taxonID	type	description	language	source
03C4E90E8B546171BA5DF9D9962DFAC9.taxon	description	urn: lsid: zoobank. org: act: 054 CFE 2 B- 7 E 7 D- 4 FB 1 - A 55 C-B 9 E 5 E 9 DB 60 F 0 Figs 2 – 4	en	Brazidec, Manuel, Aupérin, Juliette, Perrichot, Vincent (2026): A new epyrine wasp (Hymenoptera: Bethylidae) from the late Eocene of Europe. European Journal of Taxonomy 1052: 169-184, DOI: 10.5852/ejt.2026.1052.3273, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3273/14452
03C4E90E8B546171BA5DF9D9962DFAC9.taxon	diagnosis	Diagnosis Male Frons protruding anteriorly into medial projection hiding antennal foramina and lateral clypeal lobe (Figs 2 A, 3 A – B); lateral clypeal lobe inconspicuous; malar space much reduced (Fig. 3 A – B); mandible tridentate, basally with laminar expansion (Fig. 3 A); antennal foramen directed anteroventrally (Fig. 3 B); scape length 3.0 × medial width (Fig. 3 C); pedicel longer than flagellomere 1 (Fig. 3 C); flagellomere 1 as long as wide (Fig. 3 C); scape bearing sparse long setae (Fig. 3 C); lateral pronotal area concave, pronotal lobe small (Fig. 3 E); anteromesoscutum half as long as dorsal pronotal area, as long as mesoscutellum (Fig. 3 D); notaulus drop-shaped, deeply impressed posteriorly, faint anteriorly (Fig. 3 D); parapsidal signum present on half anteromesoscutum length (Fig. 3 D); foveae of mesoscuto-mesoscutellar suture separated by 6.0 × fovea width (Fig. 3 D); mesoscutellum not contacting metapectal-propodeal complex (Fig. 3 D); transepisternal line connecting with mesopleural pit (Fig. 3 E); metapostnotal median carina present (Fig. 3 D); metapostnotal-propodeal suture slender; 2 r-rs & Rs curved towards wing tip (Figs 3 F, 4); profemur strongly swollen, flattened (Fig. 3 E, G); mesotibia with longitudinal rows of spines (Fig. 3 H); mesotibial spurs longer than mesotibia diameter (Fig. 3 H).	en	Brazidec, Manuel, Aupérin, Juliette, Perrichot, Vincent (2026): A new epyrine wasp (Hymenoptera: Bethylidae) from the late Eocene of Europe. European Journal of Taxonomy 1052: 169-184, DOI: 10.5852/ejt.2026.1052.3273, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3273/14452
03C4E90E8B546171BA5DF9D9962DFAC9.taxon	etymology	Etymology The specific epithet is a patronym honouring Carsten Gröhn, who generously made the specimen available for study. The specific epithet is to be treated as a noun in the genitive case.	en	Brazidec, Manuel, Aupérin, Juliette, Perrichot, Vincent (2026): A new epyrine wasp (Hymenoptera: Bethylidae) from the late Eocene of Europe. European Journal of Taxonomy 1052: 169-184, DOI: 10.5852/ejt.2026.1052.3273, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3273/14452
03C4E90E8B546171BA5DF9D9962DFAC9.taxon	materials_examined	Type material Holotype RUSSIA • ♂ (complete specimen in amber); Yantarny, Sambia Peninsula, Kaliningrad region; GPIH 5021 (CCGG 9223). Type locality and age Yantarny, Sambia Peninsula, Kaliningrad region, Russia. Late Eocene: Priabonian, 34 – 38 Ma.	en	Brazidec, Manuel, Aupérin, Juliette, Perrichot, Vincent (2026): A new epyrine wasp (Hymenoptera: Bethylidae) from the late Eocene of Europe. European Journal of Taxonomy 1052: 169-184, DOI: 10.5852/ejt.2026.1052.3273, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3273/14452
03C4E90E8B546171BA5DF9D9962DFAC9.taxon	description	Description BODY. Stout, length 6.67 mm. HEAD. Slightly longer than wide, flattened, prognathous; LH = 1.27 mm, WH = 1.12 mm, WF = 0.90 mm, HE = 0.44 mm, OOL = 0.49 mm, VOL = 0.45 mm, WOT = 0.24 mm, DAO = 0.07 mm; frons flat to slightly convex in lateral view, imbricate, bearing sparse long setae, protruding anteriorly into medial, not elevated, frontal projection overlapping antennal foramina; frontal line absent; vertex flat to slightly convex in lateral view, imbricate, bearing sparse long setae dorsally; vertex crest bluntly angulate as seen in dorsal view; compound eye developed, barely protruding from side of head, located anteriorly on head, closer to mandible than to vertex crest, covering ca 0.3 × side of head, glabrous; gena with sparse long setae, genal setae longer than frontal setae; lateral clypeal lobe inconspicuous, median clypeal lobe outcurved, much hidden under frontal projection and scape; malar space inconspicuous, shorter than basal mandibular width; mandible curved, basally with laminar expansion, bearing setae longer than basal mandibular width, tridentate; labial palp as long as mandible; antennal foramen directed anteroventrally; antenna filiform; radicle and scape axes distinct; scape length 3.0 × medial width (length 0.44 mm), wider medially than basally and distally, basally forming flangelike expansion hiding radicle; pedicel pyriform, as long as wide (length 0.11 mm), longer than flagellomere 1; flagellomere 1 shortest flagellomere, wider than long (length 0.09 mm); flagellomeres 2 – 10 longer than wide (length ca 0.15 – 0.20 mm); flagellomere 11 longest flagellomere (length 0.25 mmº), tapering apicad; scape bearing sparse long setae, setae length about 0.5 × scape width; pedicel and flagellum bearing uniform short setae, setae length shorter than 0.5 × flagellomere width; ocellar triangle much posterad on head; anterior ocellus posterad supra-ocular line; occipital carina present. MESOSOMA. Shorter than metasoma (length 2.44 mm); propleuron not visible in dorsal view; propleural epicoxal sulcus circular; dorsal pronotal area slightly convex (length 0.77 mm), bearing sparse long setae, ecarinate, anterior and lateral margins outcurved, humeral angle not pronounced, posterior margin incurved; lateral pronotal area concave; pronotal lobe small, reaching tegula; anteromesoscutum flat to slightly convex, length 0.5 × dorsal pronotal area length (length 0.38 mm); notaulus present, drop-shaped, deeply impressed and wide posteriorly, faint and present only as line anteriorly, notauli posteriorly separated by less than distance between foveae of mesoscuto-mesoscutellar suture; parapsidal signum present as conspicuous sulcus, effaced anteriorly and posteriorly; mesoscuto-mesoscutellar suture with foveae not connected by sulcus, foveae oval, separated one to another by 6.0 × fovea width; mesoscutellum as long as anteromesoscutum (length 0.38 mm); metanotum reduced, distinct medially (i. e., mesoscutellum not contacting metapectal-propodeal complex); mesopleuron convex, with transepisternal line deeply impressed, anteriorly connecting with mesopleural pit, mesopleural pit separated from mesepimeral sulcus by about its diameter; metapectal-propodeal disc longer than wide, flat to slightly convex; metapostnotum rugulose; metapectal-propodeal disc with sculpture indistinct, anteriorly margined by transverse anterior carina of metapectal-propodeal complex, laterally margined by lateral marginal carina of metapectal-propodeal complex, posteriorly margined by transverse posterior carina of metapectal-propodeal complex; propodeum laterally slightly concave, thinly striated; metapostnotal median carina present, complete, strong; metapostnotal-propodeal suture present at least anteriorly, slender; lateroposterior propodeal angle angulate. WINGS. Fore wing developed (LFW = 3.73 mm), hyaline, uniformly micropubescent; veins C, Sc + R, M + Cu, 1 A, 1 Rs & 1 M, cu-a, 2 r-rs & Rs tubular; M + Cu shorter than A and Sc + R, forming obtuse angles with 1 Rs & 1 M and cu-a; 1 Rs & 1 M and cu-a confluent; 1 Rs & 1 M slightly incurved medially; cu-a strongly bent toward A posteriorly; 1 A not tubular distad cu-a, only thin spectral segment of 2 A visible; 2 r-rs & Rs without angulation, curved towards wing tip, distalmost section almost straight, fading before reaching margin, at distance subequal to pterostigma length; pterostigma distinct, slightly thickened; no other veins present but patterns of flexion lines visible in medial and distal sections of wing. Hind wing shorter than fore wing (length 1.72 mm), hyaline, micropubescent, posteriorly bordered by long setae; anal lobe delimitated by distinct incision, length ca 0.2 × hind wing length. LEGS. Profemur much swollen, length ca 1.8 × width, flattened, almost paddle-shaped, bearing sparse long setae; protibia fusiform, bearing uniform short setae and sparse long setae; strigil present; protarsomeres bearing uniform short setae; probasitarsomere longest, distal protarsomere longer than 0.5 × probasitarsomere length; mesofemur fusiform, length ca 2.7 × width, bearing uniform short setae; mesotibia fusiform, bearing uniform short setae and sparse spines on every faces and along distal rim, spines as long as setae, apparently organized in longitudinal rows; mesotarsomeres bearing uniform short setae; mesobasitarsomere longest; metafemur fusiform, length ca 3.5 × width, bearing uniform short setae; metatibia bearing uniform short setae; metatarsomeres bearing uniform short setae; metabasitarsomere longest; tibial spur formula 1 – 2 – 2; protibia with calcar; mesotibial spurs longer than mesotibia diameter; tarsal claws evenly curved, edentate; arolium small. METASOMA. Fusiform (length 2.96 mm), imbricate, bearing sparse long pubescence, especially near posterior margin of sternites; tergite 2 longest tergite, length 1.6 × tergite 3 length.	en	Brazidec, Manuel, Aupérin, Juliette, Perrichot, Vincent (2026): A new epyrine wasp (Hymenoptera: Bethylidae) from the late Eocene of Europe. European Journal of Taxonomy 1052: 169-184, DOI: 10.5852/ejt.2026.1052.3273, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3273/14452
03C4E90E8B586171BA61FA26969EF854.taxon	discussion	Remarks Azevedo et al. (2018: 172) proposed the replacement name Holepyris neodubius Azevedo, 2018 for Holepyris dubius Kieffer, 1904 rather than † Holepyris dubius (Brues, 1933) (Azevedo et al. 2018: 9, 171). However, this action was based on a misunderstanding of the homonymy: H. dubius Kieffer, 1904 is the senior homonym, and † H. dubius (Brues, 1933) is the junior homonym. Therefore, Holepyris neodubius Azevedo, 2018 is an unnecessary replacement name for H. dubius Kieffer, 1904. Furthermore, † H. dubius (Brues, 1933) was incorrectly named † Holepyris neodubius Azevedo, 2018 in Colombo et al. (2021 a) and † Rysepyris neodubius (Azevedo, 2018) in Colombo et al. (2022: table S 18).	en	Brazidec, Manuel, Aupérin, Juliette, Perrichot, Vincent (2026): A new epyrine wasp (Hymenoptera: Bethylidae) from the late Eocene of Europe. European Journal of Taxonomy 1052: 169-184, DOI: 10.5852/ejt.2026.1052.3273, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3273/14452
03C4E90E8B596170B8C9FE8E909BFC76.taxon	description	replacement name for † Rysepyris dubius (Brues, 1933)	en	Brazidec, Manuel, Aupérin, Juliette, Perrichot, Vincent (2026): A new epyrine wasp (Hymenoptera: Bethylidae) from the late Eocene of Europe. European Journal of Taxonomy 1052: 169-184, DOI: 10.5852/ejt.2026.1052.3273, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3273/14452
03C4E90E8B596170B8C9FE8E909BFC76.taxon	etymology	Etymology The replacement name is a patronym honouring Charles T. Brues, who originally described this Baltic amber species.	en	Brazidec, Manuel, Aupérin, Juliette, Perrichot, Vincent (2026): A new epyrine wasp (Hymenoptera: Bethylidae) from the late Eocene of Europe. European Journal of Taxonomy 1052: 169-184, DOI: 10.5852/ejt.2026.1052.3273, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3273/14452
03C4E90E8B596170B8C9FE8E909BFC76.taxon	materials_examined	Type material Holotype RUSSIA • ♂ (specimen in amber); housed in the University of Königsberg; specimen without code (lost). Locality and age Russia. Late Eocene: Priabonian, 34 – 38 Ma.	en	Brazidec, Manuel, Aupérin, Juliette, Perrichot, Vincent (2026): A new epyrine wasp (Hymenoptera: Bethylidae) from the late Eocene of Europe. European Journal of Taxonomy 1052: 169-184, DOI: 10.5852/ejt.2026.1052.3273, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3273/14452
