Amplirhagada forrestiana, Köhler, 2011

Köhler, Frank, 2011, Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae), Records of the Australian Museum 63 (2), pp. 167-202 : 183-184

publication ID	https://doi.org/ 10.3853/j.0067-1975.63.2011.1581
publication LSID	lsid:zoobank.org:pub:1BCD4085-D2B9-400D-B504-8C85C30303D6
persistent identifier	https://treatment.plazi.org/id/0384503A-FFAB-3C4F-FEA5-FB5BFB24699E
treatment provided by	Felipe
scientific name	Amplirhagada forrestiana
status	sp. nov.

Treatment

Amplirhagada forrestiana View in CoL n.sp.

Type locality. Western Australia, NW Kimberley, York Sound, Prince Frederick Harbour, 9.7 km NW of Mt. Anderdon , 4 km from coast; 14°52'50"S 125°21'30"E (RFS- 05-3; coll. V. Kessner, 05 Jun 1987) ( Fig. 1 View Figure 1 ) GoogleMaps .

Type material. Holotype WAM S34725 View Materials (preserved specimen) (Pl. 1.8; Table 1) . Paratypes AM C.472930 (2 preserved specimens), WAM S34726 View Materials (5 preserved specimens) , FMNH 220084 About FMNH (8 preserved specimens) , AM C.472931 (5 dried shells), WAM S34727 View Materials (9 dried shells) , FMNH 220083 About FMNH (12 dried shells) .

Etymology. Named after Matthew Forrest, first Kimberley explorer, who in 1879 was instructed to map out the country between the De Grey River in Western Australia and the Victoria River in the Northern Territory.

Description

Shell (Pl. 1.8; Fig. 25 A–C View Figure 25 ). Broadly conical with low spire, almost discoid. Thin, translucent. Periphery sharply angulate; upper and lower sectors of whorls rounded. Umbilicus narrowly winding, 30–80% concealed by columellar reflection. Background colour pale brownish horn; peripheral band absent or diffuse, if present moderately broad, diffuse, brown, visible on last whorls only; sub-sutural band absent or diffuse, brown, visible on last whorls only; outer and inner lip colour whitish. Protoconch 4.7 mm in diameter, comprising about 1.5 whorls, with indistinct radially elongated pustulations. Teleoconch with well-developed, regular lirae, evenly distributed across shell surface. Angle of aperture 45–60 degrees; outer lip thin, well rounded, slightly expanded, slightly reflected; basal node absent or weak. Parietal wall of inner lip inconspicuous. Average shell size 11.6±0.8 × 19.4±1.0 mm ( Table 1).

Radular and jaw morphology ( Fig. 25 D–G View Figure 25 ). Tooth formula C + 11 + 4 + 16; 150 rows of teeth (n = 1). Jaw with nine plates.

Genital morphology ( Figs 23 View Figure 23 , 24 View Figure 24 ). Penis rather straight, tubular, longer than anterior part of oviduct. Penial sheath delicate, proximally thicker. Penial of length of penial chamber. Inner penial wall entirely covered by densely in honey-comb pattern arranged, small and regular pustules. No main stimulatory pilaster. Vas deferens rather thin, entering penial sheath close to penial apex, long and winding. Vagina moderately long, distally inflated; inner vaginal wall and wall of bursa copulatrix with smooth longitudinal pilasters. Bursa copulatrix very wide, slightly extending base of spermoviduct. Free oviduct very long, tubular, coiling, comprising ¾ of length of anterior part of oviduct. Spermoviduct about as long as anterior part of oviduct.

Aestivation strategy. Free sealer.

Remarks. Listed by Solem (1991) as “ Amplirhagada NSP 25”. A single specimen from Naturalist Island, assigned to the same manuscript name in the collection of the Field Museum (FMNH 220692) was not studied. Amongst medium-sized species, shell differs from other congeners by markedly wider diameter, resulting in a discoidal shape (shell proportions and size only similar in A. discoidea ). Amplirhagada discoidea differs by having a more angulated shell periphery and a conspicuously different penial anatomy, with a large main pilaster (absent in A. forrestiana ), rather smooth inner penial wall (markedly pustulose in A. forrestiana ) and rather narrow bursa copulatrix (wide on A. forrestiana ).