Pyrops lathburii ( Kirby, 1818 ) Pyrops, Spinola, 1839

Constant, Jérôme & Pham, Hong Thai, 2022, The Lanternfly genus Pyrops in Vietnam: A new species from Central Vietnam, taxonomic changes, checklist, identification key (Hemiptera: Fulgoromorpha: Fulgoridae), European Journal of Taxonomy 813 (1), pp. 123-154 : 131-139

publication ID

https://doi.org/10.5852/ejt.2022.813.1741

publication LSID

lsid:zoobank.org:pub:4957CF5B-38A4-45F6-8313-AA80C5091E50

DOI

https://doi.org/10.5281/zenodo.6470274

persistent identifier

https://treatment.plazi.org/id/022F87A6-297A-FF86-FDDE-FAAEFDDE6710

treatment provided by

Felipe

scientific name

Pyrops lathburii ( Kirby, 1818 ) Pyrops
status

 

Pyrops lathburii ( Kirby, 1818)

Figs 6–8 View Fig View Fig View Fig , 14 View Fig

Fulgora lathburii Kirby, 1818: 450 (described, compared with Pyrops rogersi ( Distant, 1906)) [type in BMNH].

Fulgora astarte Distant, 1914: 409 (described) [type in BMNH]. syn. nov.

Fulgora lathburii – Guérin-Méneville 1829: pl. 58 fig. 2 (illustrated). — Latreille 1837: pl. 68 fig. 2 (illustrated). — Spinola 1839: 234 (as “ Fulgora lathburyi ”, treated as a dark form of Pyrops candelaria Linné, 1758 (error!)). — Westwood 1839: 139 (described). — Comte 1840: pl. 68 (listed, illustrated). — Guérin-Méneville 1844: 357 (notes), pl. 58 fig. 2 (illustrated). — White 1844: 425 (recorded from Hong-Kong). — Burmeister 1845: unnumbered page, under genus Fulgora , subgenus Pyrops (as a variety of Fulgora (Pyrops) candelaria (error!)). — Guérin-Méneville 1845: xcvi (as “ Fulgora lothburii ”, close to Pyrops cyanirostris Guérin-Méneville, 1845 ). — Stål 1870: 741 (as “ Fulgora lathburi ”, compared with Pyrops philippinus Stål, 1870 ). — Butler 1874: 97 (listed with species with orange posterior hind wings). — Gadeau de Kerville 1881: 43 (listed as “ Fulgora lalhburi ” in “Série I”, species with orange wings, in a census of light-emitting insects). — Atkinson 1885: 129 (described). — Distant 1906: 186 (keyed, nomenclature, described, as “ Fulgora lathburi ”). — Schmidt 1911: 161 (as “ Fulgora lathburi ”, compared with Pyrops peguensis Schmidt, 1911 ). — Schmidt 1919: 126 (as “ Fulgora lathburi ”, listed from Canton (= Guangzhou)). — Baker 1925: 349 (keyed inside the candelaria species group), 351 (notes), pl. 3 fig. 2 (dorsal habitus), pl. 4 fig. 6 (lateral habitus). — Lallemand 1963: 86 (transferred to the pyrorhynchus species group, keyed, described), pl. xi figs 1–3 (male terminalia).

Flata lathburii – Germar 1830: 46 (described).

Hotinus lathburii – Amyot & Audinet-Serville 1843: 491 (transferred to Hotinus Amyot & Audinet- Serville). — Walker 1851: 266 (listed, recorded from Silhet (= Sylhet, Bangladesh)). — Dohrn 1859: 57 (listed in catalogue as “ Hotinus lathburi ”). — Girard 1885: 859 (as “ H. lathburi ”, described).

Pyrops lathburii – Schaum 1850: 64 (transferred to Pyrops Spinola, 1839 ). — Nagai & Porion 1996: 25 (catalogued, recorded from Assam and Thailand), figs 194, 196, 198 (dorsal habitus). — Liang 1998: 43 (listed). — Pham 2011: 318 (recorded from Vietnam). — Constant 2015: 9 (note on intraspecific colour variation) — Constant et al. 2016: 15 (in list of species to look for in Cambodia). — Constant & Pham 2017: 18 (host plant). — Wang et al. 2018: 297 (keyed), 301 (nomenclature, measurements, description of male genitalia, diagnostic characters; records in China), figs 18–29 (colour variation, details of head, male terminalia). — Constant 2021: 14 (compared with Pyrops philippinus ( Stål, 1870)) . — Jiaranaisakul & Constant 2021: 18 (in list of species to look for in Khao Krachom Mountain, Thailand).

Laternaria lathburii – Metcalf 1947: 197 (transferred to Laternaria Linné, 1964 , catalogued).

Fulgora astarte – Distant 1918: 198 (listed from Indochina), 199 (allied to Pyrops vitalisius Distant, 1918 ). — Lallemand 1963: 85 (placed in the pyrorhynchus species group, keyed, described).

Laternaria astarte – Metcalf 1947: 186 (transferred to Laternaria Linné, 1964 , catalogued).

Pyrops astarte – Nagai & Porion 1996: 25 (transferred to Pyrops , catalogued, recorded from Vietnam and Thailand), figs 195, 197, 199 (dorsal habitus). — Pham & Ta 2004: 58 (in key to Fulgoridae of Vietnam). — Liang 1998: 42 (listed). — Pham 2011: 318 (record from Vietnam). — Constant 2021: 14 (compared with Pyrops philippinus ( Stål, 1870)) .

Note: Wang et al. (2018) erroneously mentioned that the species was attributed to the pyrorhynchus group in Constant & Pham (2017).

Material examined

Lectotype of Fulgora lathburii Kirby, 1818 ( Fig. 6 View Fig ), here designated to provide a stable reference for the taxonomy of the group

CHINA • ♀; [China]; [D. Lathbury leg.]; “63 / 41”, “Lathburii K in L. Tr.”, “Kirby.”; BMNH .

Note: The specimen was collected in China by D. Lathbury according to Kirby (1818). The code “63 / 41” refers to an entry in the Accession Register in BMNH starting on page 803 of the register and bearing a comment “These insects are the types of Mr Kirby’s Century of Insects [...]” (V. Lemaître pers. com., Jul. 2021).

Holotype of Fulgora astarte Distant, 1914 ( Fig. 7 View Fig )

VIETNAM • ♀; Indochina , [Lào Cai]; R. Vitalis leg.; “Indo-China (R. Vitalis)”, “ Fulgora astarte Type Dist.”, “Type H.T.”; BMNH .

Note: The specimen was collected in Laokay (= Lào Cai) in North Vietnam according to Distant (1914).

Additional material

LAOS • 1 ♀; Vientiane; [17°58′ N, 102°36′ E]; I.G. 23.993 ; RBINS GoogleMaps .

THAILAND • 1 ♀; Phrae Province; May 2004; ex coll. Neef de Sainval; I.G. 30.731 ; RBINS 1 ♂, 1 ♀; Chiang Rai Province; Wiang Pa Pao ; [19°20′54″ N, 99°30′24″ E]; 10–15 Apr. 2008; ex coll. A. Chaminade; I.G. 31.467 ; RBINS GoogleMaps .

VIETNAM • 3 ♂♂, 4 ♀♀; Cao Bang Province, Phia Den ; [22°34′26″ N, 105°52′29″ E]; 8 Aug. 2010; J. Constant and P. Limbourg leg.; day time; I.G. 31.668 ; RBINS GoogleMaps 5 ♂♂, 8 ♀♀; Tay Yen Tu Nature Reserve ; 21°11′10″ N, 106°43′25″ E; 7–11 Jul. 2013; J. Constant and J. Bresseel leg.; I.G. 32.454 ; RBINS GoogleMaps 3 ♂♂; Bac Giang Province, Tay Yen Tu NR , Tram Dong Thong ; 3 Jul. 2010; H.T. Pham leg.; VNMN 1 ♀; [Hai Phong Province], Cat Ba National Park ; 20°48′00″ N, 107°00′20″ E; 12–16 Jul. 2013; J. Constant and J. Bresseel leg.; day time; I.G. 32.454 ; RBINS GoogleMaps 5 ♀♀; [Vinh Phuc Province], Me Linh Biodiversity Station ; [21°23′04″ N, 105°42′44″ E]; 20–24 Aug. 2010; J. Constant and P. Limbourg leg.; I.G. 31.668 ; RBINS GoogleMaps 1 ♀; [Vinh Phuc Province], Me Linh Biodiversity Station ; [21°23′04″ N, 105°42′44″ E]; 29 Aug. 2010; J. Constant and P. Limbourg leg.; I.G. 31.668 ; RBINS GoogleMaps 2 ♂♂, 3 ♀♀; Vinh Phuc Province, Dao Tru, Lap Thach ; 17 May 2001; V.T. Hoang leg.; VNMN 3 ♀♀; Vinh Phuc Province, Me Linh ; 24 Apr. 2001; V.T. Hoang leg.; VNMN 2 ♂♂, 17 ♀♀; Vinh Phuc Province, Me Linh ; 26 May 2001; V.T. Hoang leg.; VNMN 1 ♂, 4 ♀♀; Vinh Phuc Province, Me Linh ; 23 Apr. 2007; H.T. Pham leg.; VNMN 1 ♂, 1 ♀; Vinh Phuc Province, Me Linh ; 30–31 May 2007; H.T. Pham leg.; VNMN 7 ♀♀; Vinh Phuc Province, Me Linh ; 26 Jul. 2008; H.T. Pham leg.; VNMN 2 ♀♀; Vinh Phuc Province, Me Linh ; 11 Jun. 2009; H.T. Pham leg.; VNMN 1 ♂, 10 ♀♀; Vinh Phuc Province, Me Linh ; 21 May 2010; H.T. Pham leg.; VNMN 13 ♀♀; Vinh Phuc Province, Me Linh ; 22 May 2010; H.T. Pham leg.; VNMN 3 ♂♂, 27 ♀♀; Vinh Phuc Province, Me Linh ; 4–5 Jun. 2010; H.T. Pham leg.; VNMN 9 ♀♀; Vinh Phuc Province, Me Linh ; 10 Aug. 2011; V.T. Hoang leg.; VNMN 1 ♀; Vinh Phuc Province, Me Linh ; 1 Jun. 2012; V.T. Hoang leg.; VNMN 8 ♀♀; Vinh Phuc Province, Me Linh ; 9–10 Jun. 2012; V.T. Hoang leg.; VNMN 4 ♀♀; Vinh Phuc Province, Me Linh ; 12 Aug. 2011; V.T. Hoang leg.; VNMN 1 ♀; Vinh Phuc Province, Tam Dao N.P. ; May 1998; V.T. Hoang leg.; VNMN 1 ♀; Vinh Phuc Province, Tam Dao N.P. ; Apr. 1999; V.T. Hoang leg.; VNMN 1 ♀; Vinh Phuc Province, Tam Dao N.P. ; May 2003; H.T. Pham leg.; VNMN 1 ♀; Hanoi, Hoan Kiem ; 20 Aug. 2003; V.T. Hoang leg.; VNMN 2 ♀♀; Thua Thien-Hue Province, Phong Dien , near VNMN station; 16°35′12″ N, 107°20′31″ E; 8–9 Apr. 2017; J. Constant and J. Bresseel leg.; I.G. 33.447 ; RBINS GoogleMaps 2 ♂♂, 1 ♀; Quang Ninh Province, Dong Son-Ky Thuong nature reserve ; 21°11′00.4″ N, 107°07′23.8″ E; 575 m a.s.l.; 26 Jul. 2020; H.T. Pham leg.; VNMN GoogleMaps .

Remark

Intraspecific colour variation of the disc of the hind wings exists in this species, with specimens showing bright yellow hind wings and others, milky white. The colour of the tegmina also varies to a great extent, from very dark forms, nearly black with yellow spots ringed with white, to very pale forms, nearly white with yellow spots or even nearly white with cloudy darker, brownish markings in place of the yellow spots. Conspecificity of the different forms was confirmed by the study of the male genitalia and corroborated by the fact that the different forms are sympatric, as illustrated by a series from Chiang Mai collected in June 1992 in Nagai & Porion (1996: figs 194–199). The comparison of the illustrations of the types specimens ( Figs 6A View Fig , 7A View Fig ) with the line illustration of Guérin-Méneville (1829) ( Fig. 8 View Fig ) which matches both type specimens is also very convincing.

Distribution

Northeastern India, southern China, northern Thailand, Laos, North and Central Vietnam. The species is recorded here from Laos for the first time.

Checklist of the Vietnamese species of Pyrops Spinola, 1839

Pyrops atroalbus ( Distant, 1918)

Pyrops buomvoi sp. nov.

Pyrops candelaria (Linné, 1758)

Pyrops clavatus ( Westwood, 1839)

Pyrops coelestinus (Stål, 1863)

Pyrops condorinus ( Lallemand, 1960)

Pyrops ducalis (Stål, 1863)

Pyrops itoi (Satô & Nagai, 1994)

Pyrops lathburii ( Kirby, 1818)

Pyrops spinolae (Westwood, 1842)

Pyrops viridirostris (Westwood, 1848)

Pyrops vitalisius ( Distant, 1918)

Identification key to the species of Pyrops of Vietnam, Laos, Cambodia and Southern Continental China

1. Cephalic process strongly inflated apically ( Figs 9–11 View Fig View Fig View Fig ) ................................................................... 2

– Cephalic process not strongly inflated apically ( Figs 12–19 View Fig View Fig View Fig View Fig View Fig View Fig View Fig View Fig ) ........................................................... 3

2. Abdomen black ventrally ( Fig. 9B View Fig ); tegmina pale yellow-white on disc and with 3 black spots in costal area before nodal line ( Fig. 9A View Fig ); cephalic process yellow ( Fig. 9 View Fig ) .......................................... ........................................................................................................ Pyrops atroalbus ( Distant, 1918)

– Abdomen red ventrally ( Figs 10B View Fig , 11B View Fig ); tegmina largely black on disc ( Fig. 10A View Fig ), or in the pale forms, bluish white on disc and without black spots in costal area ( Fig. 11A, F View Fig ); cephalic process red-brown to black ( Figs 10–11 View Fig View Fig ) ................................................. Pyrops clavatus ( Westwood, 1839)

3. Head entirely green, hind wings bright yellow basally or entirely yellow-orange ( Figs 12–13 View Fig View Fig ) ..... 4

– Head not entirely green, hind wings yellow, white or blue basally ( Figs 14–19 View Fig View Fig View Fig View Fig View Fig View Fig ) ............................ 5

4. Hind wings bright yellow with apical third black ( Fig. 12A View Fig ); ground colour of distal half of tegmina black ( Fig. 12A View Fig ); anterior and median legs largely black ( Fig. 12A–C View Fig ) ............................................ ................................................................................................ Pyrops viridirostris (Westwood, 1848)

– Hind wings entirely bright yellow-orange ( Fig. 13A View Fig ); ground colour of distal half of tegmina green ( Fig. 13A View Fig ); anterior and median legs green ( Fig. 13A–C View Fig ) ............ Pyrops itoi (Satô & Nagai, 1994)

5. Cephalic process black or brown-black dorsally, yellowish ventrally ( Figs 4–5 View Fig View Fig , 14–15 View Fig View Fig ); pro- and mesonotum with median black or dark brown markings ( Figs 4–5 View Fig View Fig , 14–15 View Fig View Fig ) .................................... 6

– Cephalic process yellow-red to red-brown dorsally, paler, yellow to dark red ventrally ( Figs 16–19 View Fig View Fig View Fig View Fig ); pronotum unicolourous, mesonotum sometimes with small brown-black markings on sides ( Figs 16– 19 View Fig View Fig View Fig View Fig ) ..................................................................................................................................................... 8

6. Apex of cephalic process coloured like the rest of the process ( Figs 4–5 View Fig View Fig , 15 View Fig ) ................................. 7

– Apex of cephalic process yellow to orange-red, strongly contrasting ( Figs 6–7 View Fig View Fig , 14 View Fig ) ........................ ............................................................................................................ Pyrops lathburii ( Kirby, 1818)

7. Tegmina narrower: LTg/BTg = 2.9 ( Figs 4A View Fig , 5A View Fig ); spots on basal half of tegmina not fused into bands ( Figs 4A View Fig , 5A View Fig ); cephalic process narrower (LPr/BPrH> 9) ( Figs 4–5 View Fig View Fig ); hind wings bright yellow ( Fig. 4A View Fig ) or white basally ( Fig. 5A View Fig ) ...................................... Pyrops condorinus ( Lallemand, 1960)

– Tegmina broader: LTg/BTg = 2.53 ( Fig. 15A View Fig ); spots on basal half of tegmina fused into transverse bands ( Fig. 15A View Fig ); cephalic process broader (LPr/BPrH <8) ( Fig. 15 View Fig ); hind wings always bright yellow basally ( Fig. 15A View Fig ) ....................................................................... Pyrops spinolae (Westwood, 1842)

8. Hind wings white with sutural margin bordered with black up to the anal lobe ( Fig. 1A View Fig ); cephalic process very narrow (LPr/BPrH = 19) ( Fig. 1 View Fig ) ............................................ Pyrops buomvoi sp. nov.

– Hind wings white, bright yellow or blue with only apical third black ( Figs 16A View Fig , 17A View Fig , 18A View Fig , 19A View Fig ); cephalic process broader (LPr/BPrH <10) ( Figs 16–19 View Fig View Fig View Fig View Fig ) ................................................................. 9

9. Hind wings bright yellow basally ( Fig. 16A View Fig ); cephalic process bright red, yellow ventrally ( Fig. 16 View Fig ); yellow spots on membrane of tegmina ( Fig. 16A View Fig ) ......................... Pyrops candelaria (Linné, 1758)

– Hind wings white or blue basally ( Figs 17A View Fig , 18A View Fig , 19A View Fig ); cephalic process dark red-brown and dark red ventrally or dark red, orange ventrally and at apex ( Figs 17–19 View Fig View Fig View Fig ); yellow spots on membrane of tegmina ( Figs 18–19 View Fig View Fig ) or not ( Fig. 17 View Fig ) ............................................................................................ 10

10. Tegmina without yellow spots on membrane ( Fig. 17A View Fig ); cephalic process dark red, orange ventrally and at apex ( Fig. 17 View Fig ); hind wings pale blue to white basally ( Fig. 17A View Fig ); basal half of tegmina whitish with white-margined green spots in transverse rows, sometimes fused in bands ( Fig. 17A View Fig ) ............................................................................................... Pyrops ducalis (Stål, 1863)

– Tegmina with yellow spots on membrane ( Figs 18A View Fig , 19A View Fig ); cephalic process dark red-brown, dark red ventrally ( Figs 18–19 View Fig View Fig ); hind wings white or blue basally; basal half of tegmina dark green or dark blue-green with yellow spots ( Figs 18A View Fig , 19A View Fig ) ................................................................................11

11. Hind wings blue basally (sometimes faded to nearly white in collections specimens) ( Fig. 18A View Fig ); cephalic process very elongate and strongly curved dorsad ( Fig. 18 View Fig ); ground colour of tegmina dark blue-green ( Fig. 18A View Fig ) ....................................................................... Pyrops coelestinus (Stål, 1863)

– Hind wings white basally ( Fig. 19A View Fig ); cephalic process very elongate and curved dorsad ( Fig. 19 View Fig ); ground colour of tegmina green ( Fig. 19A View Fig ) .................................... Pyrops vitalisius ( Distant, 1918)

Identification key to the species of Pyrops of Vietnam, Laos, Cambodia and Southern Continental China

1. Cephalic process strongly inflated apically ( Figs 9–11 View Fig View Fig View Fig ) ................................................................... 2

– Cephalic process not strongly inflated apically ( Figs 12–19 View Fig View Fig View Fig View Fig View Fig View Fig View Fig View Fig ) ........................................................... 3

2. Abdomen black ventrally ( Fig. 9B View Fig ); tegmina pale yellow-white on disc and with 3 black spots in costal area before nodal line ( Fig. 9A View Fig ); cephalic process yellow ( Fig. 9 View Fig ) .......................................... ........................................................................................................ Pyrops atroalbus ( Distant, 1918)

– Abdomen red ventrally ( Figs 10B View Fig , 11B View Fig ); tegmina largely black on disc ( Fig. 10A View Fig ), or in the pale forms, bluish white on disc and without black spots in costal area ( Fig. 11A, F View Fig ); cephalic process red-brown to black ( Figs 10–11 View Fig View Fig ) ................................................. Pyrops clavatus ( Westwood, 1839)

3. Head entirely green, hind wings bright yellow basally or entirely yellow-orange ( Figs 12–13 View Fig View Fig ) ..... 4

– Head not entirely green, hind wings yellow, white or blue basally ( Figs 14–19 View Fig View Fig View Fig View Fig View Fig View Fig ) ............................ 5

4. Hind wings bright yellow with apical third black ( Fig. 12A View Fig ); ground colour of distal half of tegmina black ( Fig. 12A View Fig ); anterior and median legs largely black ( Fig. 12A–C View Fig ) ............................................ ................................................................................................ Pyrops viridirostris (Westwood, 1848)

– Hind wings entirely bright yellow-orange ( Fig. 13A View Fig ); ground colour of distal half of tegmina green ( Fig. 13A View Fig ); anterior and median legs green ( Fig. 13A–C View Fig ) ............ Pyrops itoi (Satô & Nagai, 1994)

5. Cephalic process black or brown-black dorsally, yellowish ventrally ( Figs 4–5 View Fig View Fig , 14–15 View Fig View Fig ); pro- and mesonotum with median black or dark brown markings ( Figs 4–5 View Fig View Fig , 14–15 View Fig View Fig ) .................................... 6

– Cephalic process yellow-red to red-brown dorsally, paler, yellow to dark red ventrally ( Figs 16–19 View Fig View Fig View Fig View Fig ); pronotum unicolourous, mesonotum sometimes with small brown-black markings on sides ( Figs 16– 19 View Fig View Fig View Fig View Fig ) ..................................................................................................................................................... 8

6. Apex of cephalic process coloured like the rest of the process ( Figs 4–5 View Fig View Fig , 15 View Fig ) ................................. 7

– Apex of cephalic process yellow to orange-red, strongly contrasting ( Figs 6–7 View Fig View Fig , 14 View Fig ) ........................ ............................................................................................................ Pyrops lathburii ( Kirby, 1818)

7. Tegmina narrower: LTg/BTg = 2.9 ( Figs 4A View Fig , 5A View Fig ); spots on basal half of tegmina not fused into bands ( Figs 4A View Fig , 5A View Fig ); cephalic process narrower (LPr/BPrH> 9) ( Figs 4–5 View Fig View Fig ); hind wings bright yellow ( Fig. 4A View Fig ) or white basally ( Fig. 5A View Fig ) ...................................... Pyrops condorinus ( Lallemand, 1960)

– Tegmina broader: LTg/BTg = 2.53 ( Fig. 15A View Fig ); spots on basal half of tegmina fused into transverse bands ( Fig. 15A View Fig ); cephalic process broader (LPr/BPrH <8) ( Fig. 15 View Fig ); hind wings always bright yellow basally ( Fig. 15A View Fig ) ....................................................................... Pyrops spinolae (Westwood, 1842)

8. Hind wings white with sutural margin bordered with black up to the anal lobe ( Fig. 1A View Fig ); cephalic process very narrow (LPr/BPrH = 19) ( Fig. 1 View Fig ) ............................................ Pyrops buomvoi sp. nov.

– Hind wings white, bright yellow or blue with only apical third black ( Figs 16A View Fig , 17A View Fig , 18A View Fig , 19A View Fig ); cephalic process broader (LPr/BPrH <10) ( Figs 16–19 View Fig View Fig View Fig View Fig ) ................................................................. 9

9. Hind wings bright yellow basally ( Fig. 16A View Fig ); cephalic process bright red, yellow ventrally ( Fig. 16 View Fig ); yellow spots on membrane of tegmina ( Fig. 16A View Fig ) ......................... Pyrops candelaria (Linné, 1758)

– Hind wings white or blue basally ( Figs 17A View Fig , 18A View Fig , 19A View Fig ); cephalic process dark red-brown and dark red ventrally or dark red, orange ventrally and at apex ( Figs 17–19 View Fig View Fig View Fig ); yellow spots on membrane of tegmina ( Figs 18–19 View Fig View Fig ) or not ( Fig. 17 View Fig ) ............................................................................................ 10

10. Tegmina without yellow spots on membrane ( Fig. 17A View Fig ); cephalic process dark red, orange ventrally and at apex ( Fig. 17 View Fig ); hind wings pale blue to white basally ( Fig. 17A View Fig ); basal half of tegmina whitish with white-margined green spots in transverse rows, sometimes fused in bands ( Fig. 17A View Fig ) ............................................................................................... Pyrops ducalis (Stål, 1863)

– Tegmina with yellow spots on membrane ( Figs 18A View Fig , 19A View Fig ); cephalic process dark red-brown, dark red ventrally ( Figs 18–19 View Fig View Fig ); hind wings white or blue basally; basal half of tegmina dark green or dark blue-green with yellow spots ( Figs 18A View Fig , 19A View Fig ) ................................................................................11

11. Hind wings blue basally (sometimes faded to nearly white in collections specimens) ( Fig. 18A View Fig ); cephalic process very elongate and strongly curved dorsad ( Fig. 18 View Fig ); ground colour of tegmina dark blue-green ( Fig. 18A View Fig ) ....................................................................... Pyrops coelestinus (Stål, 1863)

– Hind wings white basally ( Fig. 19A View Fig ); cephalic process very elongate and curved dorsad ( Fig. 19 View Fig ); ground colour of tegmina green ( Fig. 19A View Fig ) .................................... Pyrops vitalisius ( Distant, 1918)

BMNH

United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]

RBINS

Royal Belgian Institute of Natural Sciences

VNMN

Vietnam National Museum of Nature

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Fulgoridae

Genus

Pyrops

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Fulgoridae

Loc

Pyrops lathburii ( Kirby, 1818 ) Pyrops

Constant, Jérôme & Pham, Hong Thai 2022
2022
Loc

Pyrops astarte

Pham H. T. 2011: 318
Pham H. T. & Ta H. T. 2004: 58
Liang A. P. 1998: 42
Nagai S. & Porion T. 1996: 25
1996
Loc

Laternaria lathburii

Metcalf Z. P. 1947: 197
1947
Loc

Laternaria astarte

Metcalf Z. P. 1947: 186
1947
Loc

Fulgora astarte

Lallemand V. 1963: 85
Distant W. L. 1918: 198
1918
Loc

Fulgora astarte

Distant W. L. 1914: 409
1914
Loc

Pyrops lathburii

Jiaranaisakul K. & Constant J. 2021: 18
Wang W. Q. & Xu S. - L. & Qin D. - Z. 2018: 297
Constant J. & Pham H. T. 2017: 18
Constant J. & Phauk S. & Bourgoin T. 2016: 15
Constant J. 2015: 9
Pham H. T. 2011: 318
Liang A. P. 1998: 43
Nagai S. & Porion T. 1996: 25
Schaum H. R. 1850: 64
1850
Loc

Hotinus lathburii

Girard M. 1885: 859
Dohrn F. A. 1859: 57
Walker F. 1851: 266
Amyot C. & Audinet-Serville J. 1843: 491
1843
Loc

Fulgora lathburii

Lallemand V. 1963: 86
Baker C. F. 1925: 349
Schmidt E. 1919: 126
Schmidt E. 1911: 161
Distant W. L. 1906: 186
Atkinson E. T. 1885: 129
Gadeau de Kerville H. 1881: 43
Butler A. G. 1874: 97
Stal C. 1870: 741
Guerin-Meneville F. E. 1844: 357
White A. 1844: 425
Spinola M. 1839: 234
Westwood J. O. 1839: 139
1839
Loc

Flata lathburii

Germar E. F. 1830: 46
1830
Loc

Fulgora lathburii

Kirby W. F. 1818: 450
1818