Cunninghamites ubaghsii DEBEY ex UBAGHS

Greguš, Josef & Kvaček, Jiří, 2015, Revision Of Cenomanian Flora From The Maletín Sandstone, Acta Musei Nationalis Pragae Series B 71 (3 - 4), pp. 315-364 : 322-323

Cunninghamites ubaghsii DEBEY ex UBAGHS

Pl. 4, Fig. 9; Pl. 5, Fig. 1–2

1869 Pinus quenstedti HEER , p. 13, pl. 2, fig. 5–9, pl. 3, fig. 1.

1885 Cunninghamites ubaghsii DEBEY ex UBAGHS , p. 28.

2004 Cunninghamites ubaghsii (DEBEY ex UBAGHS) ; van der Ham and van Konijnenburg-van Citter, p. 28, fig. 7.

2004 Cunninghamites ubaghsii (DEBEY ex UBAGHS) ; van der Ham et al., p. 94, fig. 3–6.

2012 Cunninghamites ubaghsii (DEBEY ex UBAGHS) ; Bosma et al., p. 25, pl. 2, fig. 1.

2013 Cunninghamites ubaghsii (DEBEY ex UBAGHS) ; Halamski, p. 8, fig. 5J, 6B.

H o l o t y p e: No. b4318 (housed in the Royal Belgian Institute of Natural Sciences), ( Ubaghs 1885, p. 28; van der Ham et al. 2004, fig. 3).

T y p e l o c a l i t y: Kunrade, province Limburg ( the

Netherlands).

S t r a t i g r a p h y: Maastrichtian, Late Cretaceous.

M a t e r i a l s t u d i e d: GPIT/PL_654, 730, 757.

O c c u r r e n c e: Maletín; Krasnobród, Poland (Campanian-Maastrichtian).

D e s c r i p t i o n. Specimen no. GPIT/PL_730 (Pl. 5, Fig. 1a, b) described by Heer (1869, p. 13, pl. 2, fig. 5) as Pinus quenstedtii HEER is a 75 mm long and 12 mm wide fragment of a shoot, with helically arranged, clearly visible leaf cushions of obovate rhombic shape. Leaf cushions are 4–5 mm long and 2–3 mm wide (Pl. 5, Fig. 1b). Linear leaves up to 100 mm long leave the leaf cushions at a 45° angle. Apex of the needle is not as well apparent as the base. In the central part of the needle, there is one pronounced groove/keel. Leaf cushions bearing needles are apparent only in the upper half of the axis fragment. In the lower half of the axis, remains of leaf cushions form small helically arranged 2–3 mm wide pits. Specimen no. GPIT/PL_757 (Pl. 5, Fig. 2a, b) described by Heer (1869, p. 13, pl. 3, fig. 1–3) is a 200 mm long and 12 mm wide shoot, with helically arranged, clearly visible leaf cushions of obovate to rhombic shape (5–6 mm long and 3–4 mm wide) bearing needles up to 200 mm. Specimen no. GPIT/PL_654 (Pl. 4, Fig. 9) is a fragment of longitudinally broken shoot bearing helically arranged needles. Due to lack of apical parts of needles, their length has to be more than 40 mm.

D i s c u s s i o n. The studied specimens determined here as C. ubaghsii were originally described by Heer (1869) as Pinus quenstedtii . They have needles growing out of leaf cushions separately. Needles arranged in brachyblasts are a typical distinctive character of the genus Pinus LINNAEUS. Within the Bohemian Cretaceous Basin, the genus Pinus is known from the Cenomanian locality of Lanšperk as Pinus landsbergensis J. KVAČEK , showing typical brachyblasts (J. Kvaček 2013). C. ubaghsii differs from other species of the genus Cunninghamites in having very long needle-like leaves ( Bosma et al. 2012).