Gondwanodromia tonnoiri, Sinclair & Saigusa, 2022

Sinclair, Bradley J. & Saigusa, Toyohei, 2022, A New Aquatic Associated Genus of Trichopezinae from the Southern Hemisphere (Diptera: Empidoidea: Brachystomatidae), Records of the Australian Museum (Rec. Aust. Mus.) 74 (3), pp. 75-98 : 94

Gondwanodromia tonnoiri sp. nov.

urn:lsid:zoobank.org:act:BD29C724-927A-424A-AF77-07C0E8884D6F

Figs 48 , 50

Holotype ♂, labelled: “TAS: Double-Barrel/Ck. [42°11'S 145°57'E], Lyell Hwy; 27-/ - i-1989; rainfor./ D.Bickel; yellowpan”; “ HOLOTYPE / Gondwanodromia / tonnoiri/ Sinclair & Saigusa” ( AMS K.594169) . Paratypes: Australia. Tasmania: 1 ♀, Hartz Mts [43°14'S 146°45'E], 10.xii.1922, A. Tonnoir ( ANIC) ; 1 ♀, Cradle Mtn NP, Dove Lk [41°39'S 145°57'E], 920 m, dung pans in button grass, 16–17.xii.2003, S.A. Marshall, debu00247175 ( DEBU) .

Diagnosis. The species is readily distinguished by bare eyes, mid tibia not swollen apically and with pair of slightly stronger ventroapical setae and epandrium shallowly notched laterally.

Description. Wing length 3–3.7 mm. Male. Head. Eye bare, without ommatrichia. Frons brown. Face with bluish pruinescence; broad, subequal to width of antennal sockets. Ocellar setae long and divergent, inserted anteromedial to lateral ocelli; one shorter posterior seta; 2–3 vertical setae, outer seta arched over margin of eye, shorter than ocellar seta; occipital setae shorter and thinner than vertical setae. Antenna with scape subequal to length of pedicel; postpedicel more than 2× height of eye. Proboscis longer than head height; palpus pale brown, half-length of proboscis.

Thorax. Dark brown, with thin blue pruinescence laterally on scutum, prescutellum, scutellum and pleura. Antepronotum with several pairs of strong setae, longer laterally. Acrostichal setae uniserial with setae alternating in direction, more than half length of dorsocentral setae; 7–12 uniserial dorsocentral setae, anterior seta set off laterally from row; 1 long postpronotal seta and 1–2 shorter anterior setae; 1 presutural supra-alar seta; 3 notopleural setae, middle seta stronger and twice length of other setae; 2–3 postsutural supra-alar setae; 1 postalar seta; 2 pairs of scutellar setae, outer pair more than half length of inner pair. Laterotergite with about 5 setae. Wing. Infuscate; basal costal seta long and slender. Halter pale brown. Legs. Coxae yellowish brown, apical setae brown; remaining leg segments increasingly darker apically. Femora short and slightly swollen; fore femur with posteroventral row of slender setae; mid femur with row of short anteroventral and row of longer posteroventral setae; hind femur with erect dorsal setae near base. Fore tibia without apical spine-like setae. Mid tibia slender, not expanded subapically; with anteroventral row and posteroventral row of short setae; with pair of slightly thickened apicoventral setae. Hind tibia with stout dorsal and ventral setae, shorter than width of tibia. Mid tarsomere 1 with row of strong posteroventral setae; 1 erect posteroventral seta at basal fifth.

Abdomen. Tergite 6 with long dorsal setae on posterior half, nearly as long as tergite, setae longer than on other tergites. Sternite 7 prolonged posterolaterally into narrow lobe overlapping anterior margin of sternite 8. Tergite 8 trapezoidal, half-length of sternite 8; posterior margin wider than anterior margin, bearing numerous setae, denser on lateral margin; sternite 8 trapezoidal, with setae longer than anterior sternites with pair of posteroapical sclerites. Terminalia ( Fig. 48 ). Cercus thinly sclerotized, thumb-like and tapered apically, bearing pair of long, spine-like setae directed posteriorly. Epandrium shallowly notched laterally, anterior and posterior halves equally narrow; without dorsal bridge. Surstylus paired; anterior surstylus smoothly sclerotized, without microtrichia, forked, with lower branch triangular bearing subapical seta; upper lobe rounded apically with subapical seta; posterior surstylus clothed in microtrichia, produced apically into thin, flap-like lobe with several marginal setae mounted on small projections. Hypandrium tubular, arched and recurved apically; narrow gonocoxal apodemes medially, directed ventrally. Phallus narrow, arched within hypandrium; apex tapered, projecting anteriorly beyond hypandrium; ejaculatory apodeme fused to base of phallus.

Female. Similar to male except as follows: postpedicel less than 1.5× eye height; midleg unmodified. Syntergite 9+10 (acanthophorites) with 2 rows of strong, erect setae.

Etymology. The species is named after the collector of the first specimen of this new genus, André Léon Tonnoir.

Distribution. This species is currently known from three localities in Tasmania ( Fig. 50 ).

Figures

Figures 48–49. Male terminalia ofAustralian and New Zealand species of Gondwanodromia, lateral view: (48) G. tonnoiri sp. nov.; (49) G. wardi sp. nov. Scale bar = 0.1 mm. Abbreviations: a sur – anterior surstylus; cerc –cercus; epand – epandrium; hypd – hypandrium; p sur – posterior surstylus; ph – phallus; sur – surstylus.

Figures 50–52. Distribution of Gondwanodromia: (50) Australian species; (51) New Zealand species; (52) South American species.