Systaria decidua, Dankittipakul, Pakawin & Singtripop, Tippawan, 2011

Dankittipakul, Pakawin & Singtripop, Tippawan, 2011, Seven new species of Systaria Simon, 1897 from Southeast Asia (Araneae, Clubionidae, Systariinae), Zootaxa 2905, pp. 16-32 : 21-23

Systaria decidua sp. nov.

Figs 9–14

Type material. Holotype: 3, THAILAND: Phetchabun Province: Nam Nao National Park, 16°43.156'N 101°35.118'E, pitfall trap, 7 July 2008, leg. P. Dankittipakul, ( MHNG, Sys-08). Paratypes: 13, same data as holotype ( TNHM). THAILAND: Lamphang Province: 2Ƥ, Doi Khun Tan National Park, 1200 m, 25-27 January 2006, leg. P. Dankittipakul ( MHNG, Sys-09). Uttaradit Province: 1Ƥ, Nam Pad District, Phu Soi Dao National Park, 1500-1600 m, mixed evergreen hill forest on flat plateau, 23 December 2005, leg. P. Dankittipakul ( MHNG, Sys-10).

Diagnosis. Males of S. decidua sp. nov. are similar to those of S. bifida sp. nov. and S. acuminata sp. nov. in having a triangular cymbial tubercle but can be easily distinguished by the RTA with a sharply pointed ventral process directing horizontally and a curved dorsal process ( Figs 9–10 ). Females closely resemble those of S. hainanensis Zhang et al., 2009 but can be distinguished by a broader epigynal atrium provided with a median septum ( Figs 12–13 ).

Etymology. The specific epithet is an adjective (Latin: deciduous, -a, -um), referring to the type of forest where this new species inhabits.

Description. Male (holotype): Total length 7.84. Prosoma 3.56 long, 2.44 wide. Opisthosoma 4.28 long, 2.08 wide. Leg formula: 4123; I 10.82 (2.76, 1.48, 2.84, 2.44, 1.30); II 9.14 (2.68, 1.28, 2.48, 1.72, 0.98); III 7.80 (2.18, 0.78, 1.78, 2.14, 0.92); IV 11.70 (3.28, 1.02, 3.12, 3.16, 1.12). Spination. Femora I d1-1-0, p1-1-1, r0-1-1; II d1-2, p1-1-1, r0-0-1; III d1-2, p0-1-1, r1-1-1; IV d1-0-1, p0-1-1, r1-1-1. Tibiae: I v2-2 -2; II v2-2 -2; III d1-0-0, p1-1-0, r2- 2-2, v2-2 -2; IV d0-0-1, p1-1-0, r1-1-1, v2-2 -2. Metatarsi: I v1 -0-0; II v1-1 -1; III p1-1-2, r1-1-1-1, v1 -0-1; IV p1-1- 1, r2-2-2, v1-2. Eye size and interdistances: AME 0.12, ALE 0.08, PME 0.06, PLE 0.06; AME–AME 0.16, AME– ALE 0.12, PME–PME 0.20, PME–PLE 0.14, ALE–PLE 0.12; MOA 0.24 long, anterior width 0.28, posterior width 0.26.

Palp ( Figs 9–11 ). Ventral surface of tibia protruding retrolaterally ( Fig. 10 ), provided with dorsal outgrowth ( Fig. 11 ). RTA with triangular ventral process directed horizontally ( Fig. 9 ); dorsal process curved, its apex sharply pointed ( Fig. 10 ). Cymbial tubercle triangular, sharply pointed. Cymbial fold entirely membranous ( Fig. 9 ). Tegulum excavated meso-prolaterally, protruding baso-prolaterally. Embolus triangular, embolic base membranous, apex of embolus filiform. Conductor membranous, broad at base, bifid apically.

Female (paratype, MHNG, Sys-09): Total length 10.48. Prosoma 4.78 long, 3.30 wide. Opisthosoma 5.68 long, 3.06 wide. Leg formula: 4123; I 9.02 (2.66, 1.34, 2.52, 1.62, 0.88); II 8.50 (2.58, 1.22, 2.32, 1.56, 0.82); III 8.10 (2.86, 0.88, 1.74, 1.82, 0.80); IV 12.32 (3.56, 1.28, 2.68, 3.22, 1.58). Spination. Femora: I d1-1-1, p1-0-0, r0-1-1; II d1-2, p0-1-1, r0-1-2; III d1-2, p0-1-1, r1-2; IV d1-1-1, p1-1-1, r0-0-1. Tibiae: I v2-1 -1; II v0-1-1; III p1-1-0, r1-0-1, v2-2 -2; IV p1-1-0, r1-0-1, v2-2 -2. Metatarsi: II v1 -0-0; III p1-1-1, r1-1-2, v2-1 -1; IV p1-1-2, r1-1-2, v2-2 -2. Eye size and interdistances: AME 0.14, ALE 0.12, PME 0.10, PLE 0.10; AME–AME 0.16, AME–ALE 0.14, PME– PME 0.20, PME–PLE 0.18, ALE–PLE 0.12; MOA 0.6 long, anterior width 0.34, posterior width 0.32.

Genitalia ( Figs 12, 14 ). Epigynal atrium shallow, broad anteriorly, slightly narrower posteriorly. Genital orifices situated medially, separated by elevated median septum. Anterior bursae sclerotized, elongated, widest in the middle. Insemination ducts inflated, spherical, spermathecae of similar size. Fertilization ducts lanceolated.

Natural history. Types of S. decidua sp. nov. were collected from deciduous forests.

Distribution. Northern Thailand (Fig. 43).

Figures

FIGURES 9 – 14. Systaria decidua sp. nov., holotype male (9 – 11) and paratype female (12 – 14). 9 – 11 Left male palp (9 ventral, 10 retrolateral, 11 dorsal); 12 – 13 Epigyne, ventral; 14 Vulva, dorsal. Scale lines = 0.1 mm.