Cobalopsis adictys Grishin, 2023

Zhang, Jing, Cong, Qian & Grishin, Nick V., 2023, Supplementary Materials and Appendix, Insecta Mundi 2023 (26), pp. 1-115 : 62-63

Cobalopsis adictys Grishin , new species

https://zoobank.org/ CBF4DB85-179A-4240-912C-8B7186AFB049

( Fig. 6 part, 153–154, 380–381)

Definition and diagnosis. Phylogenetic trees reveal that a number of specimens identified as Cobalopsis dictys (Godman, 1900) (type locality in Mexico: Veracruz, Guatemala, Costa Rica, Panama, illustrated specimen from Guatemala) show prominent genetic differentiation from it ( Fig. 6): e.g., their COI barcodes differ by 3.3% (22 bp), and therefore represent a new species. This new species keys to “ Papias dictys ” (J.36.6) in Evans (1955) but differs from it by comparatively darker ventral forewing and hindwing area by inner margin, vestigial or absent forewing apical spot ( Fig. 153–154), smaller tooth on ventral margin of harpe, distal margin of harpe stronger projecting dorsad, and more angled connection between tegumen and uncus in lateral view ( Fig. 381). Due to the cryptic nature of this species, most reliable identification is achieved by DNA and a combination of the following base pairs is diagnostic in the nuclear genome: aly 1905.5.2:A148G, aly260.3.2:A28T, aly260.3.2:T51C, aly451.31.3:C84T, aly318.41.3:G78A, and COI barcode: A100T, T112C, C343T, T445C, T536C.

Barcode sequence of the holotype. Sample NVG-19018H05, GenBank OR837691, 658 base pairs: AACTTTATACTTCATCTTTGGAATTTGAGCCGGAATATTAGGAACTTCTTTAAGATTATTAATTCGAACAGAATTAGGTAATCCTGGATCTTTAATT GGTGATGATCAAATCTATAATACTATTGTAACAGCTCATGCTTTTATTATAATTTTTTTTATAGTTATACCTATTATAATTGGAGGATTTGGAAATT GATTAGTACCCCTAATACTTGGTGCTCCTGATATAGCTTTCCCACGAATAAATAATATAAGTTTTTGAATACTACCACCTTCCTTAATATTACTAAT TTCAAGAAGAATTGTAGAAAATGGGGCAGGTACTGGTTGAACTGTATACCCTCCTCTTTCTTCTAATATTGCACATCAAGGAGCATCTGTTGATTTA GCAATTTTTTCTCTTCATTTAGCTGGTATTTCTTCTATTCTAGGAGCTATTAATTTCATTACTACAATTATTAATATACGAATTAAAAATTTATCTT TTGATCAAATACCTTTATTTGTCTGATCAGTTGGAATTACAGCTTTACTACTACTATTATCTTTACCTGTACTAGCAGGAGCTATTACTATACTTCT TACTGATCGAAATTTAAATACTTCTTTTTTTGATCCTGCAGGAGGAGGAGATCCTATTTTATACCAACATTTATTT

Type material. Holotype: ♂ deposited in the National Museum of Natural History, Smithsonian Institution , Washington, DC, USA ( USNM), illustrated in Fig. 153–154, bears the following four rectangular labels, three white: [ PANAMA: VERAGUAS | Sante Fe | 8°31′N 81°05′W | 12.IX.1981 | leg. G.B.Small], [DNA sample ID: | NVG-19018H05 | c/o Nick V. Grishin], [USNMENT | {QR Code} | 01532536], and one red [HOLOTYPE ♂ | Cobalopsis | adictys Grishin ] . Paratypes: 2♂♂ from Costa Rica: Area de Conservación Guanacaste, Guanacaste Prov., Sector Pitilla [ USNM]: NVG-18064B08, 11-SRNP-33187 Sendero Nacho, 710 m, GPS 10.9845, −85.4248, eclosed 17-Nov- 2011 ; NVG-18064C02, 11-SRNP-30141 Sendero Laguna 680 m GPS 10.9888, −85.4234, eclosed 3-Mar-2011 .

Type locality. Panama: Veraguas Province, Santa Fe, GPS 8.5167, −81.0833.

Etymology. The name is derived by combining dictys with the negating prefix “a”. As a mnemonic, a longer name indicates it is a more southern species than C. dictys . The name is a noun in apposition.

Distribution. Costa Rica and Panama.