Amplirhagada carsoniana, Köhler, 2011

Köhler, Frank, 2011, Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae), Records of the Australian Museum 63 (2), pp. 167-202 : 172-173

publication ID	https://doi.org/ 10.3853/j.0067-1975.63.2011.1581
publication LSID	lsid:zoobank.org:pub:1BCD4085-D2B9-400D-B504-8C85C30303D6
persistent identifier	https://treatment.plazi.org/id/79C0B2D0-CEAD-461E-9DB6-717A14B3C5F6
taxon LSID	lsid:zoobank.org:act:79C0B2D0-CEAD-461E-9DB6-717A14B3C5F6
treatment provided by	Felipe
scientific name	Amplirhagada carsoniana
status	sp. nov.

Treatment

Amplirhagada carsoniana View in CoL n.sp.

Type locality. Western Australia, NW Kimberley, Carson Escarpment, 2.5 km N of Face Point , base of escarpment ;

14°50'20"S 126°49'10"E ( RFS 10-4 , coll. V. Kessner, 10 Jun 1987) ( Fig. 1 View Figure 1 ) GoogleMaps .

Type material. Holotype WAM S34707 View Materials (preserved specimen) (Pl. 1.2; Table 1) . Paratypes WAM S34708 View Materials (preserved specimen) , FMNH 220344 About FMNH (preserved specimen) , AM C.472920 (5 dried shells), WAM S34709 View Materials (12 dried shells) , FMNH 220343 About FMNH (17 dried shells) .

Etymology. In reference to the Carson Escarpment, where this species was found.

Description

Shell (Pl. 1.2; Fig. 5 A–C View Figure 5 ). Broadly conical with low spire, almost discoid. Thin to delicate, translucent. Periphery rounded to slightly angulate; upper and basal sectors of whorls well rounded. Umbilicus 10–30% concealed by columellar reflection. Background colour brownish horn; peripheral band absent or well marked, thin, brown, visible on most whorls; sub-sutural diffuse, thin, visible on most whorls; outer and inner lip colour whitish. Protoconch 2.6 mm in diameter, comprising about one whorl, with weak radially elongated pustulations. Teleoconch with faint, regular axial growth lines, evenly distributed across shell surface. Angle of aperture about 45 degrees; outer lip thin, well rounded, expanded, weakly reflected; basal node absent. Parietal wall of inner lip absent or very inconspicuous. Average shell size 9.8±0.6 × 16.3± 0.8 mm ( Table 1).

Radular and jaw morphology ( Fig. 5 D–G View Figure 5 ). Tooth formula C + 12–14 + 4 + 18–20; average number of rows of teeth 130 (n = 2). Jaw with ten plates.

Genital morphology ( Figs 6–7 View Figure 6 View Figure 7 ). Penis coiled within thick penial sheath; penial complex about as long as anterior part of oviduct. Length of penial retractor muscle equivalent to about half of length of penial complex. Penial verge extending about 1/5 of length of penial chamber, slender with pointed tip. Inner penial wall entirely covered by dense pustulation; pustules forming indistinct, corrugated pilasters. Main stimulatory pilaster indistinct, formed by prolonged pustules, comprising proximal half of inner penial chamber. Vas deferens entering penial sheath within proximal third of penial complex. Vagina moderately long, distally inflated. Bursa copulatrix short, reaching base of spermoviduct. Free oviduct coiled, shorter than vagina. Spermoviduct longer than anterior part of oviduct.

Aestivation strategy. Free sealer.

Remarks. Description based on dissection of one specimen. Listed by Solem (1991) as “ Amplirhagada NSP 18” to be distinct from A. drysdaleana Solem, 1981 . Both taxa are indeed very similar but A. carsoniana differs by coiled, shorter penis with shorter main pilaster, no extended basal pilasters of inner penial wall and shorter vagina. Vas deferens entering sheath half way up in A. carsoniana but within upper sector of penial sheath in A. drysdaleana . Being similar overall and occurring in close proximity to each other, both species are likely sister taxa.