Penestomidae, Simon, 1903

Lopardo, Lara & Hormiga, Gustavo, 2015, Out of the twilight zone: phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea), Zoological Journal of the Linnean Society 173 (3), pp. 527-786 : 725-727

publication ID 10.1111/zoj.12199

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family Penestomidae   (NEW RANK), generic phylogeny of

Eresidae   , and asymmetric rates of change in spinning organ evolution ( Araneae, Araneoidea, Entelegynae   ). Molecular Phylogenetics and Evolution 55: 786–804.

Miller JA, Griswold CE, Yin C. 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae: Araneoidea): systematics and diversity of micro-orbweavers. ZooKeys 11: 9–195.

Millidge AF. 1984. The taxonomy of the Linyphiidae, based chiefly on the epigynal and tracheal characters (Araneae: Linyphiidae). Bulletin of the British Arachnological Society 6: 229–267.

Müller H-G. 1987. Spiders from Colombia V. A new Mysmenopsis from the Ciénaga Grande de Santa Marta, northern Colombia (Araneida: Mysmenidae). Bulletin of the British Arachnological Society 7: 185.

Namkung J. 2002. The spiders of Korea, 1st edn. Seoul: Kyo-Hak Publishing Co.

Namkung J. 2003. The spiders of Korea, 2nd edn. Seoul: Kyo-Hak Publishing Co.

Namkung J, Lee K. 1987. A new record spider of the genus Mysmenella Brignoli, 1980 (Araneae: Mysmenidae) from Korea. Korean Arachnology 3: 45–49.

Nixon K. 1999 a. The parsimony ratchet, a new method for rapid parsimony analysis. Cladistics 15: 407–414.

Nixon K. 1999 b. WINCLADA, program and documentation. Available at:

Ono H. 2007. Eight new species of the families Hahniidae, Theridiidae, Linyphiidae and Anapidae (Arachida, Araneae) from Japan. Bulletin of the National Science Museum, Tokyo 33: 153–173.

Ono H, Chang Y, Tso I. 2007. Three new spiders of the families Theridiidae and Anapidae (Araneae) from southern Taiwan. Memoirs of the National Science Museum, Tokyo 44: 71–82.

Penney D. 2000. Miocene spiders in Dominican amber (Oonopidae, Mysmenidae). Palaeontology 43: 343–357.

Peters H. 1982. Wie spinnen der Familie Uloboridae ihre beute einspinnen und verzehren. Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg 25: 147–167.

Petrunkevitch A. 1928. Systema Aranearum. Transactions of the Connecticut Academy of Arts and Sciences 29: 1–270.

Petrunkevitch A. 1971. Chiapas amber spiders, II. University of California Publications in Entomology 63: 1–44.

Platnick NI. 2014. The world spider catalog, version 14.5. American Museum of Natural History. Available at: http://

Platnick NI, Coddington J, Forster R, Griswold CE. 1991. Spinneret Morphology and the Phylogeny of Haplogyne Spiders (Araneae, Araneomorphae). American Museum Novitates 3016: 1–73.

Platnick NI, Forster R. 1986. On Teutoniella, an American Genus of the Spider Family Micropholcommatidae (Araneae, Palpimanoidea). American Museum Novitates 2854: 1–23.

Platnick NI, Forster R. 1987. On the first American spiders of the subfamily Sternodinae (Araneae, Malkaridae). American Museum Novitates 2894: 1–12.

Platnick NI, Forster R. 1989. A revision of the temperate South American and Australasian spiders of the family Anapidae (Araneae, Araneoidea). Bulletin of the American Museum of Natural History 190: 1–139.

Platnick NI, Shadab M. 1978. A review of the spider genus Mysmenopsis (Araneae, Mysmenidae). American Museum Novitates 2661: 1–22.

Platnick NI, Shadab MU. 1979. A review of the spider genera Anapisona and Pseudanapis (Araneae, Anapidae). American Museum Novitates 2672: 1–20.

Prendini L. 2001. Species or supraspecific taxa as terminals in cladistic analysis? Groundplans versus exemplars revisited. Systematic Biology 50: 290–300.

Purcell W. 1909. Development and origin of respiratory organs in Araneae. Quarterly Journal of Microscopical Science 54: 1–110.

Rae T. 1998. The logical basis for the use of continuous characters in phylogenetic systematics. Cladistics 14: 221–228.

Ramírez MJ. 2000. Respiratory system morphology and the phylogeny of haplogyne spiders (Araneae, Araneomorphae). Journal of Arachnology 28: 149–157.

Ramírez MJ, Lopardo L, Platnick NI. 2004. Notes on Chilean anapids and their webs (Araneae, Anapidae). American Museum Novitates 3428: 1–13.

Rix MG. 2008. A new species of Micropholcomma (Araneae: Araneoidea: Micropholcommatidae) from Western Australia. Records of the Western Australian Museum 24: 343– 348.

Rix MG, Harvey MS. 2010. The spider family Micropholcommatidae (Arachnida, Araneae, Araneoidea): a relimitation and revision at the generic level. ZooKeys 36: 1–321.

Rix MG, Harvey MS, Roberts J. 2008. Molecular phylogenetics of the spider family Micropholcommatidae (Arachnida: Araneae) using nuclear rRNA genes (18S and 28S). Molecular Phylogenetics and Evolution 46: 1031–1048.

Rix MG, Harvey MS, Roberts JD. 2010. A revision of the tricellin spider genus Raveniella (Araneae: Araneoidea: Micropholcommatidae): exploring patterns of phylogeny and biogeography in an Australian biodiversity hotspot. Invertebrate Systematics 24: 209–237.

Roberts M. 1978. Contributions à l’étude de la faune terrestre des îles granitiques de l’archipel des Séchelles (Mission PL G Benoit – J J Van Mol 1972) Theridiidae, Mysmenidae and gen Theridiosoma (Araneidae) (Araneae). Revue de Zoologie Africaine 92: 902–939.

Saaristo M. 1978. Spiders (Arachnida, Araneae) from the Seychelle islands, with notes on taxonomy. Annales Zoologici Fennici 15: 99–126.

Scharff N, Coddington J. 1997. A phylogenetic analysis of the orb-weaving spider family Araneidae (Arachnida, Araneae). Zoological Journal of the Linnean Society 120: 355– 434.

Schütt K. 2000. The limits of the Araneoidea (Arachnida: Araneae). Australian Journal of Zoology 48: 135–153.

Schütt K. 2002. The limits and phylogeny of the Araneoidea (Arachnida, Araneae). Unpublished D. Phil. Thesis, University of Berlin.

Schütt K. 2003. Phylogeny of Symphytognathidae s.l. (Araneae, Araneoidea). Zoologica Scripta 32: 129–151.

Shinkai E. 1977. Spiders of Tokyo III. Acta Arachnologica, Tokyo 27: 321–336.

Sierwald P. 1989. Morphology and Ontogeny of Female Copulatory Organs in American Pisauridae, with Special Reference to Homologous Features (Arachnida: Araneae). Smithsonian Contributions to Zoology 484: 1–24.

Simon E. 1879. Arachnides nouveau de France, d’Espagne et d’Algérie. Premier mémoire. Bulletin de la Société Zoologique de France 4: 251–263.

Simon E. 1880. Matériaux pour servir à une faun arachnologique de la Nouvelle Calédonie. Annales de la Société Entomologique de Belgique 23 (C.R.): 164–175.

Simon E. 1881. Les arachnides de France. Paris 5: 1–180.

Simon E. 1893. Histoire naturelle das araignées. Paris 1: 257– 488.

Simon E. 1894. Histoire naturelle des araignées. Paris. 1: 489–760.

Simon E. 1895 a. Histoire naturelle des araignées. Paris. 1: 761–1084.

Simon E. 1895 b. Etudes arachnologiques. 26e XLI Descriptions d’espèces et de genres nouveaux de l’ordre des Araneae. Annales de la Societe Entomologique de France 64: 131– 160.

Simon E. 1897. On the spiders of the island of St Vincent, III. Proceedings of the Zoological Society of London 1897: 860– 890.

Simon E. 1899. Contribution à la faune de Sumatra. Arachnides recueillis par M J L Weyers, à Sumatra (Deuxiéme mémoire). Annales de la Societe Entomologique de Belgique 43: 78– 125.

Simon E. 1903. Descriptions d’arachnides nouveaux. Annales de la Société Entomologique de Belgique 47: 21–39.

Simon E. 1922. Description de deux arachnides cavernicoles du midi de la France. Bulletin de la Societe Entomologique de France 1922: 199–200.

Simon E. 1926. Les arachnides de France. Synopsis générale et catalogue des espèces françaises de l’ordre des Araneae; 2e partie. Paris. 6: 309–532.

Snazell R. 1986. A new Mysmena (Araneae: Mysmenidae) from Spain. Bulletin of the British Arachnological Society 7: 62–64.

Strong E, Lipscomb D. 1999. Character coding and inapplicable data. Cladistics 15: 363–371.

Sundevall CJ. 1833. Conspectus Arachnidum. Londini Gothorum, 1–39.

Thaler K. 1975. Trogloneta granulum Simon, eine weitere Reliktart der Nordostalpen (Arachnida, Aranei, ‘Symphytognathidae’). Revue Suisse De Zoologie 82: 283– 291.

Thaler K. 1995. Mysmeniola spinifera n.gen. n.sp., eine merkwürdige Kleinspinne aus Venezuela (Arachnida, Araneae: Mysmenidae). Mitteilungen der Schweizerischen Entomologischen Gesellschaft 68: 429–433.

Thaler K, Noflatscher M-T. 1990. Neue und bemerkenswerte Spinnenfunde in Südtirol (Arachnida: Aranei). Veröffentlichungen des Museum Ferdinandeum in Innsbruck 69: 169–190.

Thiele K. 1993. The Holy Grail of the perfect character: the cladistic treatment of morphometric data. Cladistics 9: 275– 304.

Trotta A. 2005. Introduzione al ragni italiani (Arachnida Araneae). Memorie della Societa Entomologica Italiana 83: 3–178.

Uhl G. 2002. Female genital morphology and sperm priority patterns in spiders (Araneae). In: Toft S, Scharff N, eds. European arachnology 2000. Aarhus: Aarhus University Press, 145–156.

Walckenaer CA. 1841. Histoire naturelle des Insects. Aptères. Paris 2: 1–549.

Wheeler W. 1995. Sequence alignment, parameter sensitivity, and the phylogenetic analysis of molecular data. Systematic Biology 44: 321–331.

Wiehle H. 1963. Über Nesticus borutzkyi Reimoser (Arach., Araneae). Senckenbergiana Biologica 44: 431–435.

Wiehle H. 1967. Meta, -eine semientelegyne Gattung der Araneae (Arach.). Senckenbergiana Biologica 48: 183–196.

Wiens J. 2001. Character analysis in morphological phylogenetics: problems and solutions. Systematic Biology 50: 689–699.

Wood HM, Griswold CE, Gillespie RG. 2012. Phylogenetic placement of pelican spiders (Archaeidae, Araneae), with insight into evolution of the ‘neck’ and predatory behaviors of the superfamily Palpimanoidea. Cladistics 28: 598– 626.

Wood HM, Matzke NJ, Gillespie RG, Griswold CE. 2013. Treating fossils as terminal taxa in divergence time estimation reveals ancient vicariance patterns in the palpimanoid spiders. Systematic Biology 62: 264–284.

Wunderlich J. 1978. Zu Taxonomie und Synonymie der Taxa Hadrotarsidae, Lucarachne Bryant 1940 und Flegia CL Koch & Berendt 1854 (Arachnida: Araneida: Theridiidae). Zoologische Beiträge 24: 25–31.

Wunderlich J. 1980 a. Sternal-organe der Theridiosomatidae – eine bisher iibersehene Autapomorphie (Arachnida: Araneae). Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg 23: 255–257.

Wunderlich J. 1980 b. Über europäische Symphytognathidae (Arach: Araneae). Verhandlungen des Naturwissenschaftlichen Vereins in Hamburg 23: 259–273.

Wunderlich J. 1986. Die Beziehungen fossiler und rezenter Radnetzspinnen-Verwandter (überfamilie Araneoidea. In: Meyer Q, ed. Spinnenfauna gestern und heute: Fossile Spinnen in Bernstein und ihre heute lebenden Verwandten. Wiesbaden: Quelle & Meyer, 93–135.

Wunderlich J. 1987. Die Spinnen der Kanarischen Inseln und Madeiras: adaptive radiation, biogeographie, revisionen und neubeschreibungen. Langen: Verlag T.

Wunderlich J. 1995. Drei bisher unbekannte Arten und Gattungen der Familie Anapidae (s.l.) aus Süd-Afrika, Brasilien und Malaysia (Arachnida: Araneae). Beiträge zur Araneologie 4: 543–551.

Wunderlich J. 1998. Beschreibung der ersten fossilen Spinnen der Unterfamilien Mysmeninae (Anapidae) und Erigoninae (Linyphiidae) im Dominikanischen Bernstein (Arachnida: Araneae). Entomologische Zeitschrift, Frankfurt am Main 108: 363–367.

Wunderlich J. 2004. The fossil spiders of the family Anapidae s. l. (Aeaneae) [sic] in Baltic, Dominican and Mexican amber and their extant relatives, with the description of the new subfamily Comarominae. Beiträge zur Araneologie 3: 1020– 1111.

Wunderlich J. 2008. On extant and fossil (Eocene) European comb-footed spiders (Araneae: Theridiidae), with notes on their subfamilies, and with descriptions of new taxa. Beiträge zur Araneologie 5: 140–469.

Wunderlich J. 2011. Some subrecent spiders (Araneae) in copal from Madagascar. Beiträge zur Araneologie 6: 445–460.

Yin C, Peng X, Bao Y. 2004. A new species of the genus Mysmenella from China (Araneae, Mysmenidae). Acta Zootaxonomica Sinica 29: 80–82.