Parena testacea (Chaudoir, 1872)

Shi, Hongliang & Liang, Hongbin, 2023, Taxonomic revision of the genus Parena Motschulsky, 1860 (Coleoptera, Carabidae, Lebiini, Metallicina), Zootaxa 5286 (1), pp. 1-144 : 38

3. Parena testacea species group

This species group contains five allopatric species, distributed all through the Oriental Realm and eastern to Papua New Guinea and the Solomon Islands. They are characterized by: dorsum pale yellow, elytra entirely yellow or with large dark median patch; tarsi black, distinctly darker than other portions of legs; antennae bicolor, basal one or three antennomeres yellow, apical antennomeres black; elytral striae not incised, replaced by lines of punctures, intervals flat; elytral sutural angles pointed or not, forming denticles or spines, outer apical angles well rounded to distinctly angulate; median lobe of aedeagus slightly stouter than previous two groups, gradually attenuate to apex, apical lamella more or less bent to dorsum.