Quedius (Raphirus) nemo, Tephens, 2014

Tephens, 2014, Contributions to the knowledge of the Quediina (Coleoptera, Staphylinidae, Staphylinini) of China. Part 48. Genus Quedius STEPHENS, 1829. Subgenus Raphirus STEPHENS, 1829. Section 12., Linzer biologische Beiträge 46 (2), pp. 1813-1818 : 1814-1816

Quedius (Raphirus) nemo nov.sp. ( Figs 1-5 )

T y p e l o c a l i t y. CHINA: Sichuan, Erlang Shan, 29°32’N 102°18’E, 2200-2300m.

T y p e m a t e r i a l. Holotype (male): CHINA: " China: Sichuan Prov. Erlangshan Mt. 29°32’N, 102°18’E, alt. 2200-2300m 13- VII-2012 PENG, DAI& YIN leg.", in SNUC. Allotype (female): CHINA: " China: Sichuan Prov. Tianquan County Erlangshan Mt. Yakou, 3.6 km 29°31’N, 102°17’E alt. 2600-2800 m 11.VII. 2012 PENG, DAI & YIN leg.", in SNUC. Paratypes: same data as holotype, 1 (ASC).

D e s c r i p t i o n. In all characters similar to Q. shunichii SMETANA, 2010 , but different by a few external characters, and particularly by the differently shaped aedoeagus. Average size larger, coloration darker: entirely piceous-black, antennae black with last segment and sometimes first two segments paler, legs piceous-black with somewhat paler tarsi.

Male. First four segments of front tarsus markedly dilated, slightly more so than those of Q. shunichii , each densely covered with tenent setae ventrally, segment 2 wider than apex of tibia (ratio 1,20) segment 4 narrower than preceding two segments. Sternite 5 with apical margin truncate in middle and in front of it with large medial field of denser punctures and setae, extending to base of sternite; sternite 6 with apical margin slightly, widely concave, with large medial subtriangular area in front it depressed and densely punctate and setose, delimited at each lateral margin by densely set, long stiff setae; sternite 7 with modification similar to that on sternite 6, except that the depressed and densely punctate and setose area is markedly more extensive; sternite 8 with wide and deep obtusely rounded medioapical emargination, large area, narrow at the emargination, but markedly widened and extended to the base of sternite depressed and densely punctate and pubescent. Genital segment with tergite 10 markedly narrowed toward broadly arcuate apex, with numerous setae at and near apical margin, otherwise asetose ( Fig. 1 ); sternite 9 with characteristic, sharply delimited basal portion, apical portion arcuate apically, setose as in Fig. 2 . Aedoeagus ( Figs 3, 4 ) similar to that of Q. shunichii , but in general more robust; anterior characteristic plate of median lobe is wider and paramere is markedly more robust and of widely fusiform shape, whereas it is narrower and more or less parallelsided in Q. shunichii ( Figs 3,4 , and Fig. 3 in SMETANA 2010: 251).

Female. First four segments of front tarsus only slightly dilated, vaguely subbilobed, each with a few tenent setae ventrally, segment two narrower than apex of tibia (ratio 0.72); segment four narrower than preceding segments. Genital segment with second gonocoxites long and narrow, each with extremely minute stylus bearing one long, strong seta; tergite 10 markedly narrowed, apically attenuated into moderately long, dagger-like apical portion; apical portion and medial portion of tergite in front of it pigmented; setae present on dagger-like apical portion and for some distance medially in front of it ( Fig. 5 ).

Length 10.0- 10.5 mm. E t y m o l o g y. The specific epithet is the Latin noun nemo, m. (nobody) in apposition.

G e o g r a p h i c a l d i s t r i b u t i o n Quedius nemo is at present known only from Erlang Shan in Sichuan.

B i o n o m i c s.Nothingisknown about the collecting circumstances of the specimens of the original series.

R e c o g n i t i o n a n d c o m m e n t s Quedius nemo may be distinguished from the closely related and similar Q. shunichii by the characters mentioned above. The male sexual characters on the abdominal sternites are similar in both species, but they are distinctly more pronounced on sternites in Q. nemo , particularly the depressions on sternites 6 and 7 being deeper and the setae delimiting them laterally being more numerous and longer.

Figures

Figs 1-10: 1-5. Quedius nemo nov.sp.: (1) tergite 10 of male genital segment; (2) sternite 9 of male genital segment; (3) aedoeagus, ventral view; (4) underside of apical portion of paramere; (5) tergite 10 of female genital segment. 6-10. Quedius egregius nov.sp. (6) tergite 10 of male genital segment; (7) sternite 9 of male genital segment; (8) aedoeagus, ventral view; (9) apical portion of median lobe of aedoeagus, lateral view; (10) underside of apical portion of paramere.