Paragrallomyia pseudoalbibasis,

Ferro, Gustavo Borges & Marshall, Stephen A., 2020, A redefinition of Paragrallomyia Hendel (Diptera: Micropezidae, Taeniapterinae) and a revision of the P. albibasis complex, Zootaxa 4822 (1), pp. 39-70: 58-60

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Paragrallomyia pseudoalbibasis

new species

Paragrallomyia pseudoalbibasis  new species

( Figs. 35–38View FIGURES 35–38)

Description: Body length 7 mm; wing 5 mm. Head and abdomen dark brown, thorax black. Palpus dark brown except for white outer face. Clypeus dark brown with sparse pale microsetulae. First flagellomere dark brown. Fronto-orbital plate dark brown. Row of katepisternal setae brownish. Fore tibia dark brown. Fore tarsus mostly white, tarsomere one brown on basal half only. Hind femur mostly brown, white on basal quarter and with a small angled preapical white ring. Tergites dark brown. Oviscape dark brown.

Head: First flagellomere covered by pale microsetulae. Frontal vitta with a darkened spot anterior to ocellar plate. Frontal vitta flat, not swollen anterior to ocellar plate. Epicephala merging immediately posterior to ocellar plate ( Fig. 35View FIGURES 35–38). Epicephalon somewhat swollen above frontal vitta level.

Thorax: Mesonotum with lines of thin brownish microsetulae. Anterior half of postpronotal lobe bare. Femur covered with sparse black microsetulae. Mid and hind tibia flattened, sulcus on outer face strongly pronounced and curved ( Fig. 37View FIGURES 35–38). Female cervical sclerite with a white oval swollen on posterior end. Cell r 4+5 open at the tip of the wing.

Female abdomen: Single spermathecal duct with an apical rounded swelling. Single spermatheca cylindrical, with sparse small finger-like swellings on apical half ( Fig. 36View FIGURES 35–38). Paired spermathecal duct swollen apically before splitting into stems. Paired spermathecal duct stems tapered on basal third, followed by a rounded swelling and a tapered apical third with small finger-like processes. Paired spermathecae oval, and with an apical nipple-like swelling. Male abdomen: Copulatory fork with small inner basal lobes, less than half as long as the arms ( Fig. 38View FIGURES 35–38). Ejaculatory apodeme bigger than epandrium and with a thin blade, length subequal to sperm pump. Phallapodeme short, not extending to anterior limit of hypandrium. Distiphallus long, longer than epandrium and with a tapered phallic bulb. Distal distiphallus short and thin, 1/4 of length and almost half of width of basal distiphallus.

Type material: Holotype: COSTA RICA. Puntarenas, Corcovado Natl. Pk., san Pedrillo , 5–50 m, 8º 37’ 15 N 83º44’6 E, 13.VIII.2001, S. A. Marshall (1 ♀, DEBU00168174View Materials, MNCR)GoogleMaps  . Paratypes: BELIZE. Cayo Dist., San Ignacio, Maya Mt. Lodge, 17º 9’ N 89º4’W, dung trap, 7–8.I.1991, S. A. Marshall (1 ♂ DEBU00151711View Materials)GoogleMaps  ; COSTA RICA. Alajuela, Volcan Tenorio, N slope nr. Bijagua Biol. Sta. , 700 m, 16–20. VI.2000, N. E. Woodley (1 ♀, DEBU00118457View Materials)  ; Alajuela, Volcan Tenorio, N slope nr. Bijagua Biol. Sta. , 700 m, rain forest, 19. VI.2000, M. Buck (1 ♀, DEBU00118536View Materials)  ; Alajuela , Volcan Tenorio, N slope, trail to laguna, 800–900 m, rain forest, 16–20. VI.2000, S. A. Marshall (2 ♀, DEBU00118536View Materials, DEBU00118537View Materials)  ; Guana, Estacion Pitilla , 9 km Santa Cecili, 700 m, ENE 1995, C. Moraga, L N 329950 380450 #4354 (1 ♂, INBIO CRI002131321, MNCR)  ; Heredia, Estac. Biol. La Selva , 18–21.IV.2000, E. M. & C. F. Fisher coll. (1 ♀, Sequence MYCRO426-18, DEBU01088958View Materials)  ; Heredia, F. La Selva , 3 km S Pto. Viejo, 10º26’N 84º01’W, 16.IV.1984, H. A. Hespenheide (1 ♀, DEBU)GoogleMaps  ; Puntarenas, Osa Peninsula 2.5 km S Rincon , ~ 50 m, 8º 42’ 1 N 83º30’50 W, 10–11.VIII.2001, S. A. Marshall (3 ♀, DEBU00168049View Materials, DEBU00168050View Materials, DEBU00168055View Materials)GoogleMaps  ; Puntarenas, Osa Peninsula 2.5 km S Rincon , ~ 50 m, 8º 42’ 1 N 83º30’50 W, trail nr. station, human dung, 10–11.VIII.2001, S. A. Marshall (4 ♀, 1 ♂, DEBU00168234View Materials, DEBU00168237View Materials, DEBU00168245View Materials, DEBU00168246View Materials, DEBU00168252View Materials)GoogleMaps  ; Puntarenas, R. F. Golfo Dulce 24 km W, Piedra Blanca , 200 m, II.92, Tp 2, P. Hanson (1 ♀, DEBU)  ; San José, San Carlos, Riosparaiso Res. , Pecari Stn. 16 km NNE Quepos, 400 m, 9º 33’ 53N 84º7’32 W, 12–15.IV.2006, S. A. Marshall (1 ♀, 1 ♂, DEBU00256639View Materials, DEBU00258115View Materials)GoogleMaps  ; San José, San Carlos, Riosparaiso Res. , along trail from Rio Negro to Quepos , 9º 33’ 43N 84º7’38 W, 23.II.2006, S. M. Paiero (1 ♀, Sequence 404-18, DEBU00256599View Materials)GoogleMaps  ; San José, San Carlos, Riosparaiso Res. , upper trail to Rio Negro , 1º forest, 26.II.2006, S. M. Paiero (2 ♂, Sequence MYCRO405-18, DEBU00256632View Materials)  ; San José, San Carlos, Riosparaiso Res. , Pecari Stn. 16 km NNE Quepos, 400 m, 9º 33’ 53N 84º7’32 W, 15.II.2003, S. A. Marshall (1 ♂, Sequence MYCRO427-18, DEBU00205712View Materials)GoogleMaps  ; San José, San Carlos, Riosparaiso Res. , Pecari Stn. 16 km NNE Quepos, 400 m, 9º 33’ 53N 84º7’32 W, 15.II.2006, S. A. Marshall (1 ♂, DEBU00206731View Materials)GoogleMaps  .

Distribution: Belize, Costa Rica.

Etymology: The specific name reflects similarity with P. albibasis  .

Comments: A very s imilar species to P. albibasis  , however P. pseudoalbibasis  can easily be recognized by the flattened mid and hind tibiae with the sulcus on the outer face strongly pronounced and curved. The long and thin ejaculatory apodeme blade also differs from the reduced blade of P. albibasis  .


Museo Nacional de Costa Rica


Timescale Adventures Research and Interpretive Center


Mykotektet, National Veterinary Institute


Ontario Insect Collection, University of Guelph


Departamento de Geologia, Universidad de Chile