Maymena, GERTSCH, 1960

Lopardo, Lara & Hormiga, Gustavo, 2015, Out of the twilight zone: phylogeny and evolutionary morphology of the orb-weaving spider family Mysmenidae, with a focus on spinneret spigot morphology in symphytognathoids (Araneae, Araneoidea), Zoological Journal of the Linnean Society 173 (3), pp. 527-786 : 778

publication ID

https://doi.org/ 10.1111/zoj.12199

DOI

https://doi.org/10.5281/zenodo.7004799

persistent identifier

https://treatment.plazi.org/id/03832D77-10BD-939B-FE81-E262FBB75B21

treatment provided by

Felipe

scientific name

Maymena
status

 

MAYMENA GERTSCH, 1960 View in CoL View at ENA

( FIGS 10–16 View Figure 10 View Figure 11 View Figure 12 View Figure 13 View Figure 14 View Figure 15 View Figure 16 , 128B, D, E View Figure 128 , 131I, J View Figure 131 , 140M–O View Figure 140 , 141A–I View Figure 141 , 147D,E View Figure 147 : CLADE C168)

Maymena Gertsch, 1960a: 30–38 View in CoL . Gertsch, 1971: 94–95. Brignoli, 1974: 224. Baert, 1990: 17. Platnick, in Eberhard, Platnick & Schuh, 1993: 11. Griswold et al., 1998: 63.

Type species

Maymena mayana (Chamberlin & Ivie, 1938) View in CoL by original designation, type material in MCZ, not examined.

Familial placement and composition

Transferred to Mysmenidae from Symphytognathidae by Forster & Platnick (1977). Maymena is closely related to Mysmenopsinae in our working phylogenetic hypothesis ( Fig. 161B View Figure 161 ). Currently, Maymena comprises 13 described species (Platnick, 2014), and it is represented here by three described plus three undescribed species (the latter species scored only for molecular data): M. mayana , M. ambita , M. rica , Maymena -MYSM-003- ARG, Maymena -MYSM-004- MEX, and Maymena - MYSM-016-ARG).

Monophyly

Morphological synapomorphies of Maymena include: posterior lateral spinnerets with an anterior flat spatulate modified seta ( Fig. 11H View Figure 11 ); aciniform gland spigots of the posterior spinnerets of two different shapes ( Fig. 13D View Figure 13 ); seta on major ampullate field with one row of long ‘branches’ ( Figs 11E View Figure 11 , 13C View Figure 13 , 16B View Figure 16 ); leg spination (i.e. macrosetae occurring on tibiae, femora, and metatarsi; Figs 140M View Figure 140 , 141C View Figure 141 ); and males with cylindrical palpal tibia ( Fig. 10A View Figure 10 ). Ambiguously optimized synapomorphies for Maymena include: males with a femoral spot at least on femur I; metatarsal clasping spine particularly proximal ( Fig. 16G View Figure 16 ); embolus rim deeply grooved ( Fig. 10H View Figure 10 ); the primary cymbial conductor apically bent over the ventral side ( Fig. 10D, G View Figure 10 ); epiandrous fusules dispersed in a row ( Figs 12B View Figure 12 , 16A View Figure 16 ); and macrosetae on female palpal tarsus ( Figs 13A View Figure 13 , 15A View Figure 15 ). Maymena is one of the few clades in this data set that is strongly supported and stable, and its monophyly is supported by most data partitions and parameter combinations. The genus is also supported by 125 molecular synapomorphies.

Diagnosis

Maymena shares with Mysmeninae the anterior flat spatulate modified seta on PLS and the aciniform gland spigots of two different shapes; with Trogloneta the presence of femoral spot on males; and with Mysmenopsinae the dispersed male epiandrous fusules, the macrosetae on female palpal tarsus, the leg spination, and the absence of tegular conductor (see also Gertsch, 1960a). Maymena differs from all other mysmenid genera by the combination of the aforementioned features, and by the presence of one row of long ‘branches’ on the major ampullate field seta, males with cylindrical palpal tibia, metatarsal clasping spine particularly proximal, seemingly interacting with the particularly distal tibial clasping spine in most species, and the male palpal morphology, with primary cymbial conductor apically bent over the ventral side. In addition, the respiratory arrangement distinguishes Maymena from other mysmenids. It consists of anterior booklungs and two long lateral tracheal tubes and two shorter median apodemes arising from a narrow posterior single spiracle (L. Lopardo, P. Michalik & G. Hormiga, unpubl. data; also Gertsch, 1960a).

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Mysmenidae

Loc

Maymena

Lopardo, Lara & Hormiga, Gustavo 2015
2015
Loc

Maymena

Gertsch, 1971: 94–95
Brignoli, 1974: 224
Baert, 1990: 17
Platnick, in Eberhard, Platnick & Schuh, 1993: 11
Griswold et al., 1998: 63
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