Agathidium rumsfeldi Miller and Wheeler, 2005

MILLER, KELLY B. & WHEELER, QUENTIN D., 2005, Slime-Mold Beetles Of The Genus Agathidium Panzer In North And Central America, Part Ii. Coleoptera: Leiodidae, Bulletin of the American Museum of Natural History 2005 (291), pp. 1-167 : 123-124

Agathidium rumsfeldi Miller and Wheeler ,

new species Figures 150 , 280–283 , 367

TYPE MATERIAL: Holotype, Ƌ in FMNH labeled ‘‘ MEXICO: Hidalgo 2.5 mi N Tlanchinol, 5200 ft vii.6–11.1973/white polypore tree fungus A. Newton/ HOLOTYPE Agathidium rumsfeldi Miller and Wheeler, 2003 [red label with black line border]’’.

TYPE LOCALITY: Mexico, Hidalgo, 2.5 mi N Tlanchinol, 5200̍.

DIAGNOSIS: This species is extremely similar to A. aztec and related species. The elytra are slightly iridescent. This species can be distinguished by the shape of the median lobe which has the operculum with a small, apical V­shaped emargination and each ramus flared laterally and distinctly pointed (fig. 280). Other differences are only slight shape differences in the median lobe (figs. 280–282).

DESCRIPTION: Body moderately small (TBL = 2.73–3.03 mm), broad, robust, rounded (PNW/TBL = 0.45–0.49), strongly contractile.

Head and pronotum red; elytra red, slightly iridescent; venter red­yellow; antennae, palpi, and legs yellow.

Head broad (MDL/OHW = 0.61–0.66), dorsal surface flattened, dorsoventrally compressed; with very fine punctures, each with a short, very fine seta, surface between punctures shiny, very lightly microreticulate; frontoclypeal suture obsolete medially; eyes not reduced, not dorsoventrally compressed; gula flat; antennomere ratios: length I:II:III = 1.6:1.0:1.3, width VII:VIII:IX = 1.0:1.0: 2.3. Pronotum very large, broad (PNL/PNW = 0.67–0.76), strongly convex, anterolateral lobes strongly produced, lateral margin broadly curved, not angulate; with fine, sparse punctures, each with a short, very fine seta, surface between punctures shiny, smooth. Elytra broad, lateral margins strongly rounded, apically rounded (SEL/ELW = 1.01–1.05); punctation and surface similar to pronotum; sutural stria present only at apex of elytron. Flight wings present, fully developed. Mesosternum moderately broad, not declivitous; medial carina well developed. Metasternum narrow (MTL/MTW = 0.16), flat medially, distinctly dorsally sloped anteriorly; oblique femoral carinae low, not prominent where meeting medially.

Male tarsi 5­5­4; pro­ and mesobasotarsomeres slightly laterally expanded, with small ventral field of spatulate setae; mandibles not modified; metafemur moderately broad, with small tooth subapically along posterior margin (fig. 150); metasternal fovea moderately large, oval with large brush of fine, long setae. Median lobe in lateral aspect slender, basally strongly curved, apical portion curved, slender, apically pointed, slightly expanded subapically, with lateral and dorsal carinae at base of apical portion between which fit the lateral lobes (fig. 282); in ventral aspect slender, at base of apical portion with prominent lateral lobes, apical portion evenly tapered for most of length, subapically slender, apex expanded (figs. 280, 281); operculum in lateral aspect curved ventrad, slender (fig. 282), in ventral aspect moderately broad, apex medially with Vshaped, shallow emargination, apices of rami pointed and divergent (fig. 280); lateral lobes moderately slender, curved basally, sinuate apically, medially abruptly expanded, apex expanded, rounded with 2 stout setae (fig. 283).

Female not examined.

ETYMOLOGY: Named after D. Rumsfeld (Taos, NM), Secretary of Defense of the United States.

DISTRIBUTION: Agathidium rumsfeldi is known only from Hidalgo and Oaxaca (fig. 367).

PARATYPE: MEXICO: Oaxaca: 27.3 mi N Ixtlan de Juarez, 10 Aug 1973, 9200̍, under oak bark, A Newton (1, FMNH).

DISCUSSION: This species has been collect­ ed from under oak bark at 5200–9200 ft elevation.

Figures

Figs. 145–172. Agathidium oniscoides­group species, left male metafemur, ventral aspect: 145, A. oculeum. 146, A. recurvatum. 147, A. impensum. 148, A. cheneyi. 149, A. tenangoense. 150, A. rumsfeldi. 151, A. hirsutum. 152, A. cortezi. 153, A. tumidiventre. 154, A. aztec. 155, A. iridescens. 156, A. dentigerum. 157, A. stephani. 158, A. akrogeneios. 159, A. appalachium. 160, A. pocahontasae. 161, A. carolinense. 162, A. gallititillo. 163, A. divaricatum. 164, A. bushi. 165, A. georgiaense. 166, A. chauliodoum. 167, A. microphthalmum. 168, A. nimbosilva. 169, A. oniscoides. 170, A. rubellum. 171, A.

Figs. 261–287. Agathidium oniscoides­group species, aedeagus: 261–263, A. oaxacaense: 261, ventral; 262, median lobe, lateral; 263, left lateral lobe, lateral. 264–266, A. oculeum: 264, ventral; 265, median lobe, lateral; 266, left lateral lobe, lateral. 267–269, A. recurvatum: 267, ventral; 268, median lobe, lateral; 269, left lateral lobe, lateral. 270–272, A. impensum: 270, ventral; 271, median lobe, lateral; 272, left lateral lobe, lateral. 273–276, A. cheneyi: 273, ventral; 274, median lobe, apex, ventral; 275, median lobe, lateral; 276, left lateral lobe, lateral. 277–279, A. tenangoense: 277, ventral; 278, median lobe, lateral; 279, left lateral lobe, lateral. 280–283, A. rumsfeldi: 280, ventral; 281, median lobe, apex, ventral; 282, median lobe, lateral; 283, left lateral lobe, lateral. 284–287, A. hirsutum: 284, ventral; 285, median lobe, apex, ventral; 286, median lobe, lateral; 287, left lateral lobe, lateral.

Figs. 367, 368. Geographic distribution of Agathidium oniscoides­group species: 367, A. hirsutum = -; A. tenangoense = •; A. iridescens = ^; A. rumsfeldi = *. 368, A. aztec = ^; A. cortezi = *; A. tumidiventre = •.