taxonID	type	description	language	source
03C5F665F30EF456FF4C277ACB8CFDFD.taxon	type_taxon	Type species: Micreremaeus subnitidus Berlese, 1913	en	Ermilov, Sergey G., Kontschán, Jenő (2025): Description of an arboreal oribatid mite species of the genus Ovobates (Acari, Oribatida, Oribatulidae), based on adult and nymphs from Vietnam. Zootaxa 5740 (1): 112-124, DOI: 10.11646/zootaxa.5740.1.9, URL: https://doi.org/10.11646/zootaxa.5740.1.9
03C5F665F30EF456FF4C277ACB8CFDFD.taxon	description	Generic traits of Ovobates Adult. With character states of Oribatulidae (Balogh & Balogh 1984; Norton & Behan-Pelletier 2009). Measurements. Medium-sized (length about 350 – 450). Integument. Heavy sculpturing and ornamentation absent. Prodorsum. Rostrum rounded. Lamella absent or faintly lineate. Sublamella, translamella, prolamella, tutorium, sublamellar porose area absent. Dorsosejugal porose area present. Rostral, lamellar, and interlamellar setae medium-sized well developed, ro and le setiform, in setiform or slightly swollen distally. Exobothridial seta short, setiform. Bothridial seta short, nearly globular. Bothridium cup-shaped. Dorsophragma small. Notogaster. Anterior margin of notogaster distinct, convex medially. Pteromorph and pleural carina absent. Octotaxic system with four or five pairs of circular porose areas. Fourteen pairs of short, setiform notogastral setae, humeral region without setae. Gnathosoma. Subcapitulum diarthric. Palp setation 0 - 2 - 1 - 3 - 9 (+ ω). Solenidion bacilliform, attached to eupathidium. Axillary saccule absent. Chelicera chelate-dentate, with two setae. Lateral podosomal and epimeral regions. Epimeral setal formula 3 - 1 - 2 [3] - 1 [2,3]. Sejugal apodemal border usually connected with genital aperture. Humeral porose area Am absent, Ah present. Pedotectum I represented by medium-sized lamina, pedotectum II represented by small lamina. Discidium present. Circumpedal carina absent. Anogenital region. Four pairs of genital, one pair of aggenital, two pairs of anal, and three pairs of adanal setae. Adanal lyrifissure located anterior to anal aperture. Marginal porose area band-like. Legs. All legs with three strong claws. Porose area present on tarsi I – IV, tibiae I – IV, femora I – IV, and trochanters III, IV. Chaetome reduced (e. g., pv’ and pv’ ’ absent on from all leg tarsi). Juvenile instars. With prodorsal, gastronotic, and posterolateral macrosclerites. Opisthonotal gland dark brown. Notogaster and anogenital region partially plicate. Rostral, lamellar, and interlamellar setae well developed. Bothridial seta clavate or nearly globular. Exobothridial seta present. Interlamellar seta longest, exobothridial seta shortest. All gastronotic setae short, setiform, or different in length and morphology (e. g., short, medium-sized or long; setiform, bacilliform or subflagellate). Many gastronotic setae with excentrosclerite at base. Epimeral setal formulas: LA 3 - 1 - 2; PN, DN 3 - 1 - 2 - 1; TN 3 - 1 - 2 - 1 [2]. Anogenital setal formulas, LA to TN: 0 - 1 - 2 - 3, 0 - 0 - 1 - 1, 0 - 0 - 0 - 2, 0 - 0 - 3 - 3, respectively. Deutonymph with adanal sclerite, tritonymph with adanal and anal sclerites.	en	Ermilov, Sergey G., Kontschán, Jenő (2025): Description of an arboreal oribatid mite species of the genus Ovobates (Acari, Oribatida, Oribatulidae), based on adult and nymphs from Vietnam. Zootaxa 5740 (1): 112-124, DOI: 10.11646/zootaxa.5740.1.9, URL: https://doi.org/10.11646/zootaxa.5740.1.9
03C5F665F30DF45FFF4C202AC881FE35.taxon	description	(Figs 1 – 7)	en	Ermilov, Sergey G., Kontschán, Jenő (2025): Description of an arboreal oribatid mite species of the genus Ovobates (Acari, Oribatida, Oribatulidae), based on adult and nymphs from Vietnam. Zootaxa 5740 (1): 112-124, DOI: 10.11646/zootaxa.5740.1.9, URL: https://doi.org/10.11646/zootaxa.5740.1.9
03C5F665F30DF45FFF4C202AC881FE35.taxon	materials_examined	Type material. Holotype (male) and three paratypes (two males and one female) were collected from southern Vietnam, Dong Nai Province, Dong Nai Biosphere Reserve, Cat Tien National Park, 11 ° 25 ’ N, 107 ° 25 ’ E, about 130 m a. s. l., from branches of the Ochrocarpus sp. tree at the height of 25 m, 12. XII. 2024 (collected by V. M. Salavatulin). The holotype is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz, Germany. Three paratypes are deposited in the collection of the University of Tyumen, Museum of Zoology, Tyumen, Russia. All specimens are preserved in 70 % solution of ethanol with a drop of glycerol. Additional (non-type) material. Six adults (five males and one female) and 10 nymphs (three protonymphs, four deutonymphs, and three tritonymphs) were collected in the same location and on the same tree as the holotype and paratypes. All specimens (in 70 % solution of ethanol with a drop of glycerol) are in the personal collection of the first author.	en	Ermilov, Sergey G., Kontschán, Jenő (2025): Description of an arboreal oribatid mite species of the genus Ovobates (Acari, Oribatida, Oribatulidae), based on adult and nymphs from Vietnam. Zootaxa 5740 (1): 112-124, DOI: 10.11646/zootaxa.5740.1.9, URL: https://doi.org/10.11646/zootaxa.5740.1.9
03C5F665F30DF45FFF4C202AC881FE35.taxon	diagnosis	Diagnosis. Adult. Body length 390 – 435. Opisthonotal gland colourless. Body surface sparsely microfoveolate. Rostral, lamellar, and interlamellar setae long, setiform, barbed, interlamellar seta longest. Bothridial seta nearly globular, barbed. Five pairs of porose areas circular. Fourteen pairs of notogastral setae short, setiform, thin, nearly smooth. Epimeral setal formula 3 - 1 - 3 - 3. Leg femur II with five setae, genu IV with two setae. Juvenile instars. With well-developed prodorsal, gastronotic, and posterolateral macrosclerites. Opisthonotal gland dark brown. Rostral, lamellar, and interlamellar setae setiform, barbed, interlamellar seta longest. Bothridial seta clavate, barbed. Notogastral setae c 1, c 2, da, la, and dm short, setiform, thin, roughened, c 3, lm, p 2, and p 3 medium-sized to long, setiform, barbed, dp, h 1, and h 3 medium-sized, bacilliform, roughened, lp, h 2, and p 1 long, subflagellate, barbed. All setae except lm, p 2, and p 3 with small excentrosclerite at base. Epimeral setal formula 3 - 1 - 2 - 1 (PN, DN), 3 - 1 - 2 - 2 (TN). Deutonymph with faint adanal sclerite, tritonymph with faint adanal and anal sclerites.	en	Ermilov, Sergey G., Kontschán, Jenő (2025): Description of an arboreal oribatid mite species of the genus Ovobates (Acari, Oribatida, Oribatulidae), based on adult and nymphs from Vietnam. Zootaxa 5740 (1): 112-124, DOI: 10.11646/zootaxa.5740.1.9, URL: https://doi.org/10.11646/zootaxa.5740.1.9
03C5F665F30DF45FFF4C202AC881FE35.taxon	description	Description of adult (Figs 1 – 3; 7 A – H). Measurements. Body length 390 (holotype), 390, 405 (male paratypes), 435 (female paratype). Body width 225 (holotype), 210, 225 (male paratypes), 255 (female paratype). Integument (Fig. 7 A). Body colour brown. Opisthonotal gland colourless. Body surface sparsely microfoveolate. Subcapitular mentum, lateral part of prodorsum anterior to acetabulum I, and lateral part of epimere I with faint rugosity. Lateral part of body between bothridium and acetabula I – IV densely microgranulate. Prodorsum (Figs 1 A; 2 A; 7 C, D). Rostrum broadly rounded. Lamella lineate, poorly visible, located in front of bothridium. Rostral (52 – 56), lamellar (56 – 60), and interlamellar (77 – 90) setae setiform, barbed. Bothridial seta (37 – 45) nearly globular, barbed. Exobothridial seta (22 – 26) setiform, thin, roughened. Dorsosejugal porose area oval (7 – 9 × 4), located posterolateral to insertion of interlamellar seta. Notogaster (Figs 1 A; 2 A, B; 7 E). Anterior margin of notogaster distinct, slightly convex medially. Five pairs of porose areas circular (Aa 7; A 1 – A 4 4 – 7), A 1 located between lyrifissure im and seta lp. Fourteen pairs of notogastral setae (22 – 30) setiform, nearly smooth. Opisthonotal gland opening and all lyrifissures distinct. Gnathosoma (Figs 2 C – E; 7 F). Subcapitulum size 94 – 102 × 79 – 86. Three pairs of subcapitular setae (a 22 – 26; m 19 – 26; h 34 – 37) setiform, slightly barbed, m thinner than a and h. Two pairs of adoral setae (19) setiform, barbed. Palp length 75 – 82. Palp setal formula 0 - 2 - 1 - 3 - 9 (+ ω). Palptarsus solenidion connected with eupathidium, both located on tubercle. Postpalpal seta (6) spiniform. Chelicera length 105 – 120. Cheliceral setae (cha 30 – 34; chb 22 – 26) setiform, barbed. Lateral podosomal and epimeral regions (Figs 1 B; 2 A; 7 B). Humeral porose area Ah band-like, Am not visible. Pedotectum I medium-sized, pedotectum II small. Discidium subtriangular. Circumpedal carina absent. Sejugal apodemal border complete, connected with genital aperture. Epimeral setal formula 3 - 1 - 3 - 3. Epimeral setae (1 b, 3 b, 3 c, 4 a 26 – 37; 1 a, 1 c, 2 a, 3 a, 4 b, 4 c 22 – 30) setiform, roughened. Anogenital region (Figs 1 B; 2 A, B). Anogenital setal formula 4 - 1 - 2 - 3. Genital (17 – 19), aggenital (26 – 30), anal (19 – 22), and adanal (19 – 22) setae setiform, roughened. Adanal lyrifissure oblique or longitudinal, located close and anterior to anal aperture. Marginal porose area long, band-like. Legs (Figs 3 A – D; 7 G, H). Median claw slightly thicker than lateral claws. All claws slightly barbed on dorsal side. Lateral claws with tooth ventrodistally. Dorsoparaxial porose area on femora I – IV and on trochanters III, IV, proximoventral porose area on tarsi I – IV, and distoventral porose area on tibiae I – IV well visible. Often porose area on tarsi and tibia divided into two areas (Fig. 7 H). Formulas of leg setation and solenidia: I (1 - 4 - 2 - 4 - 14) [1 - 1 - 2], II (1 - 5 - 2 - 4 - 13) [1 - 1 - 1], III (2 - 3 - 1 - 2 [3] - 12) [1 - 1 - 0], IV (1 - 2 - 2 - 2 [3] - 10) [0 - 1 - 0]. Homology of leg setae and solenidia as indicated in Table 1. Many dorsal setae of all tarsi slightly swollen distally. Seta s of tarsus I setiform, barbed (not eupathidial). Description of nymphal instars (Figs 4 – 6; 7 I – O). Measurements. Body length of PN 255 – 270, DN 330 – 360, TN 375 – 435. Body width of PN 120 – 150, DN 165 – 195, TN 195 – 240. Integument (Figs 7 I, K, N). Prodorsal macrosclerite and epimeres (except sternal region) light yellowish to light brownish. Gastronotic and posterolateral macrosclerites light brown to greyish brown. Opisthonotal gland dark brown. Legs light yellowish to greyish and slightly brown. Surface outside of sclerites mainly colorless. Cuticle densely microporose (micropores poorly visible, but clearer on macrosclerites). Sternal region, anogenital region, and anterior part of gastronotic region sparsely folded. Prodorsum (Figs 4 A – D; 7 O). Relatively short, about 1 / 2 to 1 / 3 length of gastronotic region (measured in lateral aspect). Rostrum broadly rounded, with unclear medial protrusion. Dorsal side of prodorsum with large prodorsal macrosclerite. Rostral (PN 32 – 34; DN 37 – 41; TN 49 – 56), lamellar (PN 26; DN 34 – 37; TN 49 – 56), and interlamellar (PN 67 – 71; DN 71 – 82; TN 101 – 112) setae setiform, barbed. Bothridial seta (PN 30; DN 34; TN 41) clavate, barbed, stalk and head similar in length or head slightly longer. Exobothridial seta (PN 11 – 15; DN 15 – 17; TN 19) setiform, thin, roughened. Gastronotic region (Figs 4 A – D; 5 A – C; 7 J). Gastronotic and posterolateral macrosclerites slightly bordered. Ecdysial cleavage line distinct. Notogastral setae c 1, c 2, da, la, and dm setiform, thin, roughened, c 3, lm, p 2, and p 3 setiform, barbed, dp, h 1, and h 3 bacilliform, roughened, lp, h 2, and p 1 subflagellate, barbed. Setae da, la, dm, lm, dp, lp, h 1 – h 3, and p 1 located on macrosclerite. All setae except lm, p 2, and p 3 with small excentrosclerite at base. Length of gastronotic setae in PN: c 1, c 2, da, dm, la 11 – 15, p 3 19 – 22, p 2 30 – 34, dp, h 1, h 3 34 – 41, c 3 56 – 60, lm 56 – 64, lp, h 2, p 1 139 – 150; in DN: c 1, c 2, da, dm, la 15, p 3 30, p 2 41 – 45, dp, h 1, h 3 49, c 3 60 – 64, lm 71 – 75, lp, h 2, p 1 176 – 199; in TN: c 1, c 2, da, dm, la 22, p 3 37 – 49, p 2 64 – 82, dp, h 1, h 3 49 – 56, c 3 71 – 82, lm 86 – 101, lp, h 2, p 1 176 – 199. Opisthonotal gland opening located on posterolateral macrosclerite. Cupules appearing in normal ontogenetic pattern. Gnathosoma (Figs 5 D – F). Subcapitulum size: PN, DN 64 × 56; TN 75 × 67. Length of subcapitular setae: a and m: PN 11; DN 15; TN 19; h: PN 13; DN 22; TN 30. Length of adoral seta: PN 7; DN 11; TN 13. Subcapitular and adoral setae setiform, thin, roughened. Palp length: PN 41; DN 49; TN 60. Palp setal formula in all nymphs 0 - 2 - 1 - 3 - 9 (+ ω). Palptarsus solenidion coupled with eupathidium, both located on tubercle. Length of postpalpal seta: PN, DN 4; TN 6. Postpalpal seta spiniform. Axillary saccule absent. Chelicera length: PN 67; DN 75; TN 86. Length of cheliceral setae: cha: PN 19; DN 22; TN 26; chb: PN 11; DN 15; TN 19. Seta cha setiform, barbed, seta chb setiform, slightly barbed. Epimeral region (Figs 4 D; 5 A – C; 7 N). Epimeral setal formulas: PN, DN 3 - 1 - 2 - 1; TN 3 - 1 - 2 - 2. Length of setae: in PN: 1 b, 3 b 15; others 13; in DN: 1 b, 3 b, 4 a 15 – 19; others 13 – 15; in TN: 1 b, 4 a 26 – 30; 3 b 19 – 22; others 13 – 15. All setae setiform, thin, roughened. Anogenital region (Figs 4 D; 5 A – C; 7 I). Deutonymph with longitudinally elongate adanal sclerite, tritonymph with longitudinally elongate adanal and anal sclerites, sclerites poorly visible. Ontogeny of genital (PN 9 – 11; DN 11; TN 13 – 15), aggenital (DN 15; TN 19), anal (TN 19 – 22), and adanal (DN 22; TN 26 – 30) setal formulas, PN to TN: 1 - 2 - 3, 0 - 1 - 1, 0 - 0 - 2, 0 - 3 - 3, respectively. All setae setiform, thin, roughened. Paraproctal setae absent. Adanal cupule appearing in normal ontogenetic pattern. Legs (Figs 6 A – D; 7 L, M). Claw of each leg strong, slightly barbed on dorsal side. Pretarsus of all legs well developed, longest in leg IV. Slight furrow present between setae (u) and s. Ventroparaxial porose area on femora I – IV well visible. Porose area on trochanters and tibiae absent. Proximoventral porose area on tarsi I – III well visible. Formulas of leg setation and solenidia: PN I (0 - 2 - 2 - 3 - 12) [1 - 1 - 2], II (0 - 2 - 2 - 2 - 11) [1 - 1 - 1], III (0 - 2 - 1 - 1 - 10) [1 - 1 - 0], IV (0 - 0 - 0 - 0 - 7) [0 - 0 - 0]; DN I (0 - 2 - 2 - 3 - 12) [1 - 1 - 2], II (0 - 2 - 2 - 2 - 11) [1 - 1 - 1], III (1 - 2 - 1 - 1 - 10) [1 - 1 - 0], IV (0 - 2 - 1 - 1 - 10) [0 - 1 - 0]; TN I (1 - 4 - 2 - 3 - 14) [1 - 1 - 2], II (1 - 4 - 2 - 3 - 13) [1 - 1 - 1], III (1 - 2 - 1 - 1 [2] - 12) [1 - 1 - 0], IV (0 - 2 - 1 - 1 [2] - 10) [0 - 1 - 0]. Homology of leg setae and solenidia as indicated in Table 1.	en	Ermilov, Sergey G., Kontschán, Jenő (2025): Description of an arboreal oribatid mite species of the genus Ovobates (Acari, Oribatida, Oribatulidae), based on adult and nymphs from Vietnam. Zootaxa 5740 (1): 112-124, DOI: 10.11646/zootaxa.5740.1.9, URL: https://doi.org/10.11646/zootaxa.5740.1.9
03C5F665F30DF45FFF4C202AC881FE35.taxon	discussion	Remarks. The adult of the known species of Ovobates — O. ochrocarpicus sp. nov., O. subnitidus (see Berlese 1913; Mahunka 1994), and O. khaustovi (see Ermilov & Salavatulin, 2025) — differs from each other by the body surface, the morphology of the interlamellar seta, the number of the notogastral porose areas, the colour of the opisthonotal gland, and the epimeral and leg setations (Table 2). The nymphal instars of O. ochrocarpicus sp. nov. and O. khaustovi (see Ermilov & Salavatulin, 2025) differ from each other by the morphology of the bothridial and interlamellar setae, the length and morphology of the notogastral setae, and the epimeral and leg setations (Table 3).	en	Ermilov, Sergey G., Kontschán, Jenő (2025): Description of an arboreal oribatid mite species of the genus Ovobates (Acari, Oribatida, Oribatulidae), based on adult and nymphs from Vietnam. Zootaxa 5740 (1): 112-124, DOI: 10.11646/zootaxa.5740.1.9, URL: https://doi.org/10.11646/zootaxa.5740.1.9
03C5F665F30DF45FFF4C202AC881FE35.taxon	etymology	Etymology. The species name ochrocarpicus refers to the tree (Ochrocarpus sp.) on which the new species was found.	en	Ermilov, Sergey G., Kontschán, Jenő (2025): Description of an arboreal oribatid mite species of the genus Ovobates (Acari, Oribatida, Oribatulidae), based on adult and nymphs from Vietnam. Zootaxa 5740 (1): 112-124, DOI: 10.11646/zootaxa.5740.1.9, URL: https://doi.org/10.11646/zootaxa.5740.1.9
