Naddia Fauvel, 1867

SMETANA, ALEŠ & DAVIES, ANTHONY, 2000, Reclassification of the North Temperate Taxa Associated with Staphylinus Sensu Lato, Including Comments on Relevant Subtribes of Staphylinini (Coleoptera: Staphylinidae), American Museum Novitates 3287, pp. 1-1 : 1-

Naddia Fauvel, 1867 , 3, 6–11, 16, 17, 19, 22, 38

Nausicotus Sharp, 1884 , 18

nemotoi Naomi, 1992 [ Ocypus (Pseudocypus) ], 44

Neobisnius Ganglbauer, 1895 , 14

neomexicanus (Bernhauer and Schubert, 1914) [= Platydracus sepulchralis (Erichson, 1839) ], 40

Neotasgius J. Müller, 1925 [= Platydracus (Platydracus) ], 24, 39

nepalensis (Scheerpeltz, 1976) [ Platydracus ], 40

nepalicus (Coiffait, 1981) [ Ocypus (Pseudocypus) ], 44

nepalicus (Coiffait, 1982) (Parontholestes) [ Thoracostrongylus ], 23, 39

nero (Faldermann, 1835) [= Ocypus (Matidus) nitens Schrank, 1781 ], 34, 43

nevadensis (J. Müller, 1926) [subsp. of Ocypus (Pseudocypus) picipennis (Fabricius, 1792) ], 45

nigrellus (Horn, 1879) [ Dinothenarus (Parabemus) ], 41

nigrinus (Lucas, 1849) [ Tasgius (Rayacheila) ], 46

nigroaeneus Sharp, 1889 [ Ocypus (Pseudocypus) ], 44

nitens Schrank, 1781 [ Ocypus (Matidus) , and as subsp.], 34, 43

Nordus Blackwelder, 1952 , 17

nubigena Smetana, 1965 [ Ocypus (Matidus) ], 43

Nudabemus Coiffait, 1982 [= Ocypus (Ocypus) ], 31, 32, 43

nudicollis Coiffait, 1977 (Ocypus) [incertae sedis], 47

obscuripes (Bernhauer, 1900) ( Ocypus , rufipes var.) [= Tasgius (Rayacheila) eppelsheimianus (Jakobson, 1909) ], 37

obscuroaeneus Fairmaire, 1852 [ Ocypus (Pseudocypus) , and as subsp.], 44

ochropus (J. Müller, 1950) [subsp. of Ocypus (Matidus) nitens (Schrank, 1781) ], 43

oculatus (Bernhauer, 1929) (Staphylinus) [= Platydracus (Platydracus) oculosus , nom. nov.], 25,40

oculatus (Fabricius, 1775) (Staphylinus) [ Creophilus ], 25

oculatus O. F. Müller, 1776 [ Staphylinus ], 25

oculatus (Sharp, 1874) [ Ontholestes ], 38

oculosus , nom. nov. [for Platydracus oculatus (Bernhauer, 1929) ], 25, 40

Ocypus Leach, 1819 [and subgenus], 2, 3, 6–11, 15–17, 21, 28, 31, 35, 42

ohkurai Hayashi, 1978 [ Agelosus ], 45

ohtsukaorum Naomi, 1992 [ Ocypus (Pseudocypus) ], 44

olens (O. Müller, 1764) (Staphylinus) [ Ocypus (Ocypus) , and as subsp.], 31, 42

olivaceus (Cameron, 1928) [ Abemus ], 23, 39

olympicus (Baudi, 1857) [ Tasgius (Rayacheila) ], 46

Ontholestes Ganglbauer, 1895 , 2, 3, 7–11, 16, 17, 19, 22, 38

opaciceps (Scheerpeltz, 1976) [ Platydracus ], 40

opacus Roth, 1851 (Ocypus) [incertae sedis], 47

ophthalmicus (Scopoli, 1763) (Staphylinus) [ Ocypus (Ocypus) , and as subsp.], 31, 38, 42

orientalis Bernhauer, 1906 [ Ontholestes ], 38

orientalis (Bernhauer and Schubert, 1914) (Staphylinus) [= Ocypus (Pseudocypus) orientis , nom. nov.], 34, 44

orientalis (Motschulsky, 1858) (Staphylinus) [= Creophilus maxillosus (Linne´)], 34

orientis , nom. nov. [for Ocypus (Pseudocypus) orientalis (Bernhauer and Schubert, 1914) ], 34, 44

ormayi (Reitter, 1887) [ Ocypus (Matidus) ], 43

ornaticauda (LeConte, 1863) [ Staphylinus ], 41

ottomanus Fauvel, 1900 [ Ocypus (Matidus) ], 43

Paederinae , 7, 8

paganus Sharp, 1874 [= Platydracus brevicornis (Motschulsky, 1862) ], 39

Parabemus Reitter, 1909 [subgenus of Dinothenarus ], 10, 11, 16, 20, 26, 27, 41

Paralapsodus Coiffait, 1974 [= Tasgius (Rayacheila) ], 36, 37, 46

paramurinus Li and Chen, 1993 [ Ontholestes ], 38

Paraquedius Casey, 1915 (subgen. of Quedius ), 14, 15

Paratasgius Jarrige, 1952 [= Tasgius (Tasgius) ], 36, 45

Paraxenopygus Bernhauer, 1911 , 18

Parocypus Bernhauer, 1915 [= Dinothenarus (Parabemus) ], 26–28, 41

Parontholestes Coiffait, 1982 [= Thoracostrongylus ], 23, 39

parvulus Sharp, 1874 [ Ocypus (Aulacocypus) ], 45

parvulus , nom. nov. [for Platydracus parvus (Cameron, 1932) ], 40

parvus (Cameron, 1932) (Staphylinus) [= Platydracus parvulus , nom. nov.], 40

patricius (Bernhauer, 1915) [ Platydracus (Chaetodracus) ], 25, 41

pecoudi Jarrige, 1954 ( Staphylinus ( Goerius )) [incertae sedis], 47

pedator (Gravenhorst, 1802) (Staphylinus) [ Tasgius (Tasgius) ], 7, 36, 46

pedemontanus (J. Müller, 1924) [ Ocypus (Matidus) , and as subsp.], 43

pendjabensis (Bernhauer, 1915) [ Tasgius (Rayacheila) ], 46

perniger (Scheerpeltz, 1976) [ Platydracus ], 40

pernigrus Coiffait, 1964 [= Ocypus (Ocypus) ophthalmicus atrocyaneus (Fairmaire, 1860)], 42

perreaui Coiffait, 1984 [ Platydracus ], 40

peyerimhoffi (J. Müller, 1926) [ Tasgius (Tasgius) ], 46

Philonthina , 6–12, 13, 14–17

Philonthus Stephens, 1829 , 7­10, 13

Philothalpus Kraatz, 1857 , 17, 18

phoenicurus (Nordmann, 1837) [ Platydracus ], 40

Physetops Mannerheim, 1830 , 2, 7–11, 16, 17, 19, 28, 42

picipennis (Fabricius, 1792) [ Ocypus (Pseudocypus) , and as subsp.], 3, 38, 44, 45

pindensis (Coiffait, 1964) [= Ocypus (Pseudocypus) picipennis picipennis (Fabricius, 1792) ], 45

pinorum (Casey, 1915) [ Platydracus ], 40

plagiicollis (Fairmaire, 1891) [ Ocypus (Pseudocypus) ], 45

planipennis (Aube´, 1842) [ Tasgius (Tasgius) ], 46

Platydracus Thomson, 1858 [and subgenus], 2, 6, 8–11, 15, 16, 18, 22, 24, 25, 38, 39

plebejus (Bernhauer, 1915) [ Platydracus ], 40

pleuralis (LeConte, 1861) [ Dinothenarus (Parabemus) ], 27, 41

pliginskii (Bernhauer, 1915) [ Tasgius (Rayacheila) ], 46

Plociopterus Kraatz, 1857 , 7, 17, 18

Polyphematiana Strand, 1914 , 18

ponticus (Coiffait, 1978) [= Ocypus (Pseudocypus) picipennis picipennis (Fabricius, 1792) ], 45

ponticus Smetana, 1968 [ Ocypus (Matidus) ], 43

praelongus (Mannerheim, 1830) [ Platydracus ], 40

praetorius (Bernhauer, 1915) [ Tasgius (Tasgius) ], 46

prainae (Eppelsheim, 1895) [ Dinothenarus (Parabemus) ], 41

pratti (Scheerpeltz, 1962) [ Platydracus ], 40

primigenius (J. Müller, 1923) [subsp. of Ocypus (Matidus) syriacus Baudi, 1848 ], 43

Protabemus Scheerpeltz, 1966 [= Dinothenarus (Dinothenarus) ], 26, 41

Protocypus J. Müller, 1923 [= Ocypus (Pseudocypus) ], 31, 32, 43

Protogoerius Coiffait, 1956 , 6–11, 16, 17, 21, 30, 42

proximus Kirschenblat, 1936 [ Ontholestes ], 38

pseudoalpestris (J. Müller, 1926) [subsp. of Ocypus (Matidus) brevipennis (Heer, 1839) ], 42

pseudobrevipennis Coiffait, 1964 [= Ocypus (Matidus) chevrolatii (Baudi, 1848) ], 43

Pseudocypus Mulsant and Rey, 1876 [subgenus of Ocypus ], 2, 3, 6–8, 11, 16, 21, 31, 32, 43

pseudolens Coiffait, 1964 [= Ocypus (Ocypus) curtipennis Motschulsky, 1849 ], 42

pseudopaganus (Bernhauer, 1914) [ Platydracus , and as subsp.], 40

pseudopatricius (J. Müller, 1926) [subsp. of Platydracus pseudopaganus (Bernhauer, 1914) ], 40

Pseudotasgius Seidlitz, 1891 [= Tasgius (Tasgius) ], 36, 45

pubescens (DeGeer, 1774) (Staphylinus) [ Dinothenarus (Dinothenarus) ], 26, 41

pullus Hochhuth, 1849 [ Ocypus (Matidus) ], 34, 43

pyrenaeus (J. Müller, 1924) [subsp. of Ocypus (Matidus) pedemontanus (J. Müller, 1924) ], 43

quadraticeps (Casey, 1924) [= Platydracus zonatus (Gravenhorst, 1802) ], 41

quadraticeps (Ménétriés, 1832) [ Ocypus (Matidus) ], 43

quadrimaculatus (Cameron, 1932) [ Ocypus (Ocypus) ], 31, 32, 42

Quediina , 6–10, 12– 14, 15, 17

Quedionuchus Sharp, 1884 (subgen. of Quedius ),8, 14, 15

Quedius Stephens, 1829 , 14, 15

queinneci Coiffait, 1984 ( Ocypus ?) [incertae sedis], 47

Rabigus Mulsant and Rey, 1876 , 10

rambouseki (J. Müller, 1925) [= Ocypus (Pseudocypus) nigroaeneus Sharp, 1889 ], 44

Raphirus Stephens, 1829 (subgen. of Quedius ), 14

Rayacheila Motschulsky, 1845 [subgenus of Tasgius ], 2, 6–8, 11, 16, 20, 36, 37, 46

reimoseri (Bernhauer, 1906) [ Ocypus (Matidus) ], 43

reitterianus (Bernhauer, 1933) [ Platydracus ], 40

rhaeticus Eppelsheim, 1873 [ Ocypus (Matidus) ], 42

riojanus Hozman, 1977 [ Platydracus ], 40

robustus Coiffait, 1964 [subsp. of Ocypus (Matidus) milleri Quedenfeldt, 1882 ], 43

rodopensis Coiffait, 1971 [subsp. of Ocypus (Ocypus) ophthalmicus (Scopoli, 1763) ], 42

rodopensis (Coiffait, 1971) [= Ocypus (Pseudocypus) picipennis picipennis (Fabricius, 1792) ], 45

rossii Jarrige, 1954 [= Ocypus (Pseudocypus) sericeicollis (Ménétriés, 1832) ], 35, 45

rubricornis (Adám, 1987) [ Staphylinus ], 41

rubripennis (Reiche and Saulcy, 1856) [ Tasgius (Rayacheila) ], 46

ruficeps (Cameron, 1932) [ Dinothenarus (Dinothenarus) ], 41

ruficornis Bernhauer, 1913 [= Staphylinus rubricornis (Adám, 1987) ], 41

rufipennis Bernhauer, 1915 [ Naddia ], 38

rufipes Eppelsheim, 1884 (Ocypus) [= Tasgius (Rayacheila) eppelsheimianus (Jakobson, 1909) ], 37

rufipes Latreille, 1806 (Astrapaeus) [= Tasgius (Tasgius) pedator (Gravenhorst, 1802) ], 36, 37

rutilicauda (Horn, 1879) [ Platydracus ], 40

sabi Naomi, 1992 [ Ocypus (Pseudocypus) ], 45

sachalinensis (Matsumura, 1911) [ Platydracus ], 40

sagaris Smetana, 1992 [ Dinothenarus (Dinothenarus) ], 41

sahlbergi (J. Müller, 1926) [subsp. of Tasgius (Rayacheila) melanarius (Heer, 1839) ], 46

saphyrinus (LeConte, 1861) [ Dinothenarus (Parabemus) ], 42

Scariphaeus Erichson, 1839 , 15

schatzmayri (J. Müller, 1923) [subsp. of Ocypus (Pseudocypus) obscuroaeneus Fairmaire, 1852 ], 44

scutiger Sharp, 1889 [ Ocypus (Pseudocypus) ], 45

semenowi Reitter, 1887 [ Ocypus (Pseudocypus) ], 45

semialatus (J. Müller, 1904) [subsp. of Ocypus (Matidus) nitens (Schrank, 1781) ], 43

semipurpureus (Kraatz, 1859) [ Platydracus ], 40

septentrionalis Watanabe, 1984 [ Ocypus (Pseudocypus) ], 45

sepulchralis (Erichson, 1839) [ Platydracus ], 40

sericeicollis (Ménétriés, 1832) (Pseudocypus) [ Ocypus (Pseudocypus) ], 35, 45

sericeomicans (Bernhauer, 1931) [ Ocypus (Pseudocypus) ], 45

serotinus (Adám, 1992) [ Ocypus (Pseudocypus) ], 45

sharpi (Fauvel, 1901) [ Platydracus ], 40

sibiricus (Gebler, 1830) [ Dinothenarus (Parabemus) ], 42

sikkimensis (Bernhauer, 1920) [ Tasgius (Tasgius) ], 46

sikkimensis Coiffait, 1985 (Ocypus) [incertae sedis], 47

sikkimensis Wendeler, 1927 [ Creophilus ], 38

silensis Fiori, 1894 [= Ocypus (Matidus) italicus (Aragona, 1830) ], 43

similis: (Fabricius, 1792) [= Ocypus (Matidus) nitens (Schrank, 1781) ], 34, 43

similis (Paykull, 1789) (Staphylinus) [ Tasgius (Rayacheila) ], 34

simulator Eppelsheim, 1878 [ Ocypus (Matidus) ], 34, 35, 43

simulator Kirschenblat, 1936 [ Ontholestes ], 38

smetanai Naomi, 1992 [ Ocypus (Pseudocypus) ], 45

Smilax Laporte, 1835 , 10, 14

solarii (J. Müller, 1923) [ Ocypus (Matidus) ], 43

solskyi (Fauvel, 1875) (Staphylinus) [ Tasgius (Rayacheila) , and as subsp.], 36, 46

sparsus (Cameron, 1932) [ Platydracus ], 40

speculifrons (Bernhauer, 1939) [ Platydracus ], 40

spilleri Bernhauer, 1923 [ Physetops ], 42

springeri J. Müller, 1923 [subsp. of Staphylinus erythropterus Linne´, 1758], 41

Staphylinina , 6–15, 16 –18, 22

Staphylinus Linne´, 1758, 2, 3, 6–8, 10, 11, 16– 18, 20, 25, 27, 41

stercorarius (Olivier, 1795) [ Platydracus and as subsp.], 9, 24, 40

stevensi (Cameron, 1932) [ Wasmannellus ], 45

Strouhalium Scheerpeltz, 1962 , 15

Styngetus Sharp, 1884 , 17, 18

subaeneipennis (Scheerpeltz, 1976) [ Platydracus ], 40

subaenescens Wollaston, 1864 [ Ocypus (Pseudocypus) ], 45

subaeneus (Roth, 1851) [ Platydracus ], 40

subauronotatus Coiffait, 1977 [ Platydracus ], 40

submarmorellus (Schubert, 1908) [ Platydracus ], 40

submetallicus (LeConte, 1861) [= Platydracus tarsalis (Mannerheim, 1843) ], 40

subtilis Tikhomirova, 1973 ( Ocypus ( Pseudocypus )) [ Ocypus (Aulacocypus) ], 45

subviridis (Bernhauer, 1933) [ Platydracus ], 40

sumakowi (Bernhauer, 1911) [ Platydracus ], 40

suturalis Matsumura, 1911 ( Staphylinus ( Ocypus )) [incertae sedis], 47

svozili M. Dvořák, 1984 [ Ocypus (Pseudocypus) ], 45

sylvaticus Wollaston, 1865 [ Ocypus (Pseudocypus) ], 31, 45

syriacus Baudi, 1848 [ Ocypus (Matidus) , and as subsp.], 43

szechuanensis (Bernhauer, 1935) [ Dinothenarus (Parabemus) ], 42

Tachinoporus Cameron, 1928 , 10 Tachyporini , 9, 10, 14 taiwanensis Shibata, 1979 [ Naddia ], 38 tarsalis (Mannerheim, 1843) [ Platydracus ], 40 Tasgius Stephens, 1829 [and as subgenus], 2, 3,

6, 7–11, 16, 17, 20, 36, 45 tataricus (Pallas, 1773) (Staphylinus) [ Physetops ],

28, 42 tauricus (J. Müller, 1932) [= Ocypus (Pseudocy­

pus) mus (Brulle´, 1832)], 44 temporalis (Casey, 1915) [ Platydracus ], 41 tenebricosus (Gravenhorst, 1846) [ Ocypus (Mati­

dus)], 10, 43

tenuicornis (Kraatz, 1859) [ Ontholestes ], 38

teruelensis (Coiffait, 1976) [= Ocypus (Pseudocypus) picipennis nevadensis (J. Müller, 1926)], 45

tessellatus (Geoffroy, 1785) [ Ontholestes ], 38

testaceipes Fairmaire, 1887 (Ocypus) [incertae sedis], 47

teter (Bondroit, 1913) [= Platydracus aureofasciatus (Motschulsky, 1862) ], 39

Thinopinus LeConte, 1852 , 8, 9, 11, 12, 16, 17, 18, 22

Thoracostrongylus Bernhauer, 1915 , 8–11, 16, 17, 19, 23, 39

tomentosus (Baudi, 1869) (Staphylinus) [= Ocypus (Pseudocypus) orientis , nom. nov.],

tomentosus (Gravenhorst, 1802) (Staphylinus) [ Platydracus ], 34, 41

torvus Smetana, 1965 [ Ocypus (Matidus) ], 43

transadriaticus (J. Müller, 1926) [= Ocypus (Pseudocypus) mus (Brulle´, 1832)], 44

transversiceps (Luze, 1904) [ Tasgius (Tasgius) ], 46

trapezensis Coiffait, 1964 [ Ocypus (Matidus) ], 43

Triacrina, 12

Triacrus Nordmann, 1837 , 18

Trichocosmetes Kraatz, 1859 , 18

Trichoderma Stephens, 1829 [= Dinothenarus ], 2

tricinctus (Aragona, 1830) [ Tasgius (Rayacheila) ], 46

tricolor (Coiffait, 1974) [ Tasgius (Rayacheila) ], 46

Trigonopselaphus Gemminger and Harold, 1868 , 18

triplicans (Casey, 1924) [= Platydracus viridanus (Horn, 1879) ], 41

tristis Bernhauer, 1920 [ Wasmannellus ], 35, 45

turcicus (Bernhauer, 1923) [ Ocypus (Matidus) ], 43

Tympanophorus Nordmann, 1837 , 17, 18