Foveacheles (Mediostella) carnichensis Zacharda, 2011
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publication ID |
https://doi.org/ 10.1080/00222933.2010.535915 |
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persistent identifier |
https://treatment.plazi.org/id/0382502C-9005-FFFC-FE9F-FED5E31425F0 |
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treatment provided by |
Felipe |
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scientific name |
Foveacheles (Mediostella) carnichensis Zacharda |
| status |
sp. nov. |
Foveacheles (Mediostella) carnichensis Zacharda , sp. nov.
( Figures 4–7 View Figure 4 View Figure 5 View Figure 6 View Figure 7 )
Material examined
Holotype. Adult female, Italy, Friuli, Prealpi Carniche. Clauzetto (Pordenone); Grotta sotto il Ponte sul Rio Molino Cave , coll. E. Piva, 28 June 1998;
A,B A / G B,D,E C F
deposited in the Museum of Biological Diversity, Ohio State University , Type No. OSAL0007415 View Materials .
Diagnosis
Cheliceral digits slender, elongated, masticatory surface of fixed digit smooth and with small prebasal ventrolateral fissure lying slightly proximad of insertion of proximal cheliceral seta; movable digit serrated along approximately distal two-thirds of masticatory surface and with blunt prebasal tubercle. Rhagidial organ I comprises four long, slender rhagidial solenidia lying obliquely in separate depressions. Rhagidial organ II with four rhagidial solenidia lying in separate, slightly oblique depressions, some may be confluent. Small spiniform famulus (ε) subtends proximal rhagidial solenidion. Spiniform solenidion on genu III and tibia IV dorsal, dorsoproximal.
Description
Adult female (one examined). Length of idiosoma 1040 µm, ratio of leg I length to idiosomal length 1.23.
Gnathosoma . Subcapitulum broadly oval ( Figure 5F View Figure 5 ), subtriangular; ratio of length to breadth 1.50, distal hypostomal lips with spiniform internal and serrate external malar processes, adoral setae nude and only slightly overlapping apex of subcapitulum, proximal subcapitular setae pubescent, external pair slightly longer than internal pair. Dorsal surface of chelicera with distinct saddle-shaped depression at level of bases of digits, cheliceral digits slender, dorsal surface of fixed digit with distinct narrow rim above insertion of distal cheliceral seta, fixed digit smooth along masticatory surface and with small prebasal ventrolateral fissure ( Figure 5A View Figure 5 ), movable digit serrated along approximately distal two-thirds of masticatory surface, with blunt prebasal tubercle ( Figure 5B View Figure 5 ). Chelicera with two setae, proximal seta inserted just distad of articulation of movable digit, tip of proximal seta not reaching insertion of distal seta, tip of distal seta slightly overlaps apex of fixed digit. Length of chelicera 264 µm, dorsoventral width 112 µm, length of movable digit 102 µm, length of proximal and distal cheliceral setae 26 and 39 µm, respectively, distance between their insertions 33 µm. Ratios: cheliceral length to dorsoventral width 2.35, length of movable digit to length of chelicera 0.38, length of movable digit to dorsoventral width of chelicera 0.91. Palpal tarsus shorter than femorogenu ( Figure 5C View Figure 5 ), ratio of length to width of tarsus 2.66. Length of palpal trochanter, femorogenu, tibia and tarsus 39, 132, 49, 105 µm, respectively. Number of setae and solenidia (in brackets) on palpal trochanter, femorogenu, tibia and tarsus 0-2-3-10(1), respectively, tarsal solenidion spiniform, erect.
Prodorsum. Naso well-developed, with pair of internal vertical setae v 1. Bothridial setae sc 1 filiform, finely pubescent, slightly overlapping disjugal suture ( Figure 4A View Figure 4 ). Length of setae: v 1 59, v 2 45, sc 1 112, sc 2 132 µm.
Opisthosomal dorsum and anal region. Cupules; ia located laterally between insertions of setae c 1 and d 1, im lateral and just anterior to setae e 1, ip distad of insertions of f 2 setae, about one third of distance between insertions of e 1 and f 1, ih positioned ventrolaterally, laterodistad of insertions of adanal setae ad 1. Setae c 1, d 1, e 1 short, reach about one-third of distance to insertion of successive seta, seta f 1 reaches about two-thirds of distance to insertion of h 1. Length of setae: c 1 49, c 2 138, d 1 46, e 1 46, f 1 82, f 2 63, h 1 119, h 2 69, ps 1 99, ps 2 56, ps 3 36, ad 1 25 µm.
Podosoma. Coxae I, II, III, IV with 3-1-5-3 finely pubescent setae, respectively ( Figure 4 B View Figure 4 ).
Genital region. Genital valves each with five finely pubescent genital setae (g) of similar length, about 23 µm, arranged evenly along lateral edge of valve. Five pairs of aggenital setae (ag) of similar length, about 36 µm. Length of genital valves 148 µm.
Legs. Leg I 1280 µm long, about 1.23 times as long as idiosoma. Empodia of all legs setulose and reaching to tips of claws. Number of setae and solenidia (solenidia and famulus (ε) bracketed), respectively, on legs I-II-III-IV (Figures 6,7): trochanters 1-1-2-2; basifemora + telofemora 5+5-6+5-4+4-3+4; genua 11(1)-9(1)-7(1)-6; tibiae 11(2)-7(2)-7(2)-7(1); tarsi 19(4+ ε)-16(3+ ε)-14-14. Genua I and II each with one erect spiniform distiventral solenidion (σ); genu III with one small spiniform dorsal medioproximal solenidion. Tibia I with one erect spiniform dorsoproximal solenidion, and one dorsodistal rhagidial solenidion (Φ); tibia II with one spiniform erect dorsoproximal solenidion, and one lanceolate dorsodistal solenidion recessed in deep pit with small surface pore ( Figure 5G View Figure 5 ); tibia III with two adjacent erect spiniform dorsomedial solenidia; tibia IV with one erect spiniform dorsoproximal solenidion. Tarsus I robust, its tip abruptly tapered in lateral view ( Figure 6A View Figure 6 ), ratio length to width 5.20, with four rhagidial solenidia (ω) lying obliquely in separate depressions dorsodistally; stellate famulus (ε) inserted between second and third rhagidial solenidia antiaxially ( Figure 5D View Figure 5 ); tarsus II with four rhagidial solenidia lying in separate or confluent, slightly oblique depressions. Small spiniform famulus (ε) subtending proximal rhagidial solenidion ( Figure 5E View Figure 5 ).
Affinities
Foveacheles (M.) carnichensis n. sp. resembles F. (M.) canestrini (Berlese and Trouessart, 1889) from which it can be distinguished by the following principal characteristics. (1) The spiniform famulus (ε) in rhagidial organ II subtends the proximal rhagidial solenidion; in F. (M.) canestrini it lies between the first and second rhagidial solenidion antiaxially. (2) The cheliceral fixed digit is slender; in F. (M.) canestrini it is robust, broad. (3) The spiniform medioproximal solenidion on tibia IV is positioned dorsally; in F. (M.) canestrini it is positioned ventrally.
Foveacheles (M.) carnichensis sp. nov. is also similar to F. (M.) pribilovi Zacharda, 1996 from which it can be distinguished by the following morphological characters. (1) The cheliceral fixed digit is slender; in F. (M.) pribilovi it is robust, broad. (2) The medioproximal spiniform solenidion on genu III is positioned dorsally; in F. (M.) pribilovi it is positioned ventrally.
Etymology
The epithet carnichensis reflects the name of the Prealpi Carniche Mountains where the mite was collected.
Key to adults of Foveacheles (Mediostella)
1. Masticatory surface of cheliceral fixed digit with distinct, sharply pointed cusp. Nude adoral setae short, not reaching apex of subcapitulum. Palpal tibia with two pubescent setae. Nearctic region............................
.................................... F. (M.) vancouverensis Zacharda, 1986 View in CoL
Masticatory surface of cheliceral fixed digit smooth. Nude adoral setae long, reaching or overlapping apex of subcapitulum. Palpal tibia with three pubescent setae........................................................ 2
2. Rhagidial organ II with three rhagidial solenidia. Austrian Alps.................................................... F. (M.) willmanni Zacharda, 1980 View in CoL
Rhagidial organ II with four rhagidial solenidia.......................... 3
3. Solenidia in rhagidial organ II oblique and separate, spiniform famulus (ε) inserted between 1st and 2nd proximal solenidia antiaxially. Palearctic region.......... F. (M.) canestrini (Berlese and Trouessart, 1889)( Zacharda, 1980) View in CoL
At least some rhagidial solenidia in rhagidial organ II arranged in tandem axially, separate or lying in confluent depressions, spiniform famulus (ε) subtends proximal rhagidial solenidion.................................. 4
4. Rhagidial organ II with solenidia lying in confluent depressions. Spiniform solenidion on tibia IV medioproximal, ventral. Slovakia, the High Tatras................................... F. (M) rupestris Zacharda (1980)
Rhagidial organ II with solenidia lying in separate depressions; spiniform solenidion on tibia IV medioproximal, lateroventral or dorsoproximal
....................................................................... 5
5. Coxa III with five pubescent setae. Spiniform solenidion on tibia IV dorsoproximal. Masticatory surface of cheliceral movable digit denticulate and with blunt prebasal tubercle. Northern Italy...............................
............................................... F. (M.) carnichensis sp. nov.
Coxa III with six pubescent setae. Spiniform solenidion on tibia IV lateroventral. Masticatory surface of cheliceral movable digit denticulate, but without blunt prebasal tubercle. Nearctic region...................................
.......................................... F. (M.) pribilovi Zacharda (1996) View in CoL
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