Philothis (Farabius) hexeris Reichardt, 1930

Lackner, Tomáš, 2010, Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae), Acta Entomologica Musei Nationalis Pragae (suppl.) 50, pp. 1-254 : 176-181

publication ID	https://doi.org/ 10.5281/zenodo.4272127
DOI	https://doi.org/10.5281/zenodo.4341995
persistent identifier	https://treatment.plazi.org/id/0385915E-FF24-0991-6037-FA66CFD2FDCD
treatment provided by	Felipe
scientific name	Philothis (Farabius) hexeris Reichardt, 1930
status

Treatment

Philothis (Farabius) hexeris Reichardt, 1930 View in CoL

( Figs. 34 View Figs , 39 View Figs , 65 View Figs , 99 View Figs , 130 View Figs , 174 View Figs , 559–576 View Figs View Figs )

Philothis (Farabius) hexeris Reichardt, 1930: 299 View in CoL .

Philothis (Farabius) hexeris: REICHARDT (1941) View in CoL : 340, 343, 409, Fig. 175B; KRYZHANOVSKIJ View Figs & REICHARDT (1976): 245, 249, Fig. 485 View Figs ; MAZUR (1984): 110; OLEXA (1990): 153, Figs. 4 View Fig , 13, 20, 26 View Figs , 48, 49 View Figs ; MAZUR (1997): 268; MAZUR (2004): 96.

Type locality. Turkmenistan, Farab.

Type material examined. HOLOTYPE: spec., ‘Buchara occ. / Farab 2.iv.[1]911 / A Hohlbeck [printed-written] // Philothis (Farabius) / hexeris sp. n. / Holotypus / Reichardt det. [printed-written] // Coll. Semenov-Tian-Shansky [printed] // Holotypus [red lablel, printed] // two golden round labels’ ( ZIN).

Additional material examined. UZBEKISTAN: Chiva [= Khiva], Karakum, 1.–5.v.1979, 1 ♁ 2 spec., A. Olexa lgt .; Buchara , Kyzylkum, 30.iv.1979, 1 ♁, A. Olexa lgt .; ditto, but 27.iv.1979, 1 ♁ 1♀ 1 spec. ( TLAN) .

Redescription. Body length: PEL: 2.25–2.50 mm; APW: 0.875–1.00 mm; PPW: 1.75–1.875 mm; EL: 1.375–1.50 mm; EW: 1.875 –2.075 mm.

Body ( Figs. 559–560 View Figs ) shortly oval, moderately convex from above, underside very convex, cuticle reddish brown, shiny; legs, mouthparts and antennal scape rufopiceous.

Antennal scape ( Fig. 562 View Figs ), with numerous long setae; club comparatively small, round, without visible articulation, apical third imbricate, with thick dense short yellow sensilla concealing apical sensory area; basal surface glabrous; sensory structures of antennal club ( Fig. 34 View Figs ) in form of one apical sensory area and one stipe-shaped vesicle situated under it.

Mouthparts. Mandibles ( Fig. 99 View Figs ) stout, slightly curved, mandibular apex blunt; sub-apical tooth on inner margin of left mandible moderately large, perpendicular; disc of labrum ( Fig. 65 View Figs ) slightly depressed medially, almost smooth; labral process and labral fold significantly developed, completely concealing epipharynx; terminal labial palpomere elongated, its width about one-third its length; mentum almost square-shaped, imbricate, anterior margin ( Fig. 130 View Figs ) slightly shallowly emarginate medially with tiny median notch surrounded with few long setae; lateral margins with one row of sparse shorter ramose setae; disc of mentum with several long setae, almost as long as those of anterior margin. Cardo of maxilla with several short setae on lateral margin; stipes triangular, with numerous much longer setae; terminal maxillary palpomere elongated, its width about one-third its length, approximately three times as long as penultimate.

Clypeus ( Fig. 562 View Figs ) rectangular, smooth, rounded laterally; frontal stria represented by short vague fragments above antennal insertions; supraorbital stria absent; frontal disc ( Fig. 562 View Figs ) smooth; eyes flattened, invisible from above.

Pronotal sides ( Fig. 559 View Figs ) with a shallow longitudinal excavation, weakly convergent anteriorly; apical angles prominent, anterior pronotal margin with a deep incision for head, almost straight medially; marginal pronotal stria well impressed, carinate, shortly interrupted behind head (at times complete); disc moderately convex, entirely smooth except for a row of small round punctures along pronotal base; disc with shallow depressions and bulges, in medio-lateral part with a single deep fovea on each side; pronotal hypomeron with long yellow setae; scutellum very small, almost invisible.

Elytral humeri weakly developed, elytral epipleura smooth; marginal epipleural stria complete, thin; marginal elytral stria well impressed, carinate, continued as complete apical elytral stria. Humeral elytral stria vaguely impressed, surrounded by tiny strioles, at times indistinguishable; inner subhumeral stria rather well developed, at times parallel to first dorsal elytral stria; outer subhumeral stria shortly impressed on basal fourth (at times absent). Only first dorsal elytral stria developed, shortened on basal fourth, almost reaching elytral apex, apically bent inwardly; other dorsal elytral striae absent; sutural elytral stria vaguely impressed, apically connected with apical elytral stria, on basal half slightly distanced from elytral suture, interspace between it and elytral suture on basal third forming a longitudinal bulge, continued along elytral base as a bisinuate basal elytral stria surrounding a vague low bulge, occasionally obliterated. Elytral disc with dense, round, deep punctation, interspaces smooth, punctation becomes sparser towards elytral suture; area around scutellum and on transverse basal bulge smooth; along elytral apex punctures confluent.

Propygidium completely exposed, approximately 1.5 times as broad as long, with dense shallow confluent punctures; pygidium distinctly longer than broad, convex, with similar punctation.

Anterior margin of median portion of prosternum ( Fig. 564 View Figs ) obtuse-angulate; marginal prosternal stria well impressed, complete; prosternal process substrigulate, with dense long setae, gradually sloping down anteriorly; carinal prosternal striae present only on prosternal apophysis, strongly convergent anteriorly; lateral prosternal striae absent.

Anterior margin of mesoventrite ( Fig. 566 View Figs ) bisinuate; discal marginal mesoventral stria well impressed, anteriorly almost straight, slightly carinate, distanced from anterior margin of mesoventrite; laterally deeply impressed, carinate; disc of mesoventrite with scattered microscopic punctures; meso-metaventral sutural stria ( Fig. 566 View Figs ) well impressed, almost straight, slightly projected outwardly. Intercoxal disc of metaventrite with scattered microscopic punctures, almost smooth; lateral metaventral stria ( Fig. 565 View Figs ) well impressed, carinate, obliquely arcuate, shortened apically; lateral disc of metaventrite ( Fig. 565 View Figs ) concave, with oblong shallow punctures, fringed with long dense setae; metepisternum + fused metepimeron ( Fig. 565 View Figs ) densely setose, punctation almost unrecognizable beneath the setae.

Intercoxal disc of the first abdominal sternite almost completely striate laterally, with scattered microscopic punctation, becoming more prominent along apical margin; lateral discs of all visible abdominal sternites setose laterally.

Protibia ( Fig. 567 View Figs ) moderately dilated, anterior margin truncate, formed by anterior margin of distal-most large triangular tooth; outer margin of protibia apart from this tooth with one more, somewhat shorter triangular tooth, topped with short rounded blunt denticle; setae of outer row confined to basal third, thin, sparse, moderately long; setae of median row thicker, more regular; protarsal groove and anterior protibial stria inconspicuous; apical margin of protibia posteriorly without denticles; outer part of posterior surface of protibia smooth, vaguely separated from smooth median part of posterior surface by indistinct ridge with few obliterated rudimentary microscopic denticles; posterior protibial stria inconspicuous; inner margin of protibia with long lamelliform dense setae, apical third of inner margin of protibia with one row of shorter, morphologically different setae; protibial spur microscopic, entombed in apical margin of protibia; protarsus rudimentary, apical tarsomere with two short thin claws, approximately half as long as apical tarsomere itself.

Mesotibia ( Fig. 561 View Figs ) slender, outer margin with one double row of sparse, morphologically different short denticles significantly growing in size and density near tarsal insertion, denticles continuous anteriorly along apical margin; setae of outer row growing near inner mesotibial margin, dense and long, covering almost entire posterior mesotibial surface; setae of median row absent; posterior mesotibial stria inconspicuous; mesotibial spur conspicuously long, slender; anterior surface of mesotibia smooth; anterior mesotibial stria thin, shortened apically; inner margin of mesotibia with rather sparse row of long lamelliform setae; mesotarsus telescope-like, diameter of tarsomeres diminishing apically, each tarsomere with two long lamelliform strongly sclerotized setae, one posteriorly and one anteriorly; claws of apical mesotarsomeres thin, hair-like, several times as long as mesotarsomeres itself. Metatibia conspicuously dilated, thickened apically; outer margin (especially on thickened apical third) with up to three rows (two rows of short rounded and thick, dense, much longer denticles) of morphologically different denticles, another row of conspicuously short round regular denticles markedly shifted from it, present on anterior surface of metatibia and only observable from ventral view. Posterior surface of metatibia entirely covered with dense rows of rather long setae, distinction between outer and median rows unclear; posterior metatibial stria inconspicuous; anterior surface of metatibia and metatarsus otherwise similar to mesotibia and mesotarsus.

Male genitalia. Eighth sternite ( Figs. 568–569 View Figs ) moderately narrowing apically, longitudinally separated medially almost along its entire length, apically with tiny inflatable membrane (velum), without setae; eighth tergite and eighth sternite fused laterally ( Fig. 570 View Figs ). Morphology of 9 th tergite ( Figs. 571–572 View Figs ) typical for the subfamily; spiculum gastrale ( Fig. 575 View Figs ) expanded on both ends. Basal piece of aedeagus ( Figs. 573–564 View Figs ) short, ratio of its length: length of parameres ca. 5.30; aedeagus slender; parameres fused along their basal two-thirds; aedeagus gently curved ventrad ( Fig. 574 View Figs ).