Systaria acuminata, Dankittipakul, Pakawin & Singtripop, Tippawan, 2011

Dankittipakul, Pakawin & Singtripop, Tippawan, 2011, Seven new species of Systaria Simon, 1897 from Southeast Asia (Araneae, Clubionidae, Systariinae), Zootaxa 2905, pp. 16-32 : 28-29

Systaria acuminata sp. nov.

Figs 32–37

Type material. Holotype: 3, THAILAND: Krabi Province: Aao Luk District, outside Waree-Rin Cave, 60 m, oil palm plantation, 13 September 2006, leg. P.J. Schwendinger ( MHNG, Sys-13).

Paratypes: 13, same data as holotype, ( TNHM). INDONESIA: Mapur Island: 23, disturbed rainforest ca. 5 km northwest of Mapur Niang, 10-50 m, 21 June 2001, leg. P.J. Schwendinger ( MHNG, SIM- 01/05, Sys-14). Sumatra: 1Ƥ, Jambi Province: Mt. Tujuh, footpath to Lake Mt. Tujuh, 1,500-2,000 m, evergreen hill forest, 20 February 2000, leg. P.J. Schwendinger ( MHNG, Sys-17).

Diagnosis. Males of S. acuminata sp. nov. can be recognized by the shape of RTA, and by the abruptly bent apex of embolus. Females closely resemble those of S. dentata sp. nov. and S. cervina in the shape of their internal genitalia but can be distinguished by the more elongate anterior bursae.

Etymology. The specific epithet is a Latin adjective (acuminatus, -a, -um), referring to a gradually tapering embolus on the male palp.

Description. Male (holotype): Total length 8.80. Prosoma 3.68 long, 2.56 wide. Opisthosoma 4.40 long, 2.20 wide. Leg formula: 4123; I 11.88 (3.04, 1.68, 2.96, 2.68, 1.52); II 10.18 (2.88, 1.46, 2.70, 1.96, 1.18); III 8.78 (2.38, 0.98, 1.98, 2.46, 0.98); IV 12.70 (3.52, 1.24, 3.28, 3.38, 1.28). Spination. Femora I d1-1-1, p1-1-1, r0-1-1; II d1-1-1, p1-1-1, r0-1-1; III d1-1-1, p1-1-0, r1-1-1; IV d1-1-1, p0-1-1, r1-1-1. Tibiae: I v2-2 -2; II v2-2 -2; III d0-1-1, p1-1-0, r2-2-2, v2-2 -2; IV d0-1-0, p1-1-0, r1-1-0, v2-2 -2. Metatarsi: I v1 -0-0; II v1-1 -1; III p1-1-2, r1-1-2, v1 -0-1; IV p1-1-1, r2-2-2, v1-1 -1. Eye size and interdistances: AME 0.14, ALE 0.10, PME 0.08, PLE 0.08; AME–AME 0.18, AME–ALE 0.16, PME–PME 0.24, PME–PLE 0.16, ALE–PLE 0.12; MOA 0.30 long, anterior width 0.30, posterior width 0.28.

Palp ( Figs 32–34 ). Ventral surface of tibia protruding retrolaterally ( Figs 32, 34 ). RTA provided with sharply pointed ventral process and flange-like dorsal one. Cymbial fold partially membranous. Cymbial tubercle sharply pointed. Embolus rostrated; embolic base membranous; apex of embolus elongated, abruptly bent subapically. Conductor spoon-shaped, with broad basal flange.

Variation. The dorsal process on the RTA of the male palp is dorsally notched in a male paratype collected from Sumatra ( Fig. 35 ).

Female (paratype, MHNG, Sys-17): Total length 10.44. Prosoma 4.78 long, 3.44 wide. Opisthosoma 5.66 long, 3.08 wide. Leg formula: 4123; I 9.26 (2.68, 1.40 2.58, 1.66, 0.94); II 8.70 (2.64, 1.28, 2.26, 1.64, 0.88); III 8.38 (2.90, 0.88, 1.78, 1.94, 0.88); IV 12.58 (3.56, 1.34, 2.82, 3.28, 1.58). Spination. Femora: I d1-1-1, p1-1-0, r1-1-0; II d1-1-1, p0-1-1, r1-1-1; III d1-1-1, p1-0-1, r1-1-1; IV d1-1-1, p1-1-1, r1-0-0. Tibiae: I v2-2 -0; II v1-1 -0; III p1-1-0, r1-0-1, v2-2 -2; IV p1-1-0, r1-1-0, v2-2 -2. Metatarsi: II v1-1 -0; III p1-1-1, r1-1-2, v2 -0-2; IV p1-1-2, r1-1-2, v2-2 -2.

Eye size and interdistances: AME 0.14, ALE 0.12, PME 0.10, PLE 0.10; AME–AME 0.16, AME–ALE 0.14, PME–PME 0.22, PME–PLE 0.16, ALE–PLE 0.14; MOA 0.26 long, anterior width 0.36, posterior width 0.34.

Genitalia ( Figs 36–37 ). Epigynal atrium deep, situated posteriorly. Copulatory orifices oval, situated on anterior margin of epigynal atrium. Anterior bursae sclerotized, elongated, widest posteriorly, distinctly narrowed anteriorly. Spermathecae enlarged, more or less spherical. Fertilization ducts acuminated.

Natural history. Types of S. acuminata sp. nov. were collected from plantation and forests not far from the coast.

Distribution. Southern Thailand and Indonesia (Fig. 43).

Figures

FIGURES 32 – 37. Systaria acuminata sp. nov., holotype male (33 – 34), paratype male (35) and paratype female (36 – 37). 32 – 34 Left male palp (32 ventral, 33 retrolateral, 34 dorsal); 35 Palpal tibia, retrolateral; 36 Epigyne, ventral; 37 Vulva, dorsal. Scale lines = 0.1 mm.