Strenorhagio grimaldi, Zhang & Yang & Ren & Shih, 2010, Zhang & Yang & Ren & Shih, 2010

Zhang, Kuiyan, Yang, Ding, Ren, Dong & Shih, Chungkun, 2010, An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera), Zoological Journal of the Linnean Society 158 (3), pp. 563-572 : 567-568

STRENORHAGIO GRIMALDI ZHANG, REN & SHIH GEN. ET SP. NOV. ( FIG. 4 )

Etymology: The specific name is dedicated to the eminent American palaeoentomologist: David Grimaldi.

Diagnosis: Hind femur and tibia slender. Crossvein r–m far proximal to beginning of vein R 2+3; basal trunk of veins M 1 and M 2 distinctly shorter than crossvein m–m; vein CuA 1 arising from cell bm, crossvein m–cu short; mouths of cells sc and r 1 subequal in width; mouth of cell m 1 slightly narrower than that of cell m 2; mouth of cell m 3 narrower than that of cell m 1.

Holotype: Part and counterpart of CNU-DIB- NN2007019P, and CNU-DIB-NN2007019C, an almost complete adult body with wings in dorsal view.

Type locality and horizon: Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China; Jiulongshan Formation, the Middle Jurassic (Aalenian–Bajocian).

Description: Body length, 17.0 mm; wing length, 13.7 mm; wing width, 4.2 mm.

Body stout. Head round, slightly narrower than thorax. Eyes large, bare; ommatidia clearly visible, with upper area composed of large facets.

Hind legs pubescent, femur and tibia slender. Wing hyaline; veins thick and strong. Vein Sc ending slightly beyond middle of wing. Veins R 1 and Rs 1 long; vein R 2+3 distinctly and strongly bent upwards at base; vein R 5 ending slightly beyond wing apex. Crossvein r–m at basal third of cell d, proximal to beginning of vein R 2+3; relative lengths of veins Rs 1, Rs 2 and Rs 3 about 6: 1: 4.5. All branches of vein M long, veins M 1 and M 2 bifurcating distal to crossvein m–m, basal trunk of veins M 1 and M 2 shorter than crossvein m–m. Vein CuA 1 arising from cell bm, crossvein m–cu short; vein CuP present. Mouths of cells sc and r 1, wide, but subequal in width; mouths of cells r 2+3 and r 4, distinctly narrower. Cell bm about as wide as cell br; cell d slender, far from wing margin. Five posterior cells present and wide open; mouth of cell m 1 slightly narrower than that of cell m 2; mouth of cell m 3 distinctly narrower than that of cell m 1, about 1/2 as wide as that of cell cua 1. Cell cup distinctly open; anal lobe small and narrow.

Abdomen cylindrical, robust, and pubescent. Eight segments visible; segment I evidently shortest.

Remarks: Strenorhagio grimaldi sp. nov. looks, more or less, like S. deviatus sp. nov. However, we can separate it from S. deviatus sp. nov. by the following features: basal trunk of veins M 1 and M 2 distinctly shorter than crossvein m–m; vein CuA 1 arising from cell bm; crossvein m–cu short. In S. deviatus sp. nov., the basal trunk of veins M 1 and M 2 is longer than crossvein m–m; vein CuA 1 arises from the infall of cells bm and d.

Figures

Figure 4. Strenorhagio grimaldi sp. nov. Holotype, body with wings in dorsal view. A, photograph. B, camera lucida drawing.