taxonID	type	description	language	source
03C4A00CE176D22EFF2DFDA0FE64FA57.taxon	description	The studied amber specimen shows the combination of characters unequivocally corresponding to adults of the subfamily Scraptiinae (Lawrence & Ślipiński 2010; Johnston et al. 2024): protrochantins exposed, head without distinct transverse carina separating occiput from posterior face of head capsule, temples present, emarginate eyes bearing interfacetal setae, pronotum and elytra without transverse strigae, pronotum with pair of basal pits or depressions, tibiae slender, and all tibial and tarsal segments lacking crenulation. The subfamily Scraptiinae (recent) is divided into two tribes: Allopodini, characterized by the absence of a lobe on the penultimate metatarsomere, and Scraptiini, which possess a lobed third metatarsomere. The Mesozoic specimen under consideration is formally assigned to the extant tribe Allopodini, based on its linear metatarsi that lack lobed tarsomeres.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE176D22FFF2DFA24FD93FE43.taxon	materials_examined	Type species: Cretoscraptella crucifera sp. nov., designated herein	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE176D22FFF2DFA24FD93FE43.taxon	diagnosis	Diagnosis. Cretoscraptella gen. nov. can be distinguished from all other described Allopodini genera of Recent (Allopoda LeConte, 1866; Pectotoma Hatch, 1965; and Pseudoscraptia Wollaston, 1868) by the following characters: distinctly smaller body size, narrow temples (in contrast to eye attaining posterior margin of head in Allopoda and eye separated by nearly the width of one eye from posterior margin of head in Pectotoma), comparatively short elytra (in contrast to elytra narrow and long, twice or more as long as wide in Pectotoma, Allopoda or Pseudoscraptia), filiform antennae (flabellate in male of Pectotoma), and not shortened last metatarsomere (in contrast to shortened last metatarsomere in Pseudoscraptia). Among extinct genera of Scraptiidae, the new genus can be distinguished by smaller body size, 1.5 mm (Scraptiomima is 2.5 – 8.6 mm long, Palaeoscraptia is 2.0 mm long, Archescraptia is 4.3 mm long) and absence of lobed sements in metatarsi. Cretoscraptella gen. nov. differs from Scraptiomima in filiform antennae with elongate anennomeres (short, almost moniliform antenna with transverse antennomeres in Scraptiomima), in comparatively shorter tarsi and shorter tibial spurs.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE176D22FFF2DFA24FD93FE43.taxon	etymology	Derivatio nominis. The name of the new genus is a compound word and combines the Creto- (referring to the Cretaceous age of the deposit), Scraptia (name of the type genus in the subfamily) and the Latin diminutive suffixella (referring to the small size of the fossil). Gender feminine.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE176D22FFF2DFA24FD93FE43.taxon	discussion	Remarks. The new genus is monotypic, represented by the type species only. Therefore the generic description considerably overlaps that of the species.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE177D22AFF2DFE10FEB1F879.taxon	description	(Figs 1 – 4)	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE177D22AFF2DFE10FEB1F879.taxon	materials_examined	Material examined. Holotype: No. JH / BUR / SCRAP / 24 [JHAC], adult, sex unknown. Complete beetle inclusion included in transparent amber piece with dimensions 15 × 12 × 3 mm. Syninclusions are represented by numerous small to minute organic particles and one specimen of Archiculicoides sp. (Diptera: Nematocera: Ceratopogonidae). Type strata. Burmese amber, mid-Cretaceous. Type locality. Hukawng Valley, Tanai Township, Myitkyina District, Kachin State, Myanmar.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE177D22AFF2DFE10FEB1F879.taxon	diagnosis	Diagnosis. As for the genus (vide supra).	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE177D22AFF2DFE10FEB1F879.taxon	description	Description. Measurements: body length (from anterior margin of head to elytral apex along midline) 1.5 mm; body maximum width across elytra 0.6 mm; head width (across eyes) 0.4 mm; head width (between eyes) 0.2 mm; pronotal length 0.3 mm; pronotal maximum width 0.6 mm; pronotal width at base 0.5 mm; elytral length 1.1 mm; elytral maximum width 0.6 mm. Body ovoid, elongate, about 2.2 × as long as wide; weakly convex dorsally and ventrally; bicolorous (as preserved): head, appendages, underside and pronotum brown, elytra with dark pattern, forming cross consisting of one vertical and two horizontal bars, i. e., one longitudinal (along suture) and two transverse (in basal and in medial parts of elytra). Pubescence: dorsum homogenously covered with fine, semirecumbent hairs, not forming patches. Head declined, abruptly constricted posteriorly, widest across eyes, with narrow temples, densely punctate, without distinct transverse carina separating occiput from posterior face of head capsule. Compound eyes large, prominent, emarginate anteriorly at antennal base, with short interfacetal setation, widely separate. Mandible short and broad, bidentate. Antennal insertions exposed from above. Antennae 11 - segmented, filiform, comparatively short, extending to about base of pronotum; scape and pedicel short, rounded; antennomeres 3 – 10 elongate, subconical; anennomere 11 longest, spindle-shaped, pointed at apex, about 3.3 × as long as wide. Maxillary palp large, about 1.5 × shorter than antenna, with apical maxillary palpomere elongate securiform. Apical labial palpomere small, expanded, with emargination apically. Pronotum distinctly transverse, 0.5 × as long as wide, widest in posterior one-quarter and distinctly narrowed anteriad, laterally margined (at least in posterior one-half); pronotal disc weakly convex, with two paired basal pits; pronotal surface densely punctate. Anterior pronotal margin arcuate; posterior margin trisinuate (concave against scutellar shield and in areas of basal pronotal pits). Posterior pronotal angles rectangular, rounded. Prosternal process incomplete, narrowed apically. Procoxal cavities narrowly separated, open externally. Scutellar shield minute, triangular, not pubescent. Elytra oval, tapered at apex, about 1.8 × as long as wide combined, moderately convex, 3.7 × as long as pronotum. Humeral angles rounded, humeri well-developed. Elytral punctation rather large, irregular, distance between punctures 1.0 – 2.0 × as large as one puncture diameter. Epipleura well-developed, widest basally, slightly narrowed posteriad, reaching abdominal ventrite 1. Mesocoxal cavities closed, narrowly separated. Metaventrite with broad longitudinal carina, finely punctate. Metathoracic wings not apparent. Legs slender. Procoxae subcontiguous, projecting well below prosternum; mesocoxae large, circular; metacoxae transverse, narrowly separated. Protrochantin exposed. Femora and tibiae subequal in length. Tibiae straight, slightly dilated apically, finely setose along outer margin, with paired well-developed apical spurs; tibial spurs flattened, equal in length, seemingly pubescent but not serrate. Tarsal formula 5 – 5 – 4, tarsomeres with sparse, fine setation ventrally; pro- and mesotarsi with penultimate tarsomeres weakly widened, but not bilobed; metatarsi linear, without distinctly lobed tarsomeres. Abdomen with five visible ventrites, finelly punctate, with very short, recumbent pubescence. Relative length ratios of ventrites 1 – 5 equal to 4: 3: 1: 1: 2 (measured medially).	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE177D22AFF2DFE10FEB1F879.taxon	etymology	Derivatio nominis. The specific epithet refers to the bicolored elytral pattern of the specimen, looking like black double-barred Jagiellonian cross on a brown background. It is formed after the Latin adjective “ crucifer, - a, - um ” (cross bearing).	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE17CD224FF2DFDE8FEB1F85F.taxon	description	(Figs 5 – 6)	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE17CD224FF2DFDE8FEB1F85F.taxon	materials_examined	Material examined. One specimen with collection number JDC- 13732 [JDC], adult, male. Complete beetle inclusion with partially exposed metathoracic wings and apical portion of aedeagus is included in transparent amber piece with dimensions 40 × 26 × 3 mm. Syninclusions are represented by three specimens of Nematocera (Diptera), few stellate Fagaceae trichomes. Strata. Baltic amber, most probable from the Eocene amber-bearing layers of Blaue Erde (Blue Earth) within the Prussian Formation; estimated age: middle-late Eocene (Bukejs et al. 2019). Locality. Yantarny village (formerly Palmnicken), Sambian (Samland) Peninsula, Kaliningrad Region, western Russia.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE17CD224FF2DFDE8FEB1F85F.taxon	description	Note. Body length of beetle JDC- 13732 (including visible part of head) 4.8 mm, maximum body width 1.3 mm. Although the specimen is partially obscured on the ventral side and the dorsal view is partially hidden by crack in amber matrix, no morphological differences were found, and the specimen is assigned to extinct Samlandotoma seidlitzi Alekseev, 2019. All tibiae of JDC- 13732 with paired pubescent apical spurs of equal length (tibial spur formula 1 - 1 - 2 of Samlandotoma is mistakenly reported in the original description), posterior pronotal angles triangular (“ obtuse, rounded ” is mistakenly reported for the holotype in the original description). Taxonomic assignment. Characters of Samlandotoma that preclude its original misplacement within Pelecotominae or Ripiphoridae have been discussed by Batelka (2021). The inclusion shows the combination of characters corresponding to adults of the subfamily Scraptiinae within Scraptiidae, namely: (1) body form elongate-oval with base of pronotum narrower than anterior elytral margin; (2) eyes emarginate anteriorly; (3) head constricted posteriorly; (4) antennal insertions exposed from above; (5) antennae 11 - segmented, non-clubbed; (6) temples present; (7) pronotum laterally margined in basal one-half of its lenght; (8) pronotum and elytra without transverse strigae; (9) procoxae subcontiguous, projecting well below prosternum; (10) protrochantins exposed; (11) elytra irregularly punctate, without transverse strigae; (12) tibiae slender, all tibial and tarsal segments lacking crenulation; (13) tibial spurs paired, equal in length, well-developed, pubescent; (14) tarsi 5 - 5 - 4 with all penultimate tarsomeres lobed; (15) tarsal claws simple; and (16) abdomen with five ventrites and ventrite 1 not much longer than ventrite 2. Interfacetal setae in compound eyes are absent, posterior pair of pits or depressions in pronotum apparently not discernible. The genus Samlandotoma shares the unique, within Scraptiidae, pectinate antennae with males of the monotypic North American genus Pectotoma Hatch, placed in the scraptiid tribe Allopodini by Young (1976) and with males of the monotypic Tasmanian Xylophilostenus Lea, 1917, placed in Scraptiidae by Lawrence (1987). The antennae of Samlandotoma consists of simple scape and pedicel (non-pectinate, conical in form), pectinate antennomeres 3 – 9, antennomeres 10 elongately triangular and antennomere 11 simple, spindle-shaped, whereas the antennomeres 3 – 10 of Pectotoma male and 4 – 10 of Xylophilostenus male are pectinate. Additionally, base of each projection on antennomeres 3 – 9 is located at distal margin of each antennomere in Samlandotoma in contrast to location of projection near the base of antennomere in Pectotoma.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE17ED221FF2DF903FB71FD4F.taxon	description	(Figs 7 – 8)	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE17ED221FF2DF903FB71FD4F.taxon	materials_examined	Material examined. Holotype, No. 1799 - 6 [CCHH], adult, male. Complete beetle inclusion is preserved in a polished piece of transparent amber with an orange shade without any further fixation. The amber piece is irregularly triangular, with maximum length about 12 mm and maximum width about 2 mm. The animal syninclusions are represented by one mite (Acari) about 0.3 mm in length. Type strata. Baltic amber, most probable from the Eocene amber-bearing layers of Blaue Erde (Blue Earth) within the Prussian Formation; estimated age: middle-late Eocene (Bukejs et al. 2019). Type locality. Yantarny village (formerly Palmnicken), Sambian (Samland) Peninsula, Kaliningrad Region, western Russia. Taxonomic assignment. This taxon in Baltic amber should be placed into Scraptiini (Scraptiidae: Scraptiinae) based on a combination of the following morphological characters (mentioned in original description or clearly visible in photographs): (1) body form elongate-oval with base of pronotum slightly narrower than anterior elytral margin; (2) eyes emarginate anteriorly at antennal base; (3) head constricted posteriorly; (4) terminal maxillary palpomere elongate oval with pointed apex, flattened, nearly as long as penultimate palpomere; (5) antennae 11 - segmented, not clubbed, filiform; (6) antennal insertions exposed from above; (7) pronotum laterally margined; (8) procoxae subcontiguous, projecting well below prosternum; (9) elytra irregularly punctate, without transverse strigae; (10) tarsi 5 - 5 - 4 with all penultimate tarsomeres lobed; (11) tibial spurs paired, equal in length, well-developed, not serrate; (12) crenulation on tibiae and tarsi absent; and (13) abdomen with five ventrites and ventrite 1 not much longer than ventrite 2. The ventral side of forebody is not clearly visible (e. g. labial palpomeres, prosternal process, and protrochantins), basal pits of pronotum seems to be inconspicuous (barely seen). Regardless of that and on the basis of the characteristics mentioned above, we propose that Eocene Quasianisoxya Alekseev should be removed from the family Melandryidae and transferred to Scraptiini within Scraptiidae.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE179D221FF2DFC3CFDA3FB93.taxon	description	1. † C. crucifera sp. nov. Kachin amber.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE179D221FF2DFB4CFD34F8CF.taxon	description	2. † A. emarginata Abdullah, 1964: 341. Baltic amber. † Genus Palaeoscraptia Abdullah, 1964: 339. 3. † P. elongata Abdullah, 1964: 340. Baltic amber. Genus Scraptia Latreille, 1806 4. † S. inclusa Ermisch, 1941: 184. Baltic amber. 5. † S. longelytrata Ermisch, 1943: 67. Baltic amber. 6. † S. ovata Guérin-Méneville, 1838: 170. Sicilian amber. 7. † S. pseudofuscula Ermisch, 1941: 183. Baltic amber. † Genus Quasianisoxya Alekseev, 2015: 171. 8. † Q. curonensis Alekseev, 2015: 172. Baltic amber. † Genus Samlandotoma Alekseev, 2019: 285 9. † S. seidlitzi Alekseev, 2019: 286. Baltic amber.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE179D222FF2DF89CFF14FDD3.taxon	description	10. † A. antica Guérin-Méneville, 1838: 170. Sicilian amber. Subgenus Anaspis Geoffroy, 1762 11. † A. (s. str.) longispina Ermisch, 1941: 182. Baltic amber. 12. † A. (s. str.) parallela Ermisch, 1941: 182. Baltic amber. Subgenus Silaria Mulsant, 1856 13. † A. (S.) parva Abdullah, 1964: 341. Baltic amber. Subgenus Spanisa Emery, 1876 14. † A. (S.) horaki Perkovsky et Odnosum, 2009: 1092. Baltic and Rovno amber. 15. † A. (S.) solodovnikovi Telnov et Perkovsky, 2025: 429. Rovno amber. Genus Cyrtanaspis Emery, 1876 16. † C. kerneggeri Perkovsky et Odnosum in Perkovsky, Odnosum, Nazarenko et Vasilenko, 2022: 209. Baltic amber.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
03C4A00CE17AD222FF2DFD8CFBEFFD07.taxon	description	17. † S. brachycornis Medvedev, 1969: 122. Baisa (Zaza formation). 18. † S. minima Ponomarenko in Ponomarenko et Ryvkin, 1990: 76. Turga suite.	en	Alekseev, Vitalii, Háva, Jiří, Bukejs, Andris (2025): First adult of false flower beetle (Coleoptera: Scraptiidae) from mid-Cretaceous amber of northern Myanmar, with taxonomical notes and a checklist of fossil representatives of the family. Zootaxa 5723 (4): 533-549, DOI: 10.11646/zootaxa.5723.4.4, URL: https://doi.org/10.11646/zootaxa.5723.4.4
