Kalophrynus palmatissimus Kiew, 1984

Zug, George R., 2015, Morphology and Systematics of Kalophrynus interlineatus-pleurostigma Populations (Anura: Microhylidae: Kalophryninae) and a Taxonomy of the Genus Kalophrynus Tschudi, Asian Sticky Frogs, Proceedings of the California Academy of Sciences 62 (5), pp. 135-190 : 173

Kalophrynus palmatissimus Kiew, 1984 Web-footed Sticky Frog

Kalophrynus palmatissimus Kiew, 1984b, Malayan Nature Journal 37:146 [type locality: “lowland dipterocarp forest at about 75 m a.s.l. at the Pasoh Forest Reserve in Negeri Sembilian ”, Malaysia (Malaya).

TYPE MATERIAL.— HOLOTYPE: British Museum of Natural History 1982.1508 . PARATYPES: BMNH 1982 .1509–1523 , FMNH two unnumbered (now 216461–462), University of Malaysia one unnumbered. All by original designation .

DEFINITION.— Medium-sized, adults 31.2– 38. 3 mm SVL (n = 19, not sexed); head moderately large 29–34 % HeadL/ SVL; head slightly broader than long 102–116 % HeadW/HeadL; snout moderately broad NA % SnW/HeadL; naris closer to snout than to eye NA % NarEye/SnEye; eye moderately large 36–48 % EyeD/HeadL; tympanum visible and smaller than eye 55–70 % Tymp/EyeD; moderate length forelimb 24–28 % Forarm/ SVL and forearm shorter than crus length 56–66 % Forarm/CrusL; hindlimb moderate NA % HndlL/ SVL, crus moderate length 41–46 % CrusL/ SVL, and NA % CrusL/ThghL; hindfoot well developed NA % HndfL/CrusL.

Vomerine teeth presence or absence not reported; palatal fold morphology not reported; tongue elongate with rounded tip.

Fingers without web; lengths 3>2>1=4; tips rounded and not dilated; subarticular tubercles prominent, number beneath each finger not reported; palmar tubercle not reported. Toes strongly although not fully webbed, web between toes 3 and four to tip of 3, toe lengths 4>3>5>2>1; tips rounded; subarticular tubercles prominent on digits 1, 2, 3, 4, none beneath 5; oval inner and small indistinct outer metatarsal tubercles.

In life, dorsum light brown to reddish brown with darker brown blotch extending from between eyes and extending posteriorly and breaking into patches on trunk; no inguinal ocelli; lores and temporal area to axilla dark brown, trunk side reddish brown; venter yellowish with two faint longitudinal dusky bars on throat and gray mottling on sides and posterior of abdomen.

ETYMOLOGY.— The specific epithet palmatissimus refers to the strongly webbed hindfeet, the greatest amount of webbing seen in the genus Kalophrynus .

DISTRIBUTION.— This species is a resident of southern Peninsular Malaysia and presently know from only three localities in the provinces of Selangor, Negri Sembilan , and Johor.

NATURAL HISTORY.— Kalophrynus palmatissimus is a terrestrial frog of lowland dipterocarp forests. Kiew (1984) provides details on breeding and tadpole development and morphology.

COMMENTS.— Preceding information extracted from Kiew (1984), Chan et al. (2011), and Dehling (2011).