Agathidium cheneyi Miller and Wheeler, 2005

MILLER, KELLY B. & WHEELER, QUENTIN D., 2005, Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part Ii. Coleoptera: Leiodidae, Bulletin of the American Museum of Natural History 2005 (291), pp. 1-167 : 121

Agathidium cheneyi Miller and Wheeler ,

new species Figures 148 , 273–276 , 366

TYPE MATERIAL: Holotype, Ƌ in CMNC labeled ‘‘ MEXICO: Chiapas: Yerbabuena Preserve 2.1 km NW Pueblo Nuevo, Solistahuacan. R.S. Anderson 92­114 23­IX­ 1992 / HOLOTYPE Agathidium cheneyi Mill­ er and Wheeler, 2003 [red label with black line border]’’. The holotype is the only specimen examined of this species.

TYPE LOCALITY: Mexico, Chiapas, Yerbabuena Preserve, 2.1 km NW Pueblo Nuevo, Solistahuacan.

DIAGNOSIS: This species is similar to A. aztec and related species. This species differs in the greater development of the oblique metasternal carinae which meet medially in a very broad, truncate flange. The median lobe differs particularly in having the operculum very short and laterally rounded with a distinct apical emargination (fig. 273).

DESCRIPTION: Body moderately small (TBL = 3.26 mm), broad, robust, rounded (PNW/TBL = 0.47), strongly contractile.

Head and pronotum red; elytra red, iridescent; venter yellow­red; antennae, palpi, and legs yellow­brown.

Head broad (MDL/OHW = 0.58), dorsal surface flattened, dorsoventrally compressed; with very fine punctures, each with a short, very fine seta, surface between punctures shiny, very lightly microreticulate; frontoclypeal suture obsolete medially; eyes moderately large, slightly compressed dorsoventrally; gula flat; antennomere ratios: length I: II:III = 1.6:1.0:1.5, width VII:VIII:IX = 1.0: 1.0:2.1. Pronotum very large, broad (PNL/ PNW = 0.71), strongly convex, anterolateral lobes strongly produced, lateral margin broadly curved, not angulate; with fine, sparse punctures, each with a short, very fine seta, surface between punctures shiny, smooth. Elytra broad, lateral margins strongly rounded, apically rounded (SEL/ELW = 1.07); punctation and surface similar to pronotum; sutural stria present in apical one­fifth of elytron. Flight wings strongly reduced. Mesosternum moderately broad, not declivitous; medial carina well developed. Metasternum narrow (MTL/MTW = 0.16), flat medially, distinctly dorsally sloped anteriorly; oblique femoral carinae moderately well developed, meeting medially in broadly truncate carina.

Male tarsi 5­5­4; pro­ and mesobasotarsomeres somewhat laterally expanded, with moderately large ventral field of spatulate setae; mandibles not modified; metafemur moderately broad, with small tooth subapically on posterior margin (fig. 148); metasternal fovea large, transversely oval with large region of long fine setae. Median lobe in lateral aspect slender, strongly curved basally, with moderate lateral and dorsal carinae at base of apical portion for reception of lateral lobe, apical portion slender, sinuate, apex very narrowly truncate (fig. 275); in ventral aspect slender, expanded at base of apical portion that has the lateral margins slightly sinuate and convergent to near apex, which is slightly expanded and rounded (figs. 273, 274); operculum broad, short, flat, with shallow V­shaped emargination apically (fig. 273); lateral lobes slender, apically sinuate, apex narrowly rounded with 2 stout setae (fig. 276).

Female not examined.

ETYMOLOGY: This species is named in hon­ or of R.B. Cheney (Casper, WY), Vice­President of the United States.

DISTRIBUTION: This species is known only from Chiapas (fig. 366).

Figures

Figs. 145–172. Agathidium oniscoides­group species, left male metafemur, ventral aspect: 145, A. oculeum. 146, A. recurvatum. 147, A. impensum. 148, A. cheneyi. 149, A. tenangoense. 150, A. rumsfeldi. 151, A. hirsutum. 152, A. cortezi. 153, A. tumidiventre. 154, A. aztec. 155, A. iridescens. 156, A. dentigerum. 157, A. stephani. 158, A. akrogeneios. 159, A. appalachium. 160, A. pocahontasae. 161, A. carolinense. 162, A. gallititillo. 163, A. divaricatum. 164, A. bushi. 165, A. georgiaense. 166, A. chauliodoum. 167, A. microphthalmum. 168, A. nimbosilva. 169, A. oniscoides. 170, A. rubellum. 171, A.

Figs. 261–287. Agathidium oniscoides­group species, aedeagus: 261–263, A. oaxacaense: 261, ventral; 262, median lobe, lateral; 263, left lateral lobe, lateral. 264–266, A. oculeum: 264, ventral; 265, median lobe, lateral; 266, left lateral lobe, lateral. 267–269, A. recurvatum: 267, ventral; 268, median lobe, lateral; 269, left lateral lobe, lateral. 270–272, A. impensum: 270, ventral; 271, median lobe, lateral; 272, left lateral lobe, lateral. 273–276, A. cheneyi: 273, ventral; 274, median lobe, apex, ventral; 275, median lobe, lateral; 276, left lateral lobe, lateral. 277–279, A. tenangoense: 277, ventral; 278, median lobe, lateral; 279, left lateral lobe, lateral. 280–283, A. rumsfeldi: 280, ventral; 281, median lobe, apex, ventral; 282, median lobe, lateral; 283, left lateral lobe, lateral. 284–287, A. hirsutum: 284, ventral; 285, median lobe, apex, ventral; 286, median lobe, lateral; 287, left lateral lobe, lateral.

Figs. 365, 366. Geographic distribution of Agathidium oniscoides­group species: 365, A. andersoni = -; A. disgregum = ^; A. invisitatum = •; A. oaxacaense = *. 366, A. impensum = *; A. oculeum = •; A. recurvatum = -; A. cheneyi = ^.