Gondwanodromia elongata, Sinclair & Saigusa, 2022

Sinclair, Bradley J. & Saigusa, Toyohei, 2022, A New Aquatic Associated Genus of Trichopezinae from the Southern Hemisphere (Diptera: Empidoidea: Brachystomatidae), Records of the Australian Museum (Rec. Aust. Mus.) 74 (3), pp. 75-98 : 81-82

Gondwanodromia elongata sp. nov.

urn:lsid:zoobank.org:act:E7A95AA3-FE78-4B7B-BEB3-90EB06D1FB49

Figs 8, 9 , 17 , 51

Holotype ♂, labelled: “ NEW ZEALAND: TO/ Pureora For. Pk./ 2.ii.1995, Kakaho / str., Link Rd. [38°34'S 175°43'E]/ B.J. Sinclair ”; “ HOLOTYPE / Gondwanodromia / elongata/ Sinclair & Saigusa [red label]” ( NZAC) . Paratypes: NEW ZEALAND. WN: 1 ♀, Cloustonville , Akatarawa R. [40°58'S 175°07'E], YPT [yellow pan trap], 3–4.ii.1995, B.J. Sinclair ( CNC) .

Diagnosis. The species is readily distinguished by the scutum with shiny vittae between setal rows, lateral scutal setae spine-like, and male terminalia with epandrium not prolonged beyond base of surstylus in lateral view.

Description. Wing length 3.1–3.7 mm. Male. Head. Eye bare; widely separated on frons and face, wider than antennal sockets; face with bluish pruinescence. Ocellar setae long and divergent, inserted anteromedial to lateral ocelli; 2 pairs of shorter posterior setae; 2 vertical setae, outer vertical seta arched over eye, longer than ocellar setae. Antennae inserted above middle of head; scape slightly longer than globular pedicel; postpedicel with broad base, gradually tapered; stylus slightly shorter than postpedicel, small basal segment and short apical peg-like sensillum ( Fig. 8 ). Palpus long and slender, two-thirds height of head, bearing scattered dark setae; labrum greatly lengthened, longer than height of head, slightly incurved; labium shorter than labrum.

Thorax. Pleura, coxae with pale silvery-grey pruinescence. Scutum brown with shiny vittae between rows of acrostichals and dorsocentrals. Antepronotum with several pairs of stout setae.Acrostichals uniserial, slightly longer behind, extending to prescutellar depression; 6 uniserial dorsocentral setae, anterior seta strong, decreasing posteriorly; lateral setae spine-like: 2 postpronotal setae; 1 presutural supra-alar seta; 2 notopleural setae, lower slender and short; 3 short postsutural supra-alar setae; 1 postalar seta; 3 pairs of scutellar setae and outer pairs shorter than median pair. Laterotergite with patch of pale setae. Wing. Broad, infuscate; basal costal seta short and stout; costa with erect dorsal setae beyond R 1; R 4 nearly straight; halter pale yellowish-brown. Legs. Coxae with pale golden setae; knees yellowish-brown. Fore femur with anteroventral and posteroventral row of short, slender setae, shorter than width of femur; 3 stout dorsal setae on basal half, shorter than width of femur. Mid femur with anteroventral and posteroventral row of stout, black setae, posteroventral setae mostly longer than width of femur; apical half with rows of stout anterior and dorsal setae. Mid tibia lacking erect ventral setae. Hind femur arched with long, erect stout dorsal and anteroventral setae. Hind tibia with stout ventral and dorsal setae, shorter than width of tibia.

Abdomen. Brown, sparsely setose with setae mostly confined to the posterior margins. Segment 7 narrow posteriorly; sternite 7 with short, triangular posterolateral extension. Terminalia ( Fig. 17 ). Lying dorsally over segments 7 and 8. Hypandrium round apically, encircling phallus (likely postgonites fused with hypandrium at apex). Phallus rounded apically with paired posterior projections; with broad membranous cap; ejaculatory apodeme broad and tapered ventrally. Epandrium oval-shaped, without narrowed and tapered posterodorsal corner. Surstylus subtriangular, apex rounded with short setae on posterior margin. Cercus thinly sclerotized, clothed in microtrichia; several long setae on anterior and posterior faces.

Female. Similar to male except as follows: Palpus subequal to height of eye. Abdominal sternites 3–4 with 2 long, lateral marginal setae; sternite 7 with two pairs of closely approximated ventral setae, length two-thirds width of segment. Terminalia [undissected] ( Fig. 9 ). Sclerites of segment 8 sclerotized and shiny; sternite 8 prolonged beyond tergite 8. Cercus elongate, fused with syntergite 9+10, shiny, well sclerotized, projecting horizontally, longer than segment 8; with fine, short dorsal setae.

Etymology. This species is named after elongate female cerci.

Distribution. This species is currently known from the North Island of New Zealand ( Fig. 51 ).

Figures

Figures 5–9. Australian and New Zealand species of Gondwanodromia, lateral view: (5) G. bulbosa sp. nov., male habitus, holotype, scale bar = 0.75 mm; (6) G. bulbosa sp. nov., male terminalia, scale bar = 0.25 mm; (7) G. colomatta sp. nov., male habitus, holotype, scale bar = 0.75 mm; (8) G. elongata sp. nov., male habitus, holotype (prior to abdomen dissection), scale bar = 0.5 mm; (9) G. elongata sp. nov., female habitus,scale bar=0.75 mm.Abbreviations:asur – anteriorsurstylus;cerc –cercus;epand –epandrium;psur – posteriorsurstylus;st– sternite.

Figures 17–18. Male terminalia of New Zealand andAustralian species of Gondwanodromia, lateral view: (17) G. elongata sp. nov.; (18) G. lutea sp. nov. Scale bar = 0.1 mm.Abbreviations: cerc – cercus; epand – epandrium; hypd – hypandrium; ph – phallus; sur – surstylus.

Figures 50–52. Distribution of Gondwanodromia: (50) Australian species; (51) New Zealand species; (52) South American species.