Promitobates weissbier, Bragagnolo, Cibele & Pinto-Da-Rocha, Ricardo, 2012

Bragagnolo, Cibele & Pinto-Da-Rocha, Ricardo, 2012, Systematic review of Promitobates Roewer, 1913 and cladistic analysis of Mitobatinae Simon, 1879 (Arachnida: Opiliones: Gonyleptidae), Zootaxa 3308, pp. 1-48 : 39-41

Promitobates weissbier sp. n.

( Figs. 6 C, 8E, 10E, 12C, 14C, 19A–C)

Type material. BRAZIL. São Paulo: Ribeirão Pires (Gruta da Quarta Divisão), 20.VIII.1992, E. Trajano & P. Gnaspini leg.; holotype 3 and 2 paratypes (1 3, 1 Ƥ) ( MZSP 17161].

Etymology. Weissbier is a kind of ale bier, made ??with wheat.

Diagnosis. It ressembles P. trapista sp. n. by coloration of dorsal scutum, chelicerae, pedipalps and legs I and III dark-yellow and apex of coxa IV, trochanter IV and femur IV black. It can be distinguished by coxa IV with bifid apophysis and color of tubercles of dorsal scutum, greenish.

Male description. Measurements (holotype). Dorsal scutum; maximum width: 4.9; total length: 5.3; prosoma length: 2.2. Leg IV; length of external apophysis of coxa: 1.2; femur IV length: 48.8. Tarsal formula: 9; 16; 11; 10.

Dorsum ( Figs. 6 C, 8E). Anterior margin of prosoma with frontal hump with one row of small tubercles. Prosoma with tubercles behind ocularium arranged in “V”. Ocularium with two high spiniform apophyses (apex rounded) and two pairs of small tubercles behind them. Area I with two rows of tubercles up to the groove II and small scattered tubercles. Area II densely tuberculated. Area III with two high apophyses (apex rounded) and small tubercles. Area IV divided by a longitudinal groove, totally recovered by tubercles. Lateral areas with rows of tubercles which are placed from ocularium to area II. Posterior margin with two or three tubercles in the corners and one pair of median spiniform tubercles. Free tergites with one row of tubercles. Anal operculum unarmed.

Chelicera. Segment I with two setiferous tubercles; II unarmed.

Pedipalp. Trochanter with a small ventral spine. Femur with a small ventral basal spine and one subapical, prolateral spine. Tibia setation: mesal IiIi; ectal IiIi. Tarsus setation: mesal IIi; ectal IIi.

Venter. Coxa I with median row of five setiferous tubercles, two apical tubercles and scattered granules; trochanter I with median tubercle. Coxa II with median row of six small setiferous tubercles and two apical tubercles; trochanter II with median tubercle. Coxa III with small setiferous scattered distributed tubercles, and two rows of tubercles, increasing in size distally; coxa IV with four spiniform tubercles on posterior margin. Genital area unarmed; free sternites with one row of tubercles; anal operculum with small granules.

Legs ( Fig. 10 E). Coxa I with anterior dorso basal apophysis; coxa II with one anterior dorso basal apophysis and one posterior; coxa III unarmed; coxa IV with few high tubercles, one internal spiniform apophysis and one external apical bifid apophysis, directed ventrally and with one basal rounded tubercle. Trochanter I–III with small scattered tubercles. Trochanter IV with one dorsal subapical apophysis, one external basal apophysis and one internal apical spiniform apophysis. Femur IV straight, long and unarmed.

Coloration (in ethanol). Dorsal scutum and apophyses of area III dark-yellow. Apex of coxa IV, trochanter IV and femur IV black. Tubercles of dorsal scutum greenish.

Penis ( Fig. 19 A–C). Ventral plate convex, apex slightly curved and distal margin corner rounded and directed ventrally; three pairs of distal setae, three pairs of basal setae, arranged in “V” and one pair of small basal ventral setae. Stylus unarmed, apex straight and swollen. Ventral process almost parallel to the stylus, apex spoon-shaped, with serrate margins directed ventrally.

Female description ( Figs. 12 C, 14C). Measurements (n=1). Dorsal scutum; maximum width: 5.3; total length: 5.7; prosoma length: 2.1. Femur IV length IV: 15.1. Tarsal formula: 7; 15; 10; 11.

Apophyses of area III taller than male. Free tergite III with one median spiniform apophysis. External apophysis of coxa IV reduced and single-branched; internal apophysis absent. Trochanter IV unarmed. Femur IV shorter than male.

Geographical distribution. ( Fig. 3 B). Only known from type locality.

Figures

FIGURE 3. Geographical distribution of Promitobates spp ..

FIGURE 6. Dorsal view of males of Promitobates A. P. a l e sp. n. (MZSP 27404); B. P. trapista sp. n. (MZSP 27406); C. P. wiessbier sp. n. (MZSP 16171); D. lager sp. n. (MNRJ 4545). Scale bar, 1 mm.

FIGURE 10. Dorsal view of male coxa, trochanter and femur IV of Promitobates A e B. P. viridigranulatus, A. MZSP 27401; B. MHNCI 6527; C. P. ale sp. n. (MZSP 27404); D. P. trapista sp. n. (MZSP 27406); E. P. weissbier sp. n. (MZSP 16171); F. P lager sp. n. (MNRJ 04545); Scale bar, 1 mm.

FIGURE 12. Dorsal view of females of Promitobates. A. P. viridigranulatus (MZSP 18128); B. P. a l e sp. n. (MZSP 18635); C. P. weissbier sp. n. (MZSP 17161, paratype); D. P. l a ge r sp. n. (MZSP 17249). Scale bar, 1 mm.

FIGURE 19. Distal part of penis of Promitobates. A – C. P. weissbier sp. n. (MZSP- 17161); A. dorsal view; B. lateral view; C. ventral view. D – F. P. l a g e r sp. n. (MZSP- 17047); D. dorsal view; E. lateral view; F. ventral view. Scale bar = 0,05 mm.