Pulicarioidea annamica (Gagnep.) Bunwong, Chantar. & S.C.Keeley, 2014

Bunwong, Sukhonthip, Chantaranothai, Pranom & Keeley, Sterling C., 2014, Revisions and key to the Vernonieae (Compositae) of Thailand, PhytoKeys 37, pp. 25-101 : 87-88

Pulicarioidea annamica (Gagnep.) Bunwong, Chantar. & S.C.Keeley comb. nov.

Pulicaria annamica Gagnep., Bull. Soc. Bot France. 68: 121. 1921.

Vernonia pulicarioides Gagnep., Fl. Indo-Chine 3: 482. 1924.

Type.

Vietnam, Annam, Da Lat plateau, Langbian mountain, A. Chevalier 30672 (P!, lectotype designated here). Fig. 10C .

Description.

Perennial herbs, 20-40 cm tall. Stems erect, conspicuously ribbed, villose. Leaves 5-10 by 1.5-2 cm, oblong or oblanceolate, margin subentire, apex acute or truncate, base cuneate, subcoriaceous; upper surface scabrous without glands; lower surface scabrous with whip-shaped hairs and capitate glands; lateral veins 4-8-paired; petioles sessile. Capitulescences terminal, solitary or few. Capitula hemispherical, 10-15 mm long, pedunculate. Receptacle glabrous. Involucres hemispherical, in 5-6 series, 10-15 mm long, 10-15 mm in diam., phyllaries imbricate, green with purple apically, margin piliferous, outer surface sericeous without glands; the outer and the middle ones lanceolate, apex acuminate; the inner ones lanceolate, apex acuminate or aristate. Florets ca. 70; corollas narrowly funnelform, purple, pubescent with hairs and glands; corolla tubes 6-7 mm long; corolla lobes 2-2.5 mm long. Anthers 3.5-4 mm long, apical appendage acute, base acute. Styles purple, 6-6.5 mm long, branches 2-2.5 mm long. Achenes subterete or oblong, 2-2.5 mm long, 3-5 ribbed, pubescent with dense twin hairs without glands, carpopodium present. Pappus in 2 series of bristles, persistent, the outer ones shorter than the inner ones, the inner ones 8-9 mm long. Pollen lophate, without micropuncta.

Distribution.

Thailand: Mae Hong Son, Chiang Mai. Myanmar, Laos, Vietnam.

Specimens examined.

Thailand, Chiang Mai, Doi Sutep Pui national park, 18°48.39'N, 98°54.90'E, 20 Oct 1999, P. Suksathan 1932 (QBG); Laos, M. Poilane 2038 (P); Laos, 4 Feb 1932, M. Poilane 20073 (P); Vietnam, Annam, Da Lat plateau, Langbian mountain, 10 Feb 1914, A. Chevalier 30672 (P); Langbian mountain, 15 Feb 1914, A. Chevalier 30847 (P); Langbian mountain, Eberhardt 1747 (P); Langbian mountain, 27 Oct 1920, E. Evrard 406 (E, P); Langbian mountain, 19 Jan 1924, E. Evrard 1057 (P); Langbian mountain, E. Evrard 1446 (P); Langbian mountain, 20 Feb 1952, Schmid 1237 (P).

Diagnostic characters.

This species is distinguished by having hemispherical capitula, sericeous phyllaries without glands, 3-5 ribed achenes, whitish flattened pappus and lophate pollen with small lacuna.

Ecology.

Evergreen and pine-oak forest, alt. 780-1600 m; flowering October to February.

Vernacular name.

Pha Ya Muang Doi (พญาม่วงดอย).

Figures

Figure 10. Morphology of Vernonieae in Thailand 6. A Okia birmanica B Okia pseudobirmanica C Pulicarioidea annamica D-E Strobocalyx arborea F Strobocalyx solanifolia G-H Struchium sparganophorum I Tarlmounia elliptica.