Urophonius granulatus, Pocock, 1898

Ojanguren-Affilastro, Andrés A., Ochoa, José A., Mattoni, Camilo I. & Prendini, Lorenzo, 2010, Systematic Revision of the granulatus Group of Urophonius Pocock, 1893 (Scorpiones, Bothriuridae), with Description of a New Species from Central Chile, American Museum Novitates 2010 (3695), pp. 1-44 : 5

Urophonius granulatus group

Figure 1; table 1

DIAGNOSIS: The four species in the granulatus group of Urophonius can be separated from other species in the genus by means of the following combination of characters. Metasomal segment I, VSM carinae distinct, usually subparallel throughout length (diverging slightly at anterior margin of segment in some species) ( figs. 3A, C, D , 4A, C, D ), except for U. somuncura , in which it is absent or reduced to a few granules ( figs. 3B , 4B ). Metasomal segment I with two or three pairs of ventral macrosetae; II and III each with three pairs of ventral macrosetae ( figs. 3 , 4 ). Telotarsal spination, leg III: 5–5/5–6; IV: 5–6/6–7. Hemispermatophore, lobe region small to medium sized; capsular concavity very well developed; area between basal lobe and external lobe concave ( fig. 8A, C, E, G ); internal lobe with small, bilobed apophysis disconnected from distal lamina ( fig. 8B, D, F, H ). Trichobothrial pattern, neobothriotaxic major Type C, with one accessory trichobothrium in V series of chela; femur with 3 trichobothria (d, i, e); patella with 19 (3 V, 2 d, i, 3 et, est, 2 em, 2 esb, and 5 eb); chela with 27 (Est, 5 Et, 5 V, Esb, 3 Eb, Dt, Db, et, est, esb, eb, dt, dst, dsb, db, ib, and it); femoral trichobothrium e situated close to a single dorsal macroseta (M1), variable with respect to macrosetae in different species (figs. 10A, 13A, 16A, 19A). Chelicera, movable finger with two well-developed subdistal teeth. Metasomal segments, ventral surfaces each with paired VL and single VM pigmentation stripes, without paired VSM stripes.

Figures

FIGURE 3. Urophonius granulatus group, sternite VII and metasomal segments I and

FIGURE 4. Urophonius granulatus group, sternite VII and metasomal segments I and II, ventral aspect. A. Urophonius granulatus Pocock, 1898, ♀ (LBRE). B. Urophonius somuncura Acosta, 2001, ♀ (MACN). C. Urophonius tregualemuensis Cekalovic, 1981, ♀ (AMNH). D. Urophonius pizarroi, n. sp., paratype ♀ (MZUC). Scale bars = 1 mm.

FIGURE 8. Urophonius granulatus group, sinistral hemispermatophore, ectal and ental aspects. A, B. Urophonius granulatus Pocock, 1898, ♂ (MACN). C, D. Urophonius somuncura Acosta, 2001, ♂ (MACN). E, F. Urophonius tregualemuensis Cekalovic, 1981, ♂ (MACN). G, H. Urophonius pizarroi, n. sp., paratype ♂ (MZUC). Scale bars = 1 mm.