Trichopezinae, Vaillant, 1981

Sinclair, Bradley J. & Saigusa, Toyohei, 2022, A New Aquatic Associated Genus of Trichopezinae from the Southern Hemisphere (Diptera: Empidoidea: Brachystomatidae), Records of the Australian Museum (Rec. Aust. Mus.) 74 (3), pp. 75-98 : 78

Key to genera of Southern Hemisphere Trichopezinae

Modified from Sinclair (2011)

1 Wing with only two longitudinal veins fully developed (R 4+5 and M 4); very long arista-like stylus, longer than thorax; body size minute, 1.1–1.4 mm ( New Zealand, Chile) .................................................... Gondwanamyia Sinclair, Cumming, Brooks, Plant, & Saigusa

—— Wing with more than two longitudinal veins fully developed; arista-like stylus if present, shorter than thorax; body size larger, greater than 3 mm ...................................................................................................................... 2

2 Wing narrow, without anal lobe ( New Zealand) .......................................... Sematopoda Collin

—— Wing broad, anal lobe developed .............................................................................................. 3

3 Laterotergite with group of setae ............................................................................................... 4

—— Laterotergite bare ...................................................................................................................... 9

4 Vein R 4+5 unbranched; postpedicel with arista-like stylus arising distinctly before apex ( Chile) ................................................................... Hyperperacera Collin

—— Vein R 4+5 branched; postpedicel with arista-like stylus or stylus arising at apex or stylus absent .................................................................................................. 5

5 Vein R 1 with setae above ( Australia, Chile, New Zealand) .................... Heterophlebus Philippi

—— Vein R 1 bare, without setae ........................................................................................................ 6

6 Fork of vein R 4+5 narrow and V-shaped; males holoptic ( Ecuador) ..................................................................................... Apalocnemis Philippi [in part]

—— Fork of vein R 4+5 broad and bell-shaped ( Fig. 20 ); males dichoptic ............................................. 7

7 Palpus drooping, with long thin setae ( Figs 3 , 30 ) ( Australia, New Zealand, South America) ................................................................. Gondwanodromia gen. nov.

—— Palpus projected obliquely, sometimes with short, apical spinelike setae .................................................................................................................................... 8

8 Postpedicel with small swollen base, abruptly tapered, prolonged apically, longer than apical stylus ( New Zealand) .............. Adipsomyia Yang, Zhang, & Zhang

—— Postpedicel ovate with slender extension bearing elongate and narrow stylus ( Argentina, Chile) ..................... undescribed genus ( Saigusa, 2006, unpubl. data)

9 R 4+5 branched; costa circumambient; males holoptic ( Australia, New Zealand, South America) .................................................... Apalocnemis Philippi [in part]

—— R 4+5 unbranched; costa greatly weakened after R 4+5; males dichoptic ( New Zealand) ............................................ undescribed genus C (see Sinclair, 2011)

Figures

Figures 19–23. Gondwanodromia mikae sp. nov.: (19) labellum, lateral view, scale bar = 0.05 mm; (20) wing, scale bar = 1.0 mm; (21) male antenna, lateral view, scale bar = 0.1 mm; (22) tarsomere 5, dorsal view, scale bar = 0.05 mm; (23) tarsomere 5, lateral view, scale bar = 0.05 mm. Abbreviations: pped – postpedicel; psdtrch – pseudotrachea.

Figures 1–4. Photographs of living Gondwanodromia and habitats:(1)Arroyo Torrontegui,Argentina; (2)Arroyo Torrontegui,Argentina, with collector Mika Sugimoto; (3) female, Altos del Lircay, Chile (photograph © Steve Marshall); (4) Dead Horse Gap trail, Kosciusko National Park, Australia; November 1994.

Figures 30–33. Gondwanodromia mikae sp. nov.: (30) male head, lateral view; (31) epandrium, dorsal view, right cercus removed; (32) terminalia, lateral view; (33) hypandrium and phallus, lateral view. Scale bars: Figs 31–33 = 0.05 mm; Fig. 30 = 0.1 mm.Abbreviations: cerc – cercus; ej apod – ejaculatory apodeme; epand – epandrium; goncx apod – gonocoxal apodeme; hypd – hypandrium; lc – lacinia; ph – phallus; plp – palpus; pped – postpedicel; sbepand scl – subepandrial sclerite; st – sternite; sur – surstylus.