Systaria insolita, Dankittipakul, Pakawin & Singtripop, Tippawan, 2011

Dankittipakul, Pakawin & Singtripop, Tippawan, 2011, Seven new species of Systaria Simon, 1897 from Southeast Asia (Araneae, Clubionidae, Systariinae), Zootaxa 2905, pp. 16-32 : 26-28

Systaria insolita sp. nov.

Figs 26–31

Type material. Holotype: 3, THAILAND: Chaiyaphum Province: Tard Tone National Park, 300 m, 8 December 2006, leg. P. Dankittipakul ( MHNG, Sys-04).

Paratypes: THAILAND: Phitsanulok Province: 2Ƥ, Nakhon Thai District, Phu Hin Rong Kla National Park, 800 m, rotten log, 20 November 2002, leg. P. Dankittipakul ( MHNG, Sys-05).

Diagnosis. Males of S. insolita sp. nov. can be recognized by a peculiar shape of the embolus which is rather short and stout ( Fig. 26 ) instead of elongate and filiform as in other males of Systaria , and by the beak-shaped dorsal process on the RTA ( Figs 27–29 ). Females can be recognized by a pair of hoods on epigynal atrium ( Fig. 30 ), and by the peculiar shape of the anterior bursae ( Fig. 31 ).

Etymology. The specific epithet is a Latin adjective (insolitus,-a, -um), referring to an unusual form of the embolus.

Description. Male (holotype): Total length 7.78. Prosoma 3.54 long, 2.42 wide. Opisthosoma 4.24 long, 2.02 wide. Leg formula: 4123; I 11.12 (2.90, 1.54, 2.80, 2.50, 1.38); II 9.38 (2.74, 1.28, 2.56, 1.80, 1.00); III 8.08 (2.22, 0.84, 1.86, 2.24, 0.92); IV 11.88 (3.36, 1.04, 3.16, 3.20, 1.12). Spination. Femora I d1-1-1, p1-1-1, r0-0-1; II d1-1- 1, p1-1-1, r1-0-1; III d1-1-1, p0-1-1, r1-0-1; IV d1-1-1, p0-1-1, r1-1-1. Tibiae: I v2-2 -2; II v2-2 -1; III d0-1-0, p1-1- 1, r2-2-2, v2-2 -2; IV d0-1-0, p1-1-0, r1-1-0, v2-2 -2. Metatarsi: I v1 -0-0; II v0-1-1; III p1-1-2, r1-1-1, v1 -0-1; IV p1-1-1, r2-2-2, v1-1 -1. Eye size and interdistances: AME 0.12, ALE 0.08, PME 0.08, PLE 0.06; AME–AME 0.16, AME–ALE 0.14, PME–PME 0.22, PME–PLE 0.16, ALE–PLE 0.12; MOA 0.28 long, anterior width 0.30, posterior width 0.28.

Palp ( Figs 26–29 ). Ventral surface of tibia with small, elevated hump ( Fig. 27 ). RTA elongated; ventral process hook-shaped, directed anteriad ( Figs 27, 29 ); dorsal process sharply pointed, directed posteriad ( Figs 26, 29 ). Cymbial fold partially membranous. Cymbial ridge slightly elevated. Embolus triangular, its apex short and stout; embolic base membranous. Conductor spoon-shaped.

Female (paratype, MHNG, Sys-05): Total length 10.58. Prosoma 4.82 long, 3.40 wide. Opisthosoma 5.76 long, 3.12 wide. Leg formula: 4123; I 9.48 (2.88, 1.38, 2.60, 1.70, 0.92); II 8.78 (2.66, 1.28, 2.32, 1.64, 0.88); III 8.32 (2.90, 0.88, 1.82, 1.94, 0.78); IV 12.70 (3.60, 1.32, 2.86, 3.32, 1.60). Spination. Femora: I d1-1-1, p1-1-0, r0-1-1; II d1-1-1, p0-1-1, r0-1-2; III d1-1-1, p1-0-1, r1-2; IV d1-1-1, p1-0-1, r0-0-1. Tibiae: I v2-2 -0; II v0-2-0; III p1-1-0, r0- 1-1, v2-2 -2; IV p1-1-0, r1-1-0, v2-2 -2. Metatarsi: II v1-1 -0; III p1-1-1, r1-1-1, v2 -0-2; IV p1-1-2, r1-1-2, v2-2 -2. Eye size and interdistances: AME 0.14, ALE 0.12, PME 0.10, PLE 0.10; AME–AME 0.16, AME–ALE 0.14, PME–PME 0.22, PME–PLE 0.16, ALE–PLE 0.12; MOA 0.26 long, anterior width 0.38, posterior width 0.36.

Genitalia ( Figs 30–31 ). Epigynal atrium deep, with pair of median hoods probably functioning as guiding ridges. Copulatory orifices situated on posterior margin of epigynal atrium. Anterior bursae sclerotized, elongated, widest posteriorly, distinctly narrowed anteriorly. Spermathecae elongate-oval. Fertilization ducts acuminated.

Natural history. Types of S. insolita sp. nov. were collected from deciduous forests. The females built retreats beneath bark of a large rotten pine tree.

Distribution. Northeastern Thailand (Fig. 43).

Figures

FIGURES 26 – 31. Systaria insolita sp. nov., holotype male (26 – 29) and paratype female (30 – 31). 26 – 27 Left male palp (26 ventral, 27 retrolateral); 28 – 29 Apex of retrolateral tibial apophysis (28 dorsal, 29 prolateral); 30 Epigyne, ventral; 31 Vulva, dorsal. Abbreviations: DP, dorsal process of RTA; VP, ventral process of RTA. Scale lines = 0.1 mm.