 135 µm B in diameter vs. 100 µm in Fig. 16), while A and B show still later stage (sporophyte ca. 200 µm). Values indicate the distance (in µm) from the first section denoted as 0 (not shown), where placental space with mucilage medium around sporophyte appears discernible. Foot is composed of a larger number of cells and conducting tissue at the seta level is better developed. Haustorial part is more blunt and looks not so efficient in penetration in gametophytic tissue, as compared with that in Fig. 16. A solid mass of cells with small dark bodies near the base of archegonium neck likely represent spermatozoids, which were late for fertilization (red arrows in ‘56’ and ‘68’). ‘20’ shows pre-exothecial cells in the upper third of sporophyte. ‘56’ is a section through the middle of the sporophyte, showing well differentiated amphithecium (am) and endothecium (en) in the upper part of sporopyte. A bar on the right indicates the level, represented in a series of transverse sections in Fig. 26. Potential archesporial tissue (‘ar?’ in ‘68’) is still almost undifferentiated. A-B: transverse sections, showing lobate foot, with outer cells with extensive ingrowths. The section ‘B’ is closer to foot base, so leaf, archegonia and axillary hairs sections at various levels are seen.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426873/files/figure.png	https://doi.org/10.5281/zenodo.15426873	Fig.21. Andreaeobryum macrosporum (A–G: TEM; H–J: LSCM; K: LM; specimen from Fig. 16). Longitudinal sections through epigonium with young sporophyte inside, showing epigonium wall and some its inner decomposing cells filling the placental space. A–B: Walls of outer epigonium cells, showing complex multistratose polysaccharide structure. C–D: Space between sporophyte and inner surface of epigonium wall, showing long cells (arrowed) floating in a medium, likely a sort of mucilage. E: Epigonium inner wall faced sporophyte apex: this is a place with especially strongly decomposed inner cells of epigonium wall. F–G: Cells in placental space near the sporophyte apex (‘H’) and closer to foot (‘I-K’): strongly incrassate cell wall fills almost the whole volume of the cell. H-K: Autofluorescence and Schiff staining indicate a high content of polysaccharide in the placental space.	Fig.21. Andreaeobryum macrosporum (A–G: TEM; H–J: LSCM; K: LM; specimen from Fig. 16). Longitudinal sections through epigonium with young sporophyte inside, showing epigonium wall and some its inner decomposing cells filling the placental space. A–B: Walls of outer epigonium cells, showing complex multistratose polysaccharide structure. C–D: Space between sporophyte and inner surface of epigonium wall, showing long cells (arrowed) floating in a medium, likely a sort of mucilage. E: Epigonium inner wall faced sporophyte apex: this is a place with especially strongly decomposed inner cells of epigonium wall. F–G: Cells in placental space near the sporophyte apex (‘H’) and closer to foot (‘I-K’): strongly incrassate cell wall fills almost the whole volume of the cell. H-K: Autofluorescence and Schiff staining indicate a high content of polysaccharide in the placental space.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426875/files/figure.png	https://doi.org/10.5281/zenodo.15426875	Fig. 22. Andreaea rupestris (A–B, F: LSCM; C: LM, no staining) and Andreaeobryum macrosporum (D–E: LM, including D, same specimen as in Fig. 19, and E, living). Longitudinal sections or views at the stage when the sporophyte is still in epigonium. A & C show a hand sections of living plants. C: view of entire epigonium. F: pan- orama of three sections. In ‘F’ two lowermost haustorial cells have a somewhat brighter fluorescence, indicating more extensive ingrowth, comparatively with other cells, both of gametophyte and sporophyte. Cells of the pseudopodium of Anrdeaea (‘F’) and of gametophyte near the foot in Andreaeobryum (‘D’) are quite different in shape, the former being much more elongate. The part of stem in the section (‘D’) is composed of different cells due to stem curvation shortly below the epigonium. E: chlorophyllose cells of the foot (taken from epigonium at the stage comparable with ‘B’ and ‘C’ in this plate).	Fig. 22. Andreaea rupestris (A–B, F: LSCM; C: LM, no staining) and Andreaeobryum macrosporum (D–E: LM, including D, same specimen as in Fig. 19, and E, living). Longitudinal sections or views at the stage when the sporophyte is still in epigonium. A & C show a hand sections of living plants. C: view of entire epigonium. F: pan- orama of three sections. In ‘F’ two lowermost haustorial cells have a somewhat brighter fluorescence, indicating more extensive ingrowth, comparatively with other cells, both of gametophyte and sporophyte. Cells of the pseudopodium of Anrdeaea (‘F’) and of gametophyte near the foot in Andreaeobryum (‘D’) are quite different in shape, the former being much more elongate. The part of stem in the section (‘D’) is composed of different cells due to stem curvation shortly below the epigonium. E: chlorophyllose cells of the foot (taken from epigonium at the stage comparable with ‘B’ and ‘C’ in this plate).	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426877/files/figure.png	https://doi.org/10.5281/zenodo.15426877	Fig. 23. Andreaeobryum macrosporum (LM). Transverse sections through almost mature, but still not opened capsule and its seta and foot, corresponding to stage shown in longitudinal section in Figs. 30A. Values indicate the distance (in µm) from the end of foot. Foot is composed of dark cells, and cells of placental space are collapsed and destroyed (they occur up to ‘690’ section, the last one where gametophyte tissue is shown). In the middle part of foot (shown in section ‘258’) outer cells of the foot are radially elongate, considerably differing from the heterogeneous cells of the central part of foot. From ‘462’ a central strand appears in the seta: it is composed by larger cells with thicker walls, contrastingly different from surrounding cells, which are filled with some material (apparently starch-like), while cells of this central strand look empty. At ‘540’ the central strand is broadened up to half of foot diameter and then it fills almost the whole width of the foot, leaving only 2–3 layers of the starch-like tissue surrounding the foot. The vaginula extends to ‘690’, although alreary at ‘540’ gametophytic tissue (‘G’, arrowed) surrounding the foot/ seta (‘S’, arrowed) is thin, eroded and hardy functional. At the level of middle part of seta, from ‘848’ to ‘1342’, the seta becomes somewhat flattened in the ratio of its own width to height, as well as cells in its central part become transversely elongate. Closer to capsule base, at ‘1352’, cells in the centre are changing to larger; in ‘1430’ rectangular cells surrounding the base of columella are corresponding to the amphithecium (cf. with ‘1450’–‘1506’ in Fig. 24).	Fig. 23. Andreaeobryum macrosporum (LM). Transverse sections through almost mature, but still not opened capsule and its seta and foot, corresponding to stage shown in longitudinal section in Figs. 30A. Values indicate the distance (in µm) from the end of foot. Foot is composed of dark cells, and cells of placental space are collapsed and destroyed (they occur up to ‘690’ section, the last one where gametophyte tissue is shown). In the middle part of foot (shown in section ‘258’) outer cells of the foot are radially elongate, considerably differing from the heterogeneous cells of the central part of foot. From ‘462’ a central strand appears in the seta: it is composed by larger cells with thicker walls, contrastingly different from surrounding cells, which are filled with some material (apparently starch-like), while cells of this central strand look empty. At ‘540’ the central strand is broadened up to half of foot diameter and then it fills almost the whole width of the foot, leaving only 2–3 layers of the starch-like tissue surrounding the foot. The vaginula extends to ‘690’, although alreary at ‘540’ gametophytic tissue (‘G’, arrowed) surrounding the foot/ seta (‘S’, arrowed) is thin, eroded and hardy functional. At the level of middle part of seta, from ‘848’ to ‘1342’, the seta becomes somewhat flattened in the ratio of its own width to height, as well as cells in its central part become transversely elongate. Closer to capsule base, at ‘1352’, cells in the centre are changing to larger; in ‘1430’ rectangular cells surrounding the base of columella are corresponding to the amphithecium (cf. with ‘1450’–‘1506’ in Fig. 24).	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426879/files/figure.png	https://doi.org/10.5281/zenodo.15426879	Fig. 24 (continued from series in Fig. 23). Transverse sections, where values indicate the distance (in µm) from the end of foot. Spore chamber starts in this series from ‘1450’, at first as several (roughly four) compartments between the branches of cross-like columella (Co). Note a strong resorption of cells of the urn wall: in capsule middle, at ‘1674’ urn wall consists of three cell layers inside the exothecium, while in capsule bottom, at ‘1480’it has up to 13 layers (cf. also with Fig. 27).	Fig. 24 (continued from series in Fig. 23). Transverse sections, where values indicate the distance (in µm) from the end of foot. Spore chamber starts in this series from ‘1450’, at first as several (roughly four) compartments between the branches of cross-like columella (Co). Note a strong resorption of cells of the urn wall: in capsule middle, at ‘1674’ urn wall consists of three cell layers inside the exothecium, while in capsule bottom, at ‘1480’it has up to 13 layers (cf. also with Fig. 27).	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426881/files/figure.png	https://doi.org/10.5281/zenodo.15426881	Fig. 25. Andreaeobryum macrosporum (TEM, same specimen as in Fig. 16-70). Longitudinal sections at stage when the sporophyte is still in epigonium. A: Central part of the capsule, placental space (PS) and epigonium wall (EW), showing the complicated structure of the latter (cf. Fig. 21). B: Cells of the ‘pre-columella’ in upper part of sporophyte in ‘A’; nuclei and chloroplasts fill most of the cell volume; transverse cells walls are strongly flexuose and porose. C: Lower part of the capsule at the level of transtition to seta, cell walls are strongly flexuose. D: Cells from the lower part of the capsule, showing strongly flexuose cell walls; plasmodesmata are seen in both transverse and longitudinal cell walls.	Fig. 25. Andreaeobryum macrosporum (TEM, same specimen as in Fig. 16-70). Longitudinal sections at stage when the sporophyte is still in epigonium. A: Central part of the capsule, placental space (PS) and epigonium wall (EW), showing the complicated structure of the latter (cf. Fig. 21). B: Cells of the ‘pre-columella’ in upper part of sporophyte in ‘A’; nuclei and chloroplasts fill most of the cell volume; transverse cells walls are strongly flexuose and porose. C: Lower part of the capsule at the level of transtition to seta, cell walls are strongly flexuose. D: Cells from the lower part of the capsule, showing strongly flexuose cell walls; plasmodesmata are seen in both transverse and longitudinal cell walls.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426883/files/figure.png	https://doi.org/10.5281/zenodo.15426883	Fig. 26. Andreaeobryum macrosporum, a10–70 and b20–96: LSCM series of transverse sections through capsule at stage when archesporial cells (ar?) are not clearly differentiated (comparable with zone marked by bar in Fig. 20-56). Values indicate the distance (in µm) from the point where differentiation into endothecium and amphithecium becomes apparent in a series of sections from the sporophyte top. Note that endothecial part of sporophyte is composed by unequal cells, likely due to sectoral structure of the endothecium, comparable with that in Andreaea (‘A’, ‘C’). Amphithecial cells keep a moderately regular structure from 2:2:1 to 4:4:2 pattern, known in double opposite arthrodontous mosses (in a10 and a14 marked by colors: red –IPL homolog, green–PPL, cyan–OPL, cf. Ignatov et al., 2015). A–C:Andreaea rupestris cap- sule at different stages of the sporogenesis. A: longitudinal section in the beginning of the archesporial cells differentiation, from Ruhland (1924). B: transverse section roughly corresponging to the stage shown in ‘C’ (from Kuhn, 1874); C: longitudinal section of capsule b96 with differentiated archesporial cells, comparable with those shown in Fig.27 (from Kuhn, 1874).	Fig. 26. Andreaeobryum macrosporum, a10–70 and b20–96: LSCM series of transverse sections through capsule at stage when archesporial cells (ar?) are not clearly differentiated (comparable with zone marked by bar in Fig. 20-56). Values indicate the distance (in µm) from the point where differentiation into endothecium and amphithecium becomes apparent in a series of sections from the sporophyte top. Note that endothecial part of sporophyte is composed by unequal cells, likely due to sectoral structure of the endothecium, comparable with that in Andreaea (‘A’, ‘C’). Amphithecial cells keep a moderately regular structure from 2:2:1 to 4:4:2 pattern, known in double opposite arthrodontous mosses (in a10 and a14 marked by colors: red –IPL homolog, green–PPL, cyan–OPL, cf. Ignatov et al., 2015). A–C:Andreaea rupestris cap- sule at different stages of the sporogenesis. A: longitudinal section in the beginning of the archesporial cells differentiation, from Ruhland (1924). B: transverse section roughly corresponging to the stage shown in ‘C’ (from Kuhn, 1874); C: longitudinal section of capsule b96 with differentiated archesporial cells, comparable with those shown in Fig.27 (from Kuhn, 1874).	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426885/files/figure.png	https://doi.org/10.5281/zenodo.15426885	Fig. 27. Andreaeobryum macrosporum (LSCM), longitudinal sections of capsule at the stage of differentiation of archesporial cells (cf. also Fig. 28), showing discontinuous layer of archesporial cells and columella connectives with the spore sac in both lateral and apical parts of the capsule (arrowed). Values indicate the distance (in µm) from the first section, denoted as 0, where archesporial cells first appear among cells of the capsule wall (0 is not shown).	Fig. 27. Andreaeobryum macrosporum (LSCM), longitudinal sections of capsule at the stage of differentiation of archesporial cells (cf. also Fig. 28), showing discontinuous layer of archesporial cells and columella connectives with the spore sac in both lateral and apical parts of the capsule (arrowed). Values indicate the distance (in µm) from the first section, denoted as 0, where archesporial cells first appear among cells of the capsule wall (0 is not shown).	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426887/files/figure.png	https://doi.org/10.5281/zenodo.15426887	Fig. 28. Andreaeobryum macrosporum (116-126: LSCM, A–F: LM). 116–126: part of the series shown in Fig. 27, focusing on the uppermost part of the capsule, where the archesporium is not continuously overarching the columella. A: transverse section of mature capsule showing connectives from columella to the urn wall; B–C: Shiff staining indicating that capsule walls and archesporial cells are rich in hexoses. D: longisection of the capsule, showing part of columella (or a connective from columella to outer spore sac) attached to the urn near the top of spore chamber. E-F: Hand section of capsule, showing columella with membranaceous projections.	Fig. 28. Andreaeobryum macrosporum (116-126: LSCM, A–F: LM). 116–126: part of the series shown in Fig. 27, focusing on the uppermost part of the capsule, where the archesporium is not continuously overarching the columella. A: transverse section of mature capsule showing connectives from columella to the urn wall; B–C: Shiff staining indicating that capsule walls and archesporial cells are rich in hexoses. D: longisection of the capsule, showing part of columella (or a connective from columella to outer spore sac) attached to the urn near the top of spore chamber. E-F: Hand section of capsule, showing columella with membranaceous projections.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437411/files/figure.png	https://doi.org/10.5281/zenodo.15437411	Fig. 29 (previous page). Andreaea rupestris (A–C: LM, D–K: LCSM). Young capsule view (A) and its longisections (B, with close up and partial spore / remove in C), illustrating connection (arrowed) of columella (Co) with the spore chamber top in ‘C’. ‘D’ and ‘Da’ illustrate columella totally overarching by mature spores (capsule at stage close to dehiscence). ‘E’–‘K’ illustrate the upper sectors of undifferentiated endothecium (arrowed) in a position distally from the archesporial tissue (ar), which may form a connection with the top of spore chamber as in ‘C’.	Fig. 29 (previous page). Andreaea rupestris (A–C: LM, D–K: LCSM). Young capsule view (A) and its longisections (B, with close up and partial spore / remove in C), illustrating connection (arrowed) of columella (Co) with the spore chamber top in ‘C’. ‘D’ and ‘Da’ illustrate columella totally overarching by mature spores (capsule at stage close to dehiscence). ‘E’–‘K’ illustrate the upper sectors of undifferentiated endothecium (arrowed) in a position distally from the archesporial tissue (ar), which may form a connection with the top of spore chamber as in ‘C’.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426889/files/figure.png	https://doi.org/10.5281/zenodo.15426889	Fig. 30. Andreaeobryum macrosporum (A–H: SM). Photographs are taken from plants stored in glutaraldehyde, without any staining, under halogen light from KL1500 LCD illuminator, on dark grey background, except ‘D’, which was on white background), additionally contrasting light absorption capacity of the spores, as compared with the urn base; note that ‘D’ and ‘E’ is the same capsule, differing only by background color. Note also bulging surface (arrowed) of the foot (F) in ‘C’. Columella (Co in ‘E’) is strongly shrunk at this stage (cf. with Fig. 24). More dense perine in abortive spores (‘G’) makes them deeper in color, comparatively with well developed spores in ‘H’, where perine layer is quite uneven.	Fig. 30. Andreaeobryum macrosporum (A–H: SM). Photographs are taken from plants stored in glutaraldehyde, without any staining, under halogen light from KL1500 LCD illuminator, on dark grey background, except ‘D’, which was on white background), additionally contrasting light absorption capacity of the spores, as compared with the urn base; note that ‘D’ and ‘E’ is the same capsule, differing only by background color. Note also bulging surface (arrowed) of the foot (F) in ‘C’. Columella (Co in ‘E’) is strongly shrunk at this stage (cf. with Fig. 24). More dense perine in abortive spores (‘G’) makes them deeper in color, comparatively with well developed spores in ‘H’, where perine layer is quite uneven.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426891/files/figure.png	https://doi.org/10.5281/zenodo.15426891	Fig. 31. Andreaeobryum macrosporum (A–C: LM, D–F: TEM). Exothecium structure. A: Lower part of capsule, showing exothecium without stomata, but with a rugose cover, probably of dried mucilage. B: Cover view of upper part of capsule, showing longitudinal thickenings, which sections are shown in ‘C–F’. C: Transverse section of the capsule wall, showing thickenings, magnified in ‘D–F’. D–F: Sections of the longitudinal thickenings in exothecium: paradermal, longitudinal (somewhat oblique) and transversal, respectively. Combination of the dark osmiophilic vermicular structures embedded in polysaccharide matrix (white medium) provides an high level of light absorption, responsible for the black color of capsules in dry state.	Fig. 31. Andreaeobryum macrosporum (A–C: LM, D–F: TEM). Exothecium structure. A: Lower part of capsule, showing exothecium without stomata, but with a rugose cover, probably of dried mucilage. B: Cover view of upper part of capsule, showing longitudinal thickenings, which sections are shown in ‘C–F’. C: Transverse section of the capsule wall, showing thickenings, magnified in ‘D–F’. D–F: Sections of the longitudinal thickenings in exothecium: paradermal, longitudinal (somewhat oblique) and transversal, respectively. Combination of the dark osmiophilic vermicular structures embedded in polysaccharide matrix (white medium) provides an high level of light absorption, responsible for the black color of capsules in dry state.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426893/files/figure.png	https://doi.org/10.5281/zenodo.15426893	Fig. 32. Andreaeobryum macrosporium spores and capsule surfaces (SEM). A: Outer surface of capsule.B: Capsule from inside; note fine papillae on the outer spore sac surface. C: Longitudinal crack of capsule wall, showing incrassate exothecium. D:A split on the lower part of capsule wall with a spore. E: Separate valve with inrolled margins. F: Capsule from inside, showing ornamentation similar to that on spores.G–I:spores, showing variation. J: Surface of spore somewhat collapsed under vacuum: smooth intine (I2) is seen among finely granulose perine (P). K–L: spore split into two halves, apparently along an internal cell wall, still not incrassate.	Fig. 32. Andreaeobryum macrosporium spores and capsule surfaces (SEM). A: Outer surface of capsule.B: Capsule from inside; note fine papillae on the outer spore sac surface. C: Longitudinal crack of capsule wall, showing incrassate exothecium. D:A split on the lower part of capsule wall with a spore. E: Separate valve with inrolled margins. F: Capsule from inside, showing ornamentation similar to that on spores.G–I:spores, showing variation. J: Surface of spore somewhat collapsed under vacuum: smooth intine (I2) is seen among finely granulose perine (P). K–L: spore split into two halves, apparently along an internal cell wall, still not incrassate.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426895/files/figure.png	https://doi.org/10.5281/zenodo.15426895	Fig. 33. Andreaeobryum macrosporum spores (A: LM; B–F: TEM). A: Longitudinal section through capsule side, showing longitudinally incrassate exothecial cells distally and rather homogeneous isodiametric cells proximally; two spores are sectioned in their middle, one section (*) through sporoderm and one sectioned spore is abortive (a). B–F: Spore sections, showing variation in chloroplast (Ch) position, evenly spread within spore or more numerous near nucleus (N). All spores have numerous light vacuoles (Va), electron-dark lipid globules (Li), especially abundant near plasmalemma, granulate two-layered perine (P1 and P2), and usually homogenous layer formed by intine (I).	Fig. 33. Andreaeobryum macrosporum spores (A: LM; B–F: TEM). A: Longitudinal section through capsule side, showing longitudinally incrassate exothecial cells distally and rather homogeneous isodiametric cells proximally; two spores are sectioned in their middle, one section (*) through sporoderm and one sectioned spore is abortive (a). B–F: Spore sections, showing variation in chloroplast (Ch) position, evenly spread within spore or more numerous near nucleus (N). All spores have numerous light vacuoles (Va), electron-dark lipid globules (Li), especially abundant near plasmalemma, granulate two-layered perine (P1 and P2), and usually homogenous layer formed by intine (I).	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426897/files/figure.png	https://doi.org/10.5281/zenodo.15426897	Fig. 34. Andreaeobryum macrosporum, spore ultrastrucure details (TEM). Transverse sections, except the ‘D’ picture, showing paradermal section, with the electron-dense outer perine (P2) surrounding the inner perine (P1). Photos show a limited variation in sporoderm, composed of more or less homogeneous layer of intine, micro-granulate inner perine and granulate outer perine; plasmalemma is accompanied with numerous osmiophilic globules. F: Spore wall, showing weak differentiation into spotted outer layer and the lighter inner layer, both likely comprising intine, which is continued to the intercellular wall. G–H: a roll-shaped inclusion in the spore wall.	Fig. 34. Andreaeobryum macrosporum, spore ultrastrucure details (TEM). Transverse sections, except the ‘D’ picture, showing paradermal section, with the electron-dense outer perine (P2) surrounding the inner perine (P1). Photos show a limited variation in sporoderm, composed of more or less homogeneous layer of intine, micro-granulate inner perine and granulate outer perine; plasmalemma is accompanied with numerous osmiophilic globules. F: Spore wall, showing weak differentiation into spotted outer layer and the lighter inner layer, both likely comprising intine, which is continued to the intercellular wall. G–H: a roll-shaped inclusion in the spore wall.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426900/files/figure.png	https://doi.org/10.5281/zenodo.15426900	Fig. 35. Andreaeobryum macrosporium, details of the milticellular spore structure (TEM). A: Total section of three-celled spore. In all cells vacuoles (Va) are seen. B: Chloroplasts are seen in the central part, surrounded by vacuoles. C: Chloroplasts (Ch) along the sporoderm and intercellular wall. D–F show junctions of sporoderm with intercellular wall and round chloroplasts near these junctions.	Fig. 35. Andreaeobryum macrosporium, details of the milticellular spore structure (TEM). A: Total section of three-celled spore. In all cells vacuoles (Va) are seen. B: Chloroplasts are seen in the central part, surrounded by vacuoles. C: Chloroplasts (Ch) along the sporoderm and intercellular wall. D–F show junctions of sporoderm with intercellular wall and round chloroplasts near these junctions.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15437415/files/figure.png	https://doi.org/10.5281/zenodo.15437415	Fig. 36. Andreaeobryum macrosporium (previous page) (TEM). Sections of multicellular spore, with eleven cells seen (sections done at a slightly different levels). Light vacuoles and grey chloroplasts are seen in almost all cells and in one cell the nucleus is seen as well. Contrary to other spores, the inner perine is not continuous (‘A’), but seems to be split due to spore growth, forming clusters like usually outer perine does; the spore has an internal pool below the place of a putative spore wall rupture in the course of sporeling. Note the putative axillary hair in between cells (‘B, D-E’), pointed to narrow channel that is broadening to a pool of intercellular space, where no cellular organelles are discernible. Note also a putative alteration of the intine at the edge of the pool (arrowed).	Fig. 36. Andreaeobryum macrosporium (previous page) (TEM). Sections of multicellular spore, with eleven cells seen (sections done at a slightly different levels). Light vacuoles and grey chloroplasts are seen in almost all cells and in one cell the nucleus is seen as well. Contrary to other spores, the inner perine is not continuous (‘A’), but seems to be split due to spore growth, forming clusters like usually outer perine does; the spore has an internal pool below the place of a putative spore wall rupture in the course of sporeling. Note the putative axillary hair in between cells (‘B, D-E’), pointed to narrow channel that is broadening to a pool of intercellular space, where no cellular organelles are discernible. Note also a putative alteration of the intine at the edge of the pool (arrowed).	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426902/files/figure.png	https://doi.org/10.5281/zenodo.15426902	Fig. 37. Andreaeobryum macrosporum (A–C, E–F: TEM) and Andreaea rupestris (D: LM), spore details. A: Section of abortive spore; the spore is flattened and angulose, putatively shaped by pressing within the sporocyte wall in meiosis. No change in sporoderm layers in the area of the tetrahedral mark is seen, however, the characteristic structure of the sporoderm with intine and two-layered perine is well expressed (cf. ‘B’); the spore inner contents is dark, without obvious structure. B: Sporoderm ultratructure of the abortive spore, a close up from ‘A’. Intine is unevenly thickened, being at places 2–2.5 times thinner than in larger and normally developed spores. C: Abortive and cracked spore with numerous bacteria inside; the spore wall is ‘beaded’, with globules, in a way somewhat similar to extant Andreaea, cf ‘D’. D: Spores from living plants of Andreaea, with rosettes of chloroplasts; note that the sporoderm has a ‘beaded’ pattern, similar to that in abortive spore in ‘C’. E: Eleven-celled spore side (shown in Fig. 36) near the outer spore sac wall. Note the similarity between perine granules and granules on the spore sac surface: both groups include both dark and light granules; numerous bacteria are seen within the urn. F: Another view on spore and the inner surface of the outer spore sac, showing similarity of granules of their surfaces. Chloroplasts in spore are concentrated shortly below the sporoderm.	Fig. 37. Andreaeobryum macrosporum (A–C, E–F: TEM) and Andreaea rupestris (D: LM), spore details. A: Section of abortive spore; the spore is flattened and angulose, putatively shaped by pressing within the sporocyte wall in meiosis. No change in sporoderm layers in the area of the tetrahedral mark is seen, however, the characteristic structure of the sporoderm with intine and two-layered perine is well expressed (cf. ‘B’); the spore inner contents is dark, without obvious structure. B: Sporoderm ultratructure of the abortive spore, a close up from ‘A’. Intine is unevenly thickened, being at places 2–2.5 times thinner than in larger and normally developed spores. C: Abortive and cracked spore with numerous bacteria inside; the spore wall is ‘beaded’, with globules, in a way somewhat similar to extant Andreaea, cf ‘D’. D: Spores from living plants of Andreaea, with rosettes of chloroplasts; note that the sporoderm has a ‘beaded’ pattern, similar to that in abortive spore in ‘C’. E: Eleven-celled spore side (shown in Fig. 36) near the outer spore sac wall. Note the similarity between perine granules and granules on the spore sac surface: both groups include both dark and light granules; numerous bacteria are seen within the urn. F: Another view on spore and the inner surface of the outer spore sac, showing similarity of granules of their surfaces. Chloroplasts in spore are concentrated shortly below the sporoderm.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426904/files/figure.png	https://doi.org/10.5281/zenodo.15426904	Fig. 38. Andreaeobryum macrosporium, (A–D: LSCM; E-H: LM, acetolized). Acetolized spores show absence of sporopolennin layer (at least continuous enough), thus perine is breaking, partly retaining on P1 layer or outermost I2 layer. A–D: cellulose fluorescence in yellow and premature spore wall material in blue (berberin and DAPI staining). A certain stratification in intine layer is seen, demonstrating its complexity. D: series of micrographs with 2 µm interval, through multicellular spore.	Fig. 38. Andreaeobryum macrosporium, (A–D: LSCM; E-H: LM, acetolized). Acetolized spores show absence of sporopolennin layer (at least continuous enough), thus perine is breaking, partly retaining on P1 layer or outermost I2 layer. A–D: cellulose fluorescence in yellow and premature spore wall material in blue (berberin and DAPI staining). A certain stratification in intine layer is seen, demonstrating its complexity. D: series of micrographs with 2 µm interval, through multicellular spore.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
03C04D57FFCDFF9BEC31FE35FDB361BD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15426906/files/figure.png	https://doi.org/10.5281/zenodo.15426906	Fig. 39. Andreaeobryum macrosporium, (TEM). A–D: Chloroplasts in spores. E: Chloroplasts in capsule wall. F: Chloroplasts in columella. A: Transverse section, showing cristae and osmiophilic globules, arranged in two bundles. B: Curved chloroplast, with variously directed cristae and bundles of osmiophilic globules. C: Longitudinal section, showing two unequal bundles of osmiophilic globules. D: Transverse section, showing cristae arranged in several compartments in various directions, and osmiophilic globules corresponding to these compartments with only moderate reguliarity. E: Urn wall chloroplast with irregular lipid globules. F: Columella chloroplast with much larger and irregularly arranged osmiophilic globules.	Fig. 39. Andreaeobryum macrosporium, (TEM). A–D: Chloroplasts in spores. E: Chloroplasts in capsule wall. F: Chloroplasts in columella. A: Transverse section, showing cristae and osmiophilic globules, arranged in two bundles. B: Curved chloroplast, with variously directed cristae and bundles of osmiophilic globules. C: Longitudinal section, showing two unequal bundles of osmiophilic globules. D: Transverse section, showing cristae arranged in several compartments in various directions, and osmiophilic globules corresponding to these compartments with only moderate reguliarity. E: Urn wall chloroplast with irregular lipid globules. F: Columella chloroplast with much larger and irregularly arranged osmiophilic globules.	2016-12-31	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.		Zenodo	biologists	Ignatov, Michael S.;Ignatova, Elena A.;Fedosov, Vladimir E.;Ivanov, Oleg V.;Ivanova, Elena I.;Kolesnikova, Maria A.;Polevova, Svetlana V.;Spirina, Ulyana N.;Voronkova, Tatyana V.			
